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Dipterocarpaceae

Last edited by Pieter B. Pelser, 18 August 2017

Anisoptera Korth.

Photos
  1. Anisoptera aurea Foxw. PJS 67 (1938) 271; --P.S.Ashton, Fl. Males. ser. 1, 9 (1982) 334. --Anisoptera curtisii auct. non Dyer ex King; --Foxw., PJS 6 c (1911) Bot. 255; --Merr., EPFP 3 (1923) 92. LUZON: Isabela, Quezon, Laguna, Camarines Norte, POLILLO. Mixed dipterocarp forest, especially on ridges to 200m.
  2. Anisoptera brunnea Foxw., PJS 6c (1911) 254; --EPFP 3 (1923) 92; --Type: FB 11292 Klemme (NY, US, iso), Luzon: Cagayan prov., Apr-1908. LUZON: Cagayan, Ilocos Norte. Photos
  3. Anisoptera costata Korth., Kruidk. (1941) 67; --P.S.Ashton, Fl. Males. ser. 1, 9 (1982) 330. --Anisoptera mindanensis Foxw., PJS 13 c (1918) Bot. 181; --EPFP 3 (1923) 92. Myanmar, Thailand, Cambodia, S Vietnam, Peninsular Malaysia, Sumatra, W Java, Borneo (SE Kalimantan, Sabah, Brunei, NE Sarawak), Philippines. MINDANAO (Zamboanga, one record). Primary lowland forest. Conservation status: Endangered (DAO 2017-11).
  4. Anisoptera thurifera (Blanco) Blume, Ann. Mus. Bot. Lugd.-Bat. 2 (1852) 42; --EPFP 3 (1923) 92; --P.S.Ashton, Fl. Males. ser. 1, 9 (1982) 333; --Dipterocarpus thurifer (Blanco) Blanco, Fl. Filip. ed. 2 (1845) 310; Fl. Filip. ed. 3 2 (1878) 212; --Mocanera thurifera Blanco, Fl. Filip. (1837) 446. --Mocanera mayapis Blanco, Fl. Filip. (1837) 449; --Dipterocarpus mayapis (Blanco) Blanco, Fl. Filip. ed. 2 (1845) 212; --Shorea mayapis (Blanco) Blume, Mus. Bot. Ludg.-Bat. 2 (1852) 33. --Anisoptera vidaliana Brandis, Journ. Linn. Soc. Bot. 31 (1895) 44. --Anisoptera tomentosa Brandis, Journ. Linn. Soc. Bot. 31 (1895) 45. --Anisoptera calophylla Perk., Frag. Fl. Philip. (1904) 22. Philippines, Sulawesi, Moluccas (Morotai, Halmahera, Bačan, Obi, Aru Is.), New Guinea. BABUYAN ISLS (CALAYAN), LUZON (widespread in most provinces), MINDORO, TICAO, MASBATE, SIBUYAN, PANAY, NEGROS. Dominant in evergreen and semi-evergreen primary forests below 750m; also occurring in some types of secondary forests. Photos

 Balanocarpus Bedd. = Hopea

Dipterocarpus C.F.Gaertn.

Last edited by Pieter B. Pelser, 18 August 2017

Photos
  1. Dipterocarpus alatus Roxb. ex G.Don., Gen. Hist. 1 (1831) 813; --Type: Roxburgh s.n. (BR), Myanmar. --Dipterocarpus philippinensis Foxw., PJS 13 c (1918) Bot. 179; --Merr., EPFP 3 (1923) 90; --P.S.Ashton, Fl. Males. ser. 1, 9 (1982) 325. Mainland SE Asia, Philippines. LUZON: Abra, Nueva Vizcaya, Bataan. Primary mixed dipterocarp forest in seasonal areas. Rare. Conservation status: Vulnerable (DAO 2017-11). Photos
  2. Dipterocarpus condorensis Pierre, Fl. For. Cochinch. 3, 14 (1889) Pl. 214; --Type from Vietnam.
  3. Dipterocarpus eurhynchus Miq., Sum. (1862) 485; --P.S.Ashton, Fl. Males. ser. 1, 9 (1982) 324. --Dipterocarpus basilanicus Foxw., PJS 13 c (1918) Bot. 179; --Merr., EPFP 3 (1923) 88. Peninsular Malaysia (E coast), Sumatra (E Aceh, Langsa), Singkep, Banka, Borneo (W Borneo, Sarawak, Brunei), SW Philippines. BASILAN. Local, on undulating land in lowland primary dipterocarp forest. Rare. Conservation status: Endangered (DAO 2017-11).
  4. Dipterocarpus gracilis Blume, Bijdr. (1825) 224; --Merr., EPFP 3 (1923) 89; --P.S.Ashton, Fl. Males. ser. 1, 9 (1982) 301. --Dipterocarpus vernicifluus (Blanco) Blanco, Fl. Filip. ed. 2 (1845) 314; --Merr., EPFP 3 (1923) 91; --Mocanera verniciflua Blanco, Fl. Filip. (1837) 451. --Dipterocarpus fulvus Blume., Mus. Bot. 2 (1852) 37. --Dipterocarpus velutinus S.Vidal, Pl. Vasc. Filip. (1886) 59. Andaman Isls, Chittagong, Myanmar, SE & Peninsular Thailand, Sumatra, Peninsular Malaysia, W Java, Borneo (Sampit and Rejang Valley eastward), Philippines. LUZON: Zambales, Bataan (Mt Mariveles, Lamao River), Bulacan (Sibul Forest, Sitio Binalangoan), Rizal (Bosoboso; Antipolo), Quezon (Umiray), POLILLO, MINDORO, MARINDUQUE, PALAWAN (Pagdanan Range, Ibangley Brookside Hill; Quezon, Barangay Tuangan;), NEGROS: Negros Occidental, MINDANAO: Zamboanga del Norte (Duhinid), Cotabato, Davao, Surigao. Often gregarious in seasonal semi-evergreen dipterocarp forest on red soils; becoming scattered, rare, and confined to fertile red soils in everwet areas, below 800m. Conservation status: Vulnerable (DAO 2017-11).
  5. Dipterocarpus grandiflorus (Blanco) Blanco, Fl. Filip. ed. 2 (1845) 314; --Merr., EPFP 3 (1923) 89; --P.S.Ashton, Fl. Males. ser. 1, 9 (1982) 317; -Mocanera grandiflora Blanco., Fl. Filip. (1837) 451. Andaman Isls, Mergui, Peninsular Thailand, Peninsular Malaysia, Singapore, NE Sumatra and adjacent islands (Simalur, Musala, Riau Archipelago, Banka, Billiton), Karimata, Borneo (SW to Sabah) and the Philippines, particularly in the more seasonal western parts. LUZON (widespread in most provinces), MINDORO, PALAWAN, SIBUYAN, BILIRAN, SAMAR, PANAY, NEGROS, MINDANAO: Misamis, Agusan. Often common and dominant in semi-evergreen lowland and medium elevation dipterocarp forests, on small islands and on coastal hills in less seasonal part. Conservation status: Vulnerable (DAO 2017-11). Photos
  6. Dipterocarpus hasseltii Blume, Fl. Jav. (1829) 22; --Merr., EPFP 3 (1923) 89; --P.S.Ashton, Fl. Males. ser. 1, 9 (1982) 306, Fig. 23, 24; --Type from Java. --Dipterocarpus subalpinus Foxw., LPB 6 (1913) 1950. Peninsular Thailand, Malesia: around periodically dry borders of the everwet zone of the Sunda Shelf, C and N Peninsular Malaysia, Sumatra, W Java, Bali, S and E Borneo to SE Sabah. Widespread in the Philippines. LUZON: Isabela, Nueva Ecija, Laguna, PALAWAN, NEGROS, BILIRAN, LEYTE, MINDANAO: Zamboanga, BASILAN. Primary forests at low and medium elevation, ascending to 900m. Conservation status: Vulnerable (DAO 2017-11). Photos
  7. Dipterocarpus kerrii King, J. As. Soc. Beng. 622 (1893) 93; --P.S.Ashton, Fl. Males. ser. 1, 9 (1982) 305; --Type from Peninsular Malaysia (Malacca). --Dipterocarpus cuneatus Foxw., PJS 13 c (1918) Bot. 178; --Merr., EPFP 3 (1923) 89; --Type: FB 21469. --Dipterocarpus perturbinatus Foxw., PJS 13 (1918) 177. --Dipterocarpus obconicus Foxw., PJS 13 c (1918) Bot. 178; --Merr., EPFP 3 (1923) 90; --Type: Elmer 13498. Andaman Isls, Myanmar, Peninsular Thailand and Malaysia, Sumatra, Borneo (Sabah: Sandakan area) and the Philippines. LUZON: Laguna, Quezon, Camarines, Albay, Sorsogon, MINDANAO: Agusan del Norte. On lateritic soils in seasonally dry lowland primary dipterocarp forests, usually below 400m. Frequently gregarious.
  8. Dipterocarpus kunstleri King., J. As. Soc. Beng. 62, 2 (1893) 96; --P.S.Ashton, Fl. Males. ser. 1, 9 (1982) 309; --Type from Malaysia. --Dipterocarpus speciosus Brandis, J. Linn. Soc. Bot. 31 (1895) 38; --Merr., EPFP 3 (1923) 90; --Type: Vidal 2160 (K), Basilan. Peninsular Malesia (excluding seasonal areas), N, W and SE Sumatra, Simalur, Marsala, Banka), Borneo (SE Borneo, Sabah, Brunei, Sarawak), Philippines. LUZON: Cagayan, Isabela, Quezon, Camarines, Albay, POLILLO, NEGROS, SAMAR, BASILAN. Primary lowland forests, on undulating or flat land, especially near streams. Conservation status: Vulnerable (DAO 2017-11). Photos
  9. Dipterocarpus pseudocornutus P.S.Ashton, Gardens' Bulletin Singapore 67 (2015) 359; --Type: Gaerlan & Sagcal PPI 10209 (K, holo), Tawi-Tawi: Seratang Languyan. Endemic to the Philippines. TAWI-TAWI.
  10. Dipterocarpus validus Blume, Ann. Mus. Bot. Lugd.-Bat. 2 (1852) 36; --P.S.Ashton, Fl. Males. ser. 1, 9 (1982) 301. --Dipterocarpus affinis Brandis, J. Linn. Soc. Bot. 31 (1895) 31; --Merr., EPFP 3 (1923) 88. --Dipterocarpus lasiopodus Perkins, Frag. Fl. Philip, (1904) 22; --Merr., EPFP 3 (1923) 90. --Dipterocarpus warburgii Brandis, J. Linn. Soc. Bot. 31 (1895) 32; --Merr., EPFP 3 (1923) 91. --Dipterocarpus caudifera Merr., PJS 29 (1926) 398. --Dipterocarpus macrorhina van Slooten, Bull. Jard. Bot. Buitenz. III 8 (1927) 300. --Dipterocarpus woodii Merr., PJS 29 (1926) 398. Borneo (Tenom in N, and in E coast of Sabah SE-wards to River Barito), Philippines. LUZON: Cagayan, Isabela, Aurora, Nueva Ecija, Quezon, Camarines, MINDORO, CEBU, LEYTE, SAMAR, MINDANAO: Zamboanga, Davao, Agusan del Norte, Surigao. Common and frequently gregarious both in primary and secondary forests, low elevation up to 300m. There is some evidence of hybridization with Dipterocarpus kunstleri in the Philippines. Photos
Dubious species:
  1. Dipterocarpus orbicularis Foxw., PJS 13 c (1918) Bot. 180; --Merr., EPFP 3 (1923) 90; --P.S.Ashton, Fl. Males. ser. 1, 9 (1982) 326. Borneo, Philippines. LUZON: Camarines Norte. Lowland mixed dipterocarp forest. Resembles Dipterocarpus grandiflorus Blanco in fruit and Dipterocarpus gracilis Blume in the indumentum.

Hopea Roxb.

Last edited by Pieter B. Pelser, 18 August 2017

Photos
  1. Hopea acuminata Merr., Govt. Lab. Publ. (Philipp.) 29 (1905) 30; --EPFP 3 (1923) 93; --Gutierrez, Act. Manil. 4A (1968) 47; --P.S.Ashton, Fl. Males. ser. 1, 9 (1982) 420. --Hopea maquilingensis Foxw., PJS 13 (1918) 184. BABUYAN ISLS (CALAYAN), LUZON (widespread in most provinces), MINDORO, SAMAR, LEYTE, MINDANAO: Zamboanga, Misamis, Cotabato, Davao. Common in semi-evergreen and evergreen forests, 100(-300-800m). Conservation status: Endangered (DAO 2017-11). Photos
  2. Hopea basilanica Foxw., PJS 6 c (1911) Bot. 260; --Merr., EPFP 3 (1923) 93; --Gutierrez, Act. Manil. 4A (1968) 53; --P.S.Ashton, Fl. Males. ser. 1, 9 (1982) 425. MINDANAO, BASILAN, SIBUTU. Primary lowland forests on undulating hills below 70m. Conservation status: Critically Endangered (DAO 2017-11). Photos
  3. Hopea brachyptera (Foxw.) Slooten, Reinwardtia 3 (1956) 317; --P.S.Ashton, Fl. Males. ser. 1, 9 (1982) 415; --Balanocarpus brachypterus Foxw., PJS 13 c (1918) Bot. 195; --Merr., EPFP 3 (1923) 101. MINDANAO: Zamboanga. Primary lowland forests. Rare. Conservation status: Critically Endangered (DAO 2017-11).
  4. Hopea cagayanensis (Foxw.) Slooten, Reinwardtia 3 (1956) 318; --P.S.Ashton, Fl. Males. ser. 1, 9 (1982) 431; --Balanocarpus cagayanensis Foxw., PJS 13 c (1918) Bot. 194; PJS 67 (1938) 285; --Merr., EPFP 3 (1923) 101. LUZON: Apayao, Cagayan. Primary lowland semi-evergreen forests. Conservation status: Critically Endangered (DAO 2017-11). Photos
  5. Hopea foxworthyi Elmer, LPB 4 (1912) 1469; --Gutierrez, Act. Manil. 4A (1968) 32; --P.S.Ashton, Fl. Males. ser. 1, 9 (1982) 397; --Merr., EPFP 3 (1923) 94. --Hopea glutinosa Elmer, LPB 8 (1912) 1470; --Type: Elmer 12289 (MO, iso), Sibuyan. SIBUYAN. Locally common on red sticky volcanic soil along ridges at 600-700m, in seasonal semi-evergreen forest. Collections cited by Foxworthy and Merrill from islands other than Sibuyan belong to Hopea malibato. Conservation status: Critically Endangered (DAO 2017-11). Photos
  6. Hopea malibato Foxw., LPB 6 (1913) 1953; --EPFP 3 (1923) 93; --Gutierrez, Act. Manil. 4A (1968) 39; --P.S.Ashton, Fl. Males. ser. 1, 9 (1982) 403. --Hopea dalingdingan Gutierrez, Kalikasan 5 (1976) 92. --Hopea woodiana Gutierrez, Act. Manil. 4-A, 2 (1968) 42. LUZON: Quezon, NEGROS, MINDANAO: Agusan del Norte. Primary evergreen dipterocarp forests in aseasonal areas. Ascending up to 600m. Conservation status: Critically Endangered (DAO 2017-11). Photos
  7. Hopea mindanensis Foxw., PJS 6 c (1911) Bot. 261; --Merr., EPFP 3 (1923) 93; --Gutierrez, Act. Manil. 4A (1968) 63; --P.S.Ashton, Fl. Males. ser. 1, 9 (1982) 435. MINDANAO: Zamboanga. Primary lowland evergreen forests. Conservation status: Critically Endangered (DAO 2017-11). Photos
  8. Hopea philippinensis Dyer, J. Bot. 16 (1878) 100; --Merr., EPFP 3 (1923) 94; --Gutierrez, Act. Manil. 4A (1968) 65; --P.S.Ashton, Fl. Males. ser. 1, 9 (1982) 434. LUZON: Laguna, Quezon, Camarines, Albay, SAMAR, BILIRAN, LEYTE, PANAY, NEGROS, SAMAR, MINDANAO: Zamboanga, Lanao, Agusan. Primary evergreen forests on hills in aseasonal areas, ascending to 500m, often common. Conservation status: Critically Endangered (DAO 2017-11). Photos
  9. Hopea plagata (Blanco) S.Vidal, Sinopsis Atlas (1883) t. 15A; Rev. Pl. Vasc. Filip. (1886) 62; --Merr., EPFP 3 (1923) 94; --Gutierrez, Act. Manil. 4A (1968) 55; --P.S.Ashton, Fl. Males. ser. 1, 9 (1982) 423; --Mocanera plagata Blanco, Fl. Filip. (1837) 447; --Dipterocarpus plagatus (Blanco) Blanco, Fl. Filip. ed. 2 (1845) 311. NE Borneo, Philippines. LUZON: Cagayan, Nueva Vizcaya, Nueva Ecija, Pangasinan, Bulacan, Bataan, Rizal, Laguna, Quezon, Camarines, Sorsogon, MINDORO, TABLAS, BOHOL, MINDANAO, BASILAN, TAWI-TAWI. Common in semi-evergreen seasonal forests at low and medium elevation, often dominant. Conservation status: Vulnerable (DAO 2017-11). Photos
  10. Hopea quisumbingiana Gutierrez, Act. Manil. 4-A, 2 (1968) 31; --P.S.Ashton, Fl. Males. ser. 1, 9 (1982) 398; --Type: BS 1623 Ramos (GH, iso), Samar: Apr-1914. SAMAR. Low elevation everwet primary forest. Conservation status: Critically Endangered (DAO 2017-11). Photos
  11. Hopea reynosoi Gutierrez, Rojo & Madulid, Philippine Journal of Systematic Biology 4 (2010) 67; --Type: E.J. Reynoso & De la Cruz PNH125470 (PNH, holo), Samar: Eastern Samar, Balangiga, Sta. Rosa, Mar-1978. Endemic to Philippines. SAMAR. Conservation status: Critically Endangered (Gutierrez et al. 2010).
  12. Hopea rudiformis P.S.Ashton, Gard. Bull. Sing. 31 (1978) 30; Fl. Males. ser. 1, 9 (1982) 409; --Soejarto et al., Prelim. Checklist Fl. Pl. Palawan (1995). Borneo (SE and W), SW Philippines. PALAWAN (Narra, Marihuera, Narra-Aboabo Road, Soejarto et al. 6587, A, F, PNH). Conservation status: Vulnerable (DAO 2017-11).
  13. Hopea samarensis Gutierrez, Kalikasan 4 (1975) 236; --P.S.Ashton, Fl. Males. ser. 1, 9 (1982) 427. SAMAR. Mixed dipterocarp forests in moist lowland valleys. Conservation status: Critically Endangered (DAO 2017-11).
  14. Hopea santosiana Gutierrez, Rojo & Madulid, Philippine Journal of Systematic Biology 4 (2010) 69; --Type: E.J. Reynoso & De la Cruz, PNH125468 (PNH, holo), Samar: Eastern Samar, Balangiga, Sta. Rosa, Mar-1978. Endemic to Philippines. SAMAR. Conservation status: Critically Endangered (Gutierrez et al. 2010).

 Isoptera Scheffer = Shorea

Parashorea Kurz

Photos
  1. Parashorea malaanonan (Blanco) Merr., Spec. Blancoanae (1918) 271; --Merr., EPFP 3 (1923) 100; --P.S.Ashton, Fl. Males. ser. 1, 9 (1982) 383; --Mocanera malaanonan Blanco, Fl. Filip. (1837) 858; --Dipterocarpus malaanonan Blanco, Fl. Filip. ed. 2 (1845) 312. Borneo (Brunei to Sabah, SE to Berau and Nunukan), Philippines. LUZON: Nueva Ecija, Bulacan, Rizal, Laguna, Quezon, Camarines, Albay, Sorsogon, POLILLO, CATANDUANES, MASBATE, PANAY, NEGROS, CEBU, LEYTE, SAMAR, MINDANAO. Primary lowland forests, common. Photos
  2. Parashorea warburgii Brandis, J. Linn. Soc. Bot. 31 (1895) 105; --Merr., EPFP 3 (1923) 100; --P.S.Ashton, Fl. Males. ser. 1, 9 (1982) 387. --Parashorea tomentella (Symington) Meijer, Acta Bot. Neerl. 12 (1963) 320. MINDANAO: Agusan del Norte, Davao. Primary lowland forests.

Pentacme A.DC. = Shorea sect. Pentacme (A.DC) P.S.Ashton

Shorea Roxb. ex C.F.Gaertn.

Last edited by Pieter B. Pelser, 27 October 2017

Photos
  1. Shorea almon Foxw., PJS 67 (1938) 313; --P.S.Ashton, Fl. Males. ser. 1, 9 (1982) 507. --Shorea eximia auct. non (Miq.) Scheff.; --Merr., EPFP 3 (1923) 96. NE Borneo (Sarawak, Sabah, NE Kalimantan), Philippines. LUZON: Bataan, Quezon, Camarines, Albay, Sorsogon, NEGROS, SAMAR, MINDANAO: Zamboanga, Misamis, Lanao, Agusan, Surigao, BASILAN. Primary forest on undulating hills at low elevation, on clay soil. According to Ashton (1982), particularly variable in Mindanao where the leaves are much smaller and the tomentum more sparse and scabrid in some trees. Conservation status: Vulnerable (DAO 2017-11). Photos
  2. Shorea arsorianoi Gutierrez, Rojo & Madulid, Philippine Journal of Systematic Biology 4 (2010) 70; --Type: Gutierrez, Reynoso & Aga, PNH123122, (PNH, holo), Mindanao: Surigao del Sur, Bislig, PICOP, Apr-1977. --Shorea virescens auct. non Parijs; --P.S.Ashton, Fl. Mal. Prec. Dipt. Gard. Bull. Sing. 31 (1978) 44; --Rojo, Kalikasan, Philipp. J. Biol. 5 (1976) 99. Endemic to Philippines. MINDANAO: Davo City, Davao del Norte, Surigao del Sur. Conservation status: Endangered (Gutierrez et al. 2010).
  3. Shorea assamica Dyer, Fl. Br. Ind. 1 (1874) 307; --P.S.Ashton, Gard. Bull. Sing. 31 (1978) 43; Fl. Males. ser. 1, 9 (1982) 491.
  4. Shorea astylosa Foxw., PJS 13 c (1918) Bot. 188, pl. 5; PJS 67 (1938) 297; --EPFP 3 (1923) 96; --P.S.Ashton, Fl. Males. ser. 1, 9 (1982) 460. LUZON: Quezon, Camarines, NEGROS, BILIRAN, SAMAR, MINDANAO: Zamboanga, Davao, Agusan. Local in primary lowland evergreen dipterocarp forests. Conservation status: Critically Endangered (DAO 2017-11).
  5. Shorea contorta S.Vidal, Sinopsis Atlas (1883) 15; Rev. Pl. Vasc. Filip. (1886) 88; --Merr., EPFP 3 (1923) 95; --P.S.Ashton, Fl. Males. ser. 1, 9 (1982) 467; --Pentacme contorta (S.Vidal) Merr. & Rolfe, PJS 3 c (1908) Bot. 115; --Merr., EPFP 3 (1923) 95; --Foxw., PJS 67 (1938) 287. --Pentacme mindanensis Foxw., PJS 13 c (1918) Bot. 185; PJS 67 (1938) 289; --Merr., EPFP 3 (1923) 95; --Pentacme paucinervis Brandis, Journ. Linn. Soc. Bot. 31 (1895) 73. BABUYAN ISLS (CALAYAN), LUZON (in most provinces), POLILLO, MINDORO, SIBUYAN, MARINDUQUE, MASBATE, NEGROS, LEYTE, SAMAR, MINDANAO: Zamboanga, Lanao, Agusan, BASILAN. Common in primary lowland forests from semi-seasonal to everwet areas, in the former often semi-gregarious. Conservation status: Vulnerable (DAO 2017-11). Photos
  6. Shorea falciferoides Foxw., PJS 13 c (1918) Bot. 189; PJS 67 (1938) 296; --P.S.Ashton, Fl. Males. ser. 1, 9 (1982) 458. --Shorea balangeran Dyer ex S.Vidal, Phan. Cuming. Philipp. (1885) 97, nom. illeg., nom. nud.; --Merr., EPFP 3 (1923) 98; --P.S.Ashton, Fl. Males. ser. 1, 9 (1982) 458. --Shorea gisok Foxw., PJS 67 (1938) 294. LUZON: Pangasinan, Zambales, Bulacan, Quezon, Camarines, Albay, SAMAR, CEBU, SIBUYAN, LEYTE, MINDANAO: Zamboanga, Agusan, BASILAN. Mixed dipterocarp forests to 1000m, in moderately seasonal to everwet areas. According to Ashton (1982), specimens from the more seasonal NE Luzon (Shorea falciferoides) have more or less smaller leaves than other Philippine collections (formerly named Shorea gisok). The latter is more or less confined to ridgetops in the everwet areas. Conservation status: Vulnerable (DAO 2017-11). Photos
  7. Shorea guiso (Blanco) Blume, Ann. Mus. Bot. Lugd.-Bat. 2 (1852) 34; --Merr., EPFP 3 (1923) 97; --Foxw., PJS 67 (1938) 292; --P.S.Ashton, Fl. Males. ser. 1, 9 (1982) 447; --Mocanera guiso Blanco, Fl. Filip. (1837) 449; --Dipterocarpus guiso (Blanco) Blanco, Fl. Filip., ed. 2 (1845) 313. Peninsular and SE Thailand, S Vietnam, Peninsular Malaysia, Sumatra (Aceh, Tapanuli, Palembang), Borneo (mainly in E), Philippines. LUZON (widespread in most provinces), MINDORO, MASBATE, PANAY, NEGROS, LEYTE, SAMAR, MINDANAO, BASILAN. Primary lowland forests on red soils, most common in slightly seasonal areas. Photos
  8. Shorea hopeifolia (Heim) Sym., Gard. Bull. S.S. 8 (1933) 150; --P.S.Ashton, Fl. Males. ser. 1, 9 (1982) 482; --Cotylelobium hopeifolia Heim, Bull. Mens. Soc. Linn. Paris I (1891) 971. --Shorea kalunti Merr., PJS 26 (1925) 475; EPFP 4 (1925) 250; --Foxw., PJS 67 (1938) 307. Sumatra, Peninsular Malaysia (seasonal areas excepted), Borneo, Philippines. MINDANAO: Zamboanga, Lanao, Davao. Primary forests on fertile clay rich soil on undulating terrain and hills below 600m.
  9. Shorea malibato Foxw. in Elmer, LPB 6 (1913) 1955; --Merr., EPFP 3 (1923) 97; --Foxw., PJS 67 (1938) 298; --P.S.Ashton, Fl. Males. ser. 1, 9 (1982) 460. LUZON: Quezon, Camarines, LEYTE, MINDANAO: Zamboanga, Agusan del Norte. Local in lowland primary forests in aseasonal areas, c. 450m in Agusan del Norte. Conservation status: Critically Endangered (DAO 2017-11).
  10. Shorea negrosensis Foxw, PJS 6 c (1911) Bot. 274; PJS 67 (1938) 315; --Merr., EPFP 3 (1923) 96; --P.S.Ashton, Fl. Males. ser. 1, 9 (1982) 502. LUZON: Cagayan, Isabela, Aurora, Nueva Ecija, Laguna, Quezon, Camarines, Albay, Sorsogon, POLILLO, NEGROS, CEBU, LEYTE, BILIRAN, SAMAR, MINDANAO: Zamboanga, Lanao, Cotabato, Bukidnon, Davao, Surigao, Agusan, BASILAN. Primary evergreen and semi-evergreen lowland forests, common and often gregarious. Conservation status: Vulnerable (DAO 2017-11).
  11. Shorea ovata Dyer ex Brandis, J. Linn. Soc. Bot. 31 (1895) 91; --P.S.Ashton, Fl. Males. ser. 1, 9 (1982) 545. --Shorea agsaboensis W.L.Stearn, Brittonia 17 (1965) 36; --Rojo, Pterocarpus 3 (1977) 70. Sumatra (W coast), Peninsular Malaysia, Borneo (Kapuas hinterland, Sarawak to SW Sabah, SE Borneo), Philippines. MINDANAO: Misamis Oriental, Agusan del Norte (Tungao), Primary forests at medium elevations to c. 900m. Conservation status: Endangered (DAO 2017-11). Photos
  12. Shorea squamata (Turcz.) Benth. & Hook.f. ex DC., Prodr. 16, 2 (1868) 632; --Foxw., PJS 67 (1938) 310; --Hopea squamata Turcz., Bull. Soc. Mosc. 31 2 (1858) 239. --Shorea palosapis (Blanco) Merr., Sp. Blancoanae (1918) 271; EPFP 3 (1923) 98; --P.S.Ashton, Fl. Males. ser. 1, 9 (1982) 517; --Dipterocarpus palosapis Blanco, Fl. Filip., ed. 2 (1845) 312. LUZON: Ilocos Norte, Cagayan, Isabela, Nueva Ecija, Bulacan, Rizal, Laguna, Quezon, Camarines, Albay, Sorsogon, POLILLO, MINDORO, MARINDUQUE, LEYTE, SAMAR, MINDANAO: Zamboanga, Misamis, Bukidnon, Davao, Agusan, Surigao, BASILAN. Primary lowland evergreen forests on fertile well-drained soil. In Mindanao ascending to 1000m. In some regions dominant. Photos
  13. Shorea polita S.Vidal, Sinopsis Atlas (1883) 15; --Merr., EPFP 3 (1923) 98; --Foxw., PJS 67 (1938) 304; --P.S.Ashton, Fl. Males. ser. 1, 9 (1982) 487. --Shorea mindanensis Foxw., PJS 13 c (1918) Bot. 192; --Merr., EPFP 3 (1923) 97. LUZON: Cagayan, Ilocos Sur, Pangasinan, Zambales, Rizal, Nueva Ecija, Quezon, MINDORO, MINDANAO: Zamboanga, Davao, BASILAN. Apparently scattered in primary semi-evergreen and evergreen lowland forests.
  14. Shorea polysperma (Blanco) Merr., Govt. Lab. Publ. (Philipp.) 27 (1905) 22; --Merr., EPFP 3 (1923) 99; --Foxw., PJS 67 (1938) 309; --P.S.Ashton, Fl. Males. ser. 1, 9 (1982) 514; --Mocanera polysperma Blanco, Fl. Filip. (1837) 448; --Dipterocarpus polysperma (Blanco) Blanco, Fl. Filip., ed. 2 (1846) 312. --Shorea warburgii Gilg, Engl. Bot. Jahrb. 18 Beibl. 45 (1894) 38; --Type: Warburgh 12399 (K), Cagayan: Penablanca. --Shorea teysmanniana auct. non Dyer ex Brandis; --Foxw., PJS 6 c (1911) Bot. 279; --Merr., EPFP 3 (1923) 99. BABUYAN ISLS (CALAYAN), LUZON: Cagayan, Ilocos Norte, Pangasinan, Nueva Ecija, Bulacan, Zambales, Bataan, Laguna, Quezon, Camarines, Albay, Sorsogon, POLILLO, MINDORO, PANAY, NEGROS, CEBU, BILIRAN, LEYTE, SAMAR, MINDANAO: Zamboanga, Misamis, Lanao, Cotabato, Bukidnon, Davao, Agusan, Surigao, BASILAN. Often common in lowland and medium elevation primary forests, ascending to 1000m (exceptionally to 1200m). Conservation status: Vulnerable (DAO 2017-11). Photos
  15. Shorea seminis (Vriese) Slooten in Merr., Pl. Elm. Born. (1929) 204; --P.S.Ashton, Fl. Males. ser. 1, 9 (1982) 451; --Hopea seminis de Vriese, Minjak Tangkawang (1861) 32. --Isoptera borneensis Scheff. ex Burck, Ann. Jard. Bot. Buitenz. 6 (1887) 222; --Merr., EPFP 3 (1923) 101. Borneo, S Philippines (Ashton 1982). Merrill (1923) also cites Malay Peninsula and Banka. MINDANAO: Zamboanga. Primary lowland forests. Conservation status: Vulnerable (DAO 2017-11).
  16. Shorea virescens Parijs, Fedde Rep. 33 (1933) 244; --Rojo, Kalikasan 5 (1976) 99; --P.S.Ashton, Fl. Males. ser. 1, 9 (1982) 492. Borneo, Philippines. SAMAR, MINDANAO: Davao del Norte, Surigao del Sur. Flat and undulating terrains and low hills below 500m.

Vatica L.

Last edited by Pieter B. Pelser, 27 October 2017

Photos
  1. Vatica elliptica Foxw., PJS 67 (1938) 329; --P.S.Ashton, Fl. Males. ser. 1, 9 (1982) 369. MINDANAO: Zamboanga, twice collected. Mature fruit not known. Conservation status: Critically Endangered (DAO 2017-11).
  2. Vatica mangachapoi Blanco, Fl. Filip. (1837) 401; --Merr., EPFP 3 (1923) 101; --P.S.Ashton, Fl. Males. ser. 1, 9 (1982) 364.
  3. Vatica maritima Slooten, Bull. Bot. Gard. Buitenz. 3, 17 (1942) 245; --P.S.Ashton, Fl. Males. ser. 1, 9 (1982) 359. Borneo (NE Kalimantan; Brunei; W Sabah), S Philippines. PALAWAN (Taytay, NE foot of Mt Capoas). Conservation status: Endangered (DAO 2017-11).
  4. Vatica mendozae Gutierrez, Rojo & Madulid, Philippine Journal of Systematic Biology 4 (2010) 72; --Type: J.P. Rojo PNH121551 (PNH, holo; CLP, iso), Quezon: Real, Kinalumbakan, 26-May-1972. Endemic to Philippines. LUZON: Quezon. Conservation status: Endangered (Gutierrez et al. 2010).
  5. Vatica odorata (Griff.) Sym., J. Mal. Br. R. As. Soc. 19 (1941) 156; --P.S.Ashton, Gard. Bull. Sing. 22 (1967) 263; Gard. Bull. Sing. 31 (1978) 23; Fl. Males. ser. 1, 9 (1982) 360; --Sunaptea odorata Griff., Not. Pl. Asiat. 4 (1854) 516.
  6. Vatica pachyphylla Merr., PJS 13 c (1918) Bot. 311; --Merr., EPFP 3 (1923) 102; --P.S.Ashton, Fl. Males. ser. 1, 9 (1982) 363; --Type FB 27102 Alhambra (PNH, holo, ?lost; photo in Foxworthy 1938), Luzon: Camarines Norte prov., Paracale. LUZON: Cagayan; Quezon, Camarines Norte, POLILLO. Scattered in aseasonal, primary dipterocarp forests below 80m. Conservation status: Critically Endangered (DAO 2017-11).
  7. Vatica rassak Blume, Ann. Mus. Bot. Lugd.-Bat. (1852) 31, incl. var. subcordata Blume; --P.S.Ashton, Gard. Bull. Sing. 31 (1978) 20; Fl. Males. ser. 1, 9 (1982) 353. Borneo (E of Rejang Valley & Sampit, commonest on E coast), SW Philippines, Sulawesi, Moluccas (Sula Isls, Morotai, Halmahera, Bacan, Obi Is., Aru Isls), New Guinea, Sudest Is. TAWI-TAWI.
  8. Vatica umbonata (Hook.f.) Burck, Ann. Jard. Bot. Buitenz. 6 (1887) 232; --P.S.Ashton, Fl. Males. ser. 1, 9 (1982) 349; --Pachynocarpus umbonatus Hook.f., Trans. Linn. Soc. 23 (1860) 159.

Acknowledgements

Jiro Adorador kindly provided some of the photos on this webpage.

Literature

Ashton, PS. 1982. Dipterocarpaceae. Flora Malesiana series 1, 9: 237-552

Ashton, PS. 2015. A few last words on Far Eastern Dipterocarps. Gardens' Bulletin Singapore 67: 357-360


DENR Administrative Order. 2017. Updated national list of threatened Philippine plants and their categories. DAO 2017-11

Foxworthy, FW. 1911. Philippine Dipterocarpaceae. Philippine Journal of Science 6 (section C) Botany: 231-285, plate 34-44.

Foxworthy, FW. 1919. Philippine Dipterocarpaceae. Philippine Journal of Science 13: 163-199, plate 1, 2.

Foxworthy, FW. 1938. Philippine Dipterocarpaceae. Philippine Journal of Science 67: 241-333, plate 1-9.

Gutierrez, HG. 1968. Acta Manillana 4A.

Gutierrez, HG, JP Rojo & DA Madulid. 2010. New dipterocarp species from the Philippines. Philippine Journal of Systematic Biology 4: 67-78.

Pelser, P.B., J.F. Barcelona & D.L. Nickrent (eds.). 2011 onwards. Co's Digital Flora of the Philippines. www.philippineplants.org

Copyright 2011, Co's Digital Flora of the Philippines

Last updated 27 October 2017