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Primulaceae

Including Aegicerataceae, Maesaceae, Myrsinaceae

Androsace L. = Primula

Aegiceras Gaertn.

Last edited by Pieter B. Pelser, 20 September 2017

Photos
  1. Aegiceras corniculatum (L.) Blanco, Fl. Filip. (1837) 79; --Merr., Fl. Manila (1912) 361; EPFP 3 (1923) 255; --Rhizophora corniculata L., Herb. Amb. (1754) 13. Photos
  2. Aegiceras floridum Roem. & Schult., Syst. 4 (1819) 512; --Merr., EPFP 3 (1923) 256. Photos

Amblyanthopsis Merr. (1923), non Mez = Loheria, Discocalyx

Ardisia Sw.

Last edited by Pieter B. Pelser, 22 September 2017

Photos
  1. Ardisia alvarezii Merr., PJS 26 (1925) 484; EPFP 4 (1925) 250. BABUYAN ISLS (CALAYAN), LUZON (Cagayan, Apayao). Forests at low elevations.
  2. Ardisia basilanensis Merr., PJS 12 c (1917) Bot. 153; --Merr., EPFP 3 (1923) 256. BASILAN, MINDANAO: Zamboanga. Primary forests at low and medium elevation.
  3. Ardisia brevipetiolata (Merr.) Merr., EPFP 3 (1923) 256; --Ardisia serrata var. brevipetiolata Merr., PJS 2 c (1907) Bot. 296. --Type: Merrill 4049, Mindoro: Mindoro Oriental prov., Baco River near base of Mt Halcon. --Ardisia sibuyanensis Elmer, LPB 4 (1912) 1497; --Merr., EPFP 3 (1923) 263. MINDORO: Mindoro Oriental, SIBUYAN: Mt Giting-giting. Primary forests, also reported along streams, 225-550m. A. sibuyanensis is a form with entire leaves.
  4. Ardisia calavitensis Merr., PJS 20 (1922) 423; --Merr., EPFP 3 (1923) 256. MINDORO. Primary forests at low and medium elevation.
  5. Ardisia castaneifolia Mez in Engl., Pflanzenreich 9 (1902) 138; --Merr., EPFP 3 (1923) 257. LUZON: Bulacan. Probably in forests at low and medium elevation.
  6. Ardisia confertiflora Merr., PJS 5 c (1910) Bot. 213; --Merr., EPFP 3 (1923) 257. BATAN, FUGA, MINDORO. Low elevation thickets. Photos
  7. Ardisia copelandii Mez, PJS 1 (1906) Suppl. 272; --Merr., EPFP 3 (1923) 257. Borneo, Philippines. MINDANAO: Lanao, Bukidnon, Davao. Primary forests, 600-1000m.
  8. Ardisia crispa (Thunb.) A.DC. in Trans. Linn. Soc. 17 (1834) 124; --Merr., EPFP 3 (1923) 257; --Bladhia crispa Thunb., Syst. Veg. ed. 14 (1784) 237. India, S China, Vietnam, Taiwan, Ryukyus, Korea, and Japan, southward to Borneo, Sumatra, Java, Philippines. LUZON: Ilocos Norte, Mountain Province, Benguet, Nueva Vizcaya, Pangasinan, Zambales. Chiefly in the mossy forest, 800-2200m.
  9. Ardisia cumingiana A.DC., Prod. 8  (1844). 136; --Merr., EPFP 3 (1923) 257. LUZON: Ilocos Sur, Benguet, Cagayan, Batangas. Ravines, up to 1500m.
  10. Ardisia curranii Merr., PJS 5 c (1910) Bot. 214; --Merr., EPFP 3 (1923) 258. LUZON: Camarines. Primary forests at low elevation.
  11. Ardisia curtipes Merr., PJS 5 c (1910) Bot. 372; EPFP 3 (1923) 258. --Ardisia dataensis Mez, Fedde Repert. 16 (1920) 414. LUZON: Ifugao, Mountain Province, Benguet. Mossy forests, 1600-2400m.
  12. Ardisia curtiflora Elmer, LPB 8 (1915) 2778; --Merr., EPFP 3 (1923) 258. MINDANAO: Agusan del Norte (Mt Urdaneta). Mossy forests, c. 1600m.
  13. Ardisia cymosa Blume, Bijdr. (1826) 689; --C.M.Hu, Blumea 44 (1999) 393. --Ardisia cagayanensis Merr., PJS 7 c (1912) Bot. 323; --Merr., EPFP 3 (1923) 256. LUZON: Cagayan, CATANDUANES. Primary forests at low elevation (as A cagayanensis Merr.).
  14. Ardisia darlingii Merr., PJS 5 c (1910) Bot. 215; EPFP 3 (1923) 258; --C.M.Hu, Blumea 44 (1999) 398. --Ardisia gitingensis Elmer, LPB 4 (1912) 1498; --Merr., EPFP 3 (1923) 258.
  15. Ardisia diffusa Merr., PJS 5 c (1910) Bot. 215; EPFP 3 (1923) 258. MINDANAO: Misamis Occidental (Mt Malindang). Forests, probably at medium elevation.
  16. Ardisia elmeri Mez, PJS 1 (1906) Suppl. 273; --Merr., EPFP 3 (1923) 258; --Type: Elmer 6493 (MO, NY, US, iso), Luzon: Benguet prov., Baguio, Jun-1904. --Ardisia fragrans Elmer, LPB 2 (1910) 664; --Merr., EPFP 3 (1923) 258; --Type: Elmer 11647 (US, lecto; BISH, GH, L, MO, NA, isolecto), Mindanao: Davao, Mt Apo, Todaya, Sep-1909. LUZON: Mountain Province, Benguet (Baguio), Rizal, Sorsogon, MINDANAO: Davao (Mt Apo). Humid lower and midmontane forests, damp ravines, 1200-1600m.
  17. Ardisia geissanthoides Mez, Fedde Repert. 16 (1920) 412; --Merr., EPFP 3 (1923) 258. MINDORO: Mindoro Oriental (Mt Halcon). Mossy forest, 1800-2000m.
  18. Ardisia ilocana Merr., PJS 14 (1919) 443; EPFP 3 (1923) 259. LUZON: Ilocos Norte (Bangui to Claveria). Forests, c. 500m.
  19. Ardisia iwahigensis Elmer, LPB 5 (1913) 1817; --Merr., EPFP 3 (1923) 259. Endemic to the Philippines. PALAWAN: Puerto Princesa (Mt Pulgar = Thumb Peak). Damp primary forests, c. 225m. Conservation status: Vulnerable (DAO 2017-11).
  20. Ardisia keithleyi Merr., PJS 10 c (1915) Bot. 335; EPFP 3 (1923) 259. MINDANAO: Zamboanga, Lanao del Sur (between Malabang and Camp Keithley). Primary forests, c. 650m.
  21. Ardisia lanceolata Roxb., Fl. Ind. ed. Carey 2 (1824) 275; --Hort. Beng. (1814) 85, nom. nud.; --Merr., EPFP 3 (1923) 259; --C.M.Hu, Blumea 44 (1999) 401. --Ardisia clementis Elmer, LPB 2 (1910) 665; --Merr., EPFP 3 (1923) 257. --Ardisia lanaensis Mez, Fedde Repert. 16 (1920) 411. Sumatra, Peninsular Malaysia, Java, Borneo, Philippines, Sulawesi. BASILAN, MINDANAO: Zamboanga, Lanao, Davao (Mt Apo). Primary forests, from the lowlands (sometimes near the seashore) to 1100m.
  22. Ardisia laxiflora Merr., PJS 12 c (1917) Bot. 153; EPFP 3 (1923) 259. MINDANAO: Lanao. Primary forests at low elevation.
  23. Ardisia loheri Merr., PJS 12 c (1917) Bot. 152; EPFP 3 (1923) 259. LUZON: Rizal, Laguna, MINDANAO: Bukidnon. Primary forests at medium elevation, ascending to 1200m.
  24. Ardisia longipetiolata Merr., PJS 14 (1919) 444; EPFP 3 (1923) 259. LUZON: Camarines Norte (Paracale). Damp primary forests at low elevation.
  25. Ardisia macgregorii Merr., PJS 5 c (1910) Bot. 217; EPFP 3 (1923) 260; --Type: BS 1722 McGregor (US, lecto), Cebu: near Toledo. LUZON: Ilocos Norte, CEBU, SAMAR. In thickets and forests at low and medium elevation.
  26. Ardisia marginata Blume, Bijdr. (1826) 688. --Ardisia apoensis Elmer, LPB 2 (1910) 663; --Merr., EPFP 3 (1923) 256. Java, Philippines. LUZON: Abra, Mountain Province, Benguet, Cagayan, Nueva Ecija, Rizal, Laguna, Quezon, Camarines, BILIRAN, LEYTE, CAMIGUIN DE MINDANAO, MINDANAO: Davao (Mt Apo). Primary forests at low and medium elevation, also on forested ridges to 1200m.
  27. Ardisia mezii Elmer, LPB 2 (1908) 440; --Merr., EPFP 3 (1923) 260. LUZON: Tayabas, Laguna, Sorsogon, MINDANAO: Bukidnon. Primary forests, 700-1200m.
  28. Ardisia mindanaensis Mez in Engl., Pflanzenreich 9 (1902) 102; --Merr., EPFP 3 (1923) 260. LEYTE, MINDANAO: Lanao, Bukidnon, Davao. Primary forests, 500-1000m.
  29. Ardisia mirandae Merr., PJS 12 c (1917) Bot. 154; EPFP 3 (1923) 260. LUZON: Camarines. Exposed peaks and ridges in the mossy forest, c. 700m.
  30. Ardisia nigro-maculata Merr., PJS 13 c (1918) Bot. 49; EPFP 3 (1923) 260. LUZON: Quezon (Mt Binuang), PANAY. Mossy forests, c. 1000m.
  31. Ardisia oreophila C.M.Hu, Blumea 47 (2002) 496; --Type: Soejarto, Fernando & Sagcal 8828 (L, holo), Palawan: Brookes Point, Mt Mantalingahan. Ridges and slopes of Sindurapi Peak, 1100m. PALAWAN. Known only from the type collection.
  32. Ardisia palawanensis Merr., PJS 5 c (1910) Bot. 220; EPFP 3 (1923) 261. PALAWAN (c. 4.8km NE of Puerto Princesa). Undershrub in lowland flat forest.
  33. Ardisia polysticta Miq., Fl. Ind. Bat. Suppl. 1 (1861) 576; --C.M.Hu, Blumea 44 (1999) 395. --Ardisia jagorii Mez in Engl., Pflanzenreich 9 (1902) 148; --Merr., EPFP 3 (1923) 259. --Ardisia pardalina Mez in Engl., Pflanzenreich 9 (1902) 148; --Merr., EPFP 3 (1923) 261. --Ardisia ramosii Merr., PJS 7 (1912) 148; --C.M.Hu, Blumea 44 (1999) 395. --Ardisia sinuato-crenata Mez in Engl., Pflanzenreich 9 (1902) 148; --Merr., EPFP 3 (1923) 263. LUZON: Ilocos Norte, Mountain Province, Benguet, Cagayan, Nueva Vizcaya, Bataan, Rizal, Laguna, Camarines, Albay, Sorsogon, MINDORO, PANAY, BILIRAN, LEYTE, SAMAR, MINDANAO. Primary forests from low to high elevation, in mossy forest ascending to 2300m (as Ardisia pardalina and Ardisia ramosii).
  34. Ardisia pulchella Mez, Fedde Repert. 16 (1920) 410; --Merr., EPFP 3 (1923) 261. LUZON: Mountain Province, Benguet. Mossy forests, 2000-2400m.
  35. Ardisia pusilla A.DC., Trans. Linn. Soc. 17 (1836) 126. --Ardisia reptans Merr., PJS 5 c (1910) Bot. 220; EPFP 3 (1923) 262. Japan, Ryukyus, Korea, China, Taiwan, Thailand, Philippines. LUZON: Pampanga (Mt Abu, along Zambales Range). Forested ravines, c. 1350m.
  36. Ardisia pyramidalis (Cav.) Pers., Syn. 1 (1805) 233; --Merr., EPFP 3 (1923) 261. Borneo, Philippines. LUZON: Cagayan to Albay, PALAWAN, MINDANAO, in most islands and provinces. Primary forests at low and medium elevation, often common. Photos
  37. Ardisia racemosopaniculata Mez, PJS 1 (1906) Suppl. 273; --Merr., EPFP 3 (1923) 262. MINDANAO: Davao and North Cotabato (Mt Apo). Primary forests, c. 1600m.
  38. Ardisia reynosoi Stone, Sida 16 (1994) 263; --Type: Argent & Reynoso 89163 (AAU, iso), Sibuyan: Mt Giting-giting, above Magdiwang on ridge leading to Mayo’s Peak, ‘grassy’ area above Camp 3, 1400m. SIBUYAN. Known only from the type collection.
  39. Ardisia rivularis Merr., PJS 13 c (1918) Bot. 50; EPFP 3 (1923) 262. LUZON: Quezon (Umiray, along Umiray River), in forests at low and medium elevation.
  40. Ardisia romanii Elmer, LPB 5 (1913) 1820; --Merr., EPFP 3 (1923) 262. Endemic to the Philippines. PALAWAN. Forests at low elevation. Conservation status: Vulnerable (DAO 2017-11). Photos
  41. Ardisia saligna Mez in Engl., Pflanzenreich 9 (1902) 143; --Merr., EPFP 3 (1923) 262. --Ardisia macropus Mez, Fedde Repert. 16 (1920) 415; EPFP 3 (1923) 260. LUZON: Ilocos Norte, Abra, Mountain Province, Benguet, Tayabas, Nueva Ecija, Bataan, Bulacan, Rizal, Laguna, Cavite, POLILLO. Primary forests at low and medium elevation, ascending to 1000m.
  42. Ardisia samarensis Merr., PJS 12 c (1917) Bot. 151; EPFP 3 (1923) 263. SAMAR (Pinipisakan). Damp primary forests at low elevation.
  43. Ardisia scabrida Mez in Engl., Pflanzenreich 9 (1902) 118; --Merr., EPFP 3 (1923) 263. --Ardisia punctata auct. non Lindl.; --Elmer, LPB 2 (1908) 439; --Type: Elmer 10385 (A, iso?), Negros: Negros Oriental prov., Cuernos Mtns, Jun-1908. NEGROS: Negros Oriental (Cuernos Mtns), CAMIGUIN, MINDANAO: Misamis, Bukidnon, Davao (Mt Apo). Forests, 600-1300m.
  44. Ardisia scalaris Mez in Engl., Pflanzenreich 9 (1902) 142; --Merr., EPFP 3 (1923) 263. LUZON: Rizal, Quezon. Primary forests along streams at low elevation. According to Merrill (1923), very similar to Ardisia saligna Mez.
  45. Ardisia serrata (Cav.) Pers., Syn. 1 (1805) 233; --Merr., EPFP 3 (1923) 263; --C.M.Hu, Blumea 44 (1999) 398; --Anguillaria serrata Cav., Icon. 6 (1800) 2. --Ardisia angustifolia A.DC., Prodr. 8 (1844) 134; --Merr., EPFP 3 (1923) 256. --Ardisia abanii Stone, Proc. Acad. Nat. Sci. Philadelphia 142 (1990) 53. --Ardisia mindorensis Merr., PJS 5 c (1910) Bot. 218; EPFP 3 (1923) 260. --Ardisia oligocarpa Merr., J. Arn. Arb. 35 (1954) 148. --Ardisia oligantha auct. non Baker; --Elmer, LPB 4 (1912) 146; --Merr., EPFP 3 (1923) 260. --Ardisia whitfordii Mez, PJS 1 (1906) Suppl. 271; --Merr., EPFP 3 (1923) 265. --Ardisia grandidens Mez in Engl., Pflanzenreich 9 (1902) 137; --Merr., EPFP 3 (1923) 259. Borneo, Philippines. LUZON: Apayao, Ifugao, Mountain Province, Benguet, Cagayan, Nueva Ecija, Rizal, Quezon, POLILLO, MINDORO, PALAWAN, SIBUYAN, PANAY, BILIRAN, LEYTE, SAMAR, MINDANAO. Primary forests from lowland to mossy, on ridges, to 2400m. Data compiled from distribution range of above-cited synonyms, excepting Ardisia abanii Stone and Ardisia oligocarpa Merr..
  46. Ardisia squamulosa C.Presl, Rel. Haenk. 2 (1830) 65; --Merr., EPFP 3 (1923) 263. --Ardisia pirifolia Mez in Engl., Pflanzenreich 9 (1902) 129; --Merr., EPFP 3 (1923) 261. --Ardisia verrucosa C.Presl, Reliq. Haenk. 2 (1835) 66; --Merr., EPFP 3 (1923) 265. ?Endemic to the Philippines. According to Merrill (1923), a tree rather than a shrub, not certainly distinct from Ardisia obovata Blume. LUZON: Ilocos Norte, Cagayan, Isabela, Pangasinan, Tayabas, Cavite, Batangas, MINDORO, PALAWAN, PANAY, MINDANAO: Davao. In most or all islands and provinces. Common in primary forests at low and medium elevation, ascending to 1000m. Note by L.L.Co: Ardisia verrucosa is hardly distinguishable from Ardisia squamulosa. Conservation status: Vulnerable (DAO 2017-11).
  47. Ardisia stichantha Stone, Pacific Sci. 47 (1993) 290; --Type: BS 29315 Ramos & Edaño (US, holo), Luzon: Rizal prov., Mt Susong-Dalaga. Aug-1917. LUZON: Rizal.
  48. Ardisia sulcata Mez in Engl., Pflanzenreich 9 (1902) 112; --Merr., EPFP 3 (1923) 264; --Type: Warburg 12286. LUZON: Apayao, Cagayan, Tayabas, Camarines, BILIRAN, LEYTE, SAMAR. Damp primary forests at low elevation.
  49. Ardisia tayabensis Merr., PJS 12 c (1917) Bot. 156; EPFP 3 (1923) 264. LUZON: Aurora (Mt Dingalan). Primary forests at or above 300m.
  50. Ardisia taytayensis Merr., J. Arnold Arbor. 35 (1954) 148. --Ardisia pachyphylla auct. non Dunn; --Merr., PJS 12 c (1917) Bot. 154; EPFP 3 (1923) 260. Endemic to the Philippines. PALAWAN, BALABAC. Dry thickets and borders of forests at low elevation. Conservation status: Vulnerable (DAO 2017-11).
  51. Ardisia tomentosa C.Presl, Reliq. Haenk. 2 (1835) 66; --Merr., EPFP 3 (1923) 265; --C.M.Hu, Blumea 44 (1999) 397. --Ardisia philippinensis A.DC., Prodr. 8 (1844) 128; --Merr., EPFP 3 (1923) 261; --Type: Cuming 1467. --Ardisia cuprea Elmer, LPB 8 (1915) 2777; --Merr., EPFP 3 (1923) 257; --Type: Elmer 13287 (C, L, NY 2 sheets, iso), Mindanao: Agusan del Norte prov., Mt Urdaneta. --Ardisia peninsula Elmer, LPB 10 (1939) 3764, nom. inval., no Latin descr.; --Type: Elmer 16615 (C, NA, NY, iso), Luzon: Sorsogon prov., Irosin, Mt Bulusan Sabah, Sarawak, Philippines. LUZON: Isabela, Nueva Ecija, Zambales, Bulacan,  Rizal, Laguna, Batangas, Camarines, Albay, Sorsogon, PANAY, MINDANAO: Agusan  del Norte, Surigao. Primary forests at low and medium elevation, ascending to 450m. Photos
  52. Ardisia villosa Roxb., Fl. Ind. ed. Carey 2 (1824) 272. Thailand, Vietnam, Hong Kong, China, Taiwan, Peninsular Malaysia, Philippines, Indonesia.
  53. Ardisia warburgiana Mez in Engl., Pflanzenreich 9 (1902) 143; --Merr., EPFP 3 (1923) 265; --C.M.Hu, Blumea 44 (1999) 396. --Ardisia proteifolia Mez in Engl., Pflanzenreich 9 (1902) 143; --Merr., EPFP 3 (1923) 261. LUZON: Mountain Province, Benguet, Cagayan, Pampanga, Zambales, Sorsogon, MINDORO, PANAY, SAMAR, MINDANAO: Lanao, Bukidnon, Davao (Mt Apo, Todaya), Agusan. Forests from low elevation to 1700m.
  54. Ardisia yatesii Merr., PJS 12 c (1917) Bot. 155; EPFP 3 (1923) 266. LUZON: Quezon near Camarines Norte border (Mt Cadig). Primary forests, 350-400m. Closely allied to Ardisia tomentosa C.Presl.
  55. Ardisia zambalensis Merr., PJS 10 c (1915) Bot. 334; EPFP 3 (1923) 266. LUZON: Zambales (Mt Tapulao = High Peak). Exposed forested ridges, 1000-1400m.

 Discocalyx (DC.) Mez

Last edited by Pieter B. Pelser, 22 September 2017

Photos
  1. Discocalyx angustifolia Mez in Engl. Pflanzenreich 9 (1902) 214; --Merr., EPFP 3 (1923) 266; --Type: Gaudichaud 299 (P). LUZON: Laguna, Quezon, ALABAT, PANAY. Low elevation damp primary forests.
  2. Discocalyx angustissima Merr., PJS 12 c (1917) Bot. 143; EPFP 3 (1923) 267.  --Discocalyx longissima Merr., PJS 17 (1920) 302; EPFP 3 (1923) 268; --Type: BS 34456 Ramos & Pascasio. LUZON: Cagayan, Nueva Ecija, Quezon, MINDANAO: Surigao. Low and medium elevation primary forests, rocky slopes. Discocalyx longissima is said to differ from Discocalyx angustissima by its entire leaf margin. The type of the former in US has minute teeth.
  3. Discocalyx brachybotrys Merr., PJS 20 (1922) 424; EPFP 3 (1923) 267; --Type: BS 37463 Ramos & Edaño. MINDANAO: Zamboanga. Low elevation forests along streams.
  4. Discocalyx cybianthoides (A.DC.) Mez in Engl. Pflanzenreich 9 (1902) 213; --Merr., EPFP 3 (1923) 267; --Badula cybianthoides A.DC., Prodr. 8 (1844) 107. Borneo, Philippines. LUZON: Ilocos Norte, Abra, Cagayan, Nueva Vizcaya, Pangasinan, Nueva Ecija, Bataan, Bulacan, Rizal, Laguna, Batangas, Quezon, Sorsogon, MINDORO, MASBATE, LEYTE. Low and medium elevation primary forests, often common. Photos
  5. Discocalyx effusa Mez in Engl. Pflanzenreich 9 (1902) 213; --Merr., EPFP 3 (1923) 267. LUZON: Nueva Ecija, Rizal, Laguna, Quezon, CATANDUANES, MINDORO, PANAY, NEGROS, MINDANAO: Davao. Low and medium elevation damp primary forests. Photos
  6. Discocalyx euphlebia Merr., PJS 12 c (1917) Bot. 144; EPFP 3 (1923) 267; --Type: BS 17646 Ramos. SAMAR. Low elevation damp primary forests.
  7. Discocalyx filipes Mez, Fedde Repert. 16 (1920) 422; --Merr., EPFP 3 (1923) 267; --Type: BS 1857 Ramos. LUZON: Rizal. Medium elevation primary forests.
  8. Discocalyx hymendroides Mez, Fedde Repert. 16 (1920) 422; --Merr., EPFP 3 (1923) 267; --Type: FB 5096 Curran. LUZON: Benguet, Cagayan, Rizal. Medium elevation damp forests, ascending to 1400m in Benguet.
  9. Discocalyx insignis Merr., PJS 5 c (1910) Bot. 221; EPFP 3 (1923) 268; --Type: FB 7616 Hutchinson, Mindanao: Agusan del Norte prov., Agusan River valley near Amparo, in forests, c. 130m, 26-Aug-1907.  --Discocalyx congestiflora Elmer, LPB 8 (1915) 2780; --Type: Elmer 13724 (BISH, iso), Mindanao: Agusan del Norte, Cabadbaran, Mt Urdaneta, Sep-1912. LUZON: Tayabas, Camarines, NEGROS: Negros Occidental (Mt Marapara), LEYTE, MINDANAO: Agusan del Norte (Agusan River valley near Amparo). Low elevation primary forests. Photos
  10. Discocalyx leytensis Merr., PJS 26 (1925) 485; EPFP 4 (1925) 250; --Type: BS 41709 Edaño (US, iso), Leyte: Mt Abucayan. Feb-1923. LEYTE. In primary forests.
  11. Discocalyx linearifolia Elmer, LPB 2 (1908) 441; --Merr.,  EPFP 3 (1923) 268; --Type: Elmer 7674 (A, iso), Luzon: Quezon prov., Lucban, May-1907. LUZON: Ilocos Norte, Abra, Kalinga, Mountain Province, Sierra Madre Range from Isabela to Quezon, Laguna, Sorsogon, POLILLO, CATANDUANES, SAMAR, SIARGAO. Photos
  12. Discocalyx longifolia Merr., PJS 7 c (1912) Bot. 325; EPFP 3 (1923) 268; --Type: BS 13181 Foxworthy & Ramos, Luzon: Quezon prov., Quinatacutan, Mar-1911. LUZON: Quezon. Low elevation forests.
  13. Discocalyx luzoniensis Merr., PJS 12 c (1917) Bot. 145; EPFP 3 (1923) 268; --Type: BS 26474 (US, iso), Luzon: Nueva Ecija prov., Mt Umingan, forested slopes between 300-400m, 16-Aug-1916. LUZON: Nueva Ecija (Mt Umingan). Primary forests, c. 350m.
  14. Discocalyx macrophylla Merr., PJS 5 c (1910) Bot. 222; EPFP 3 (1923) 268; --Type: FB 17237 Curran (US, iso), Luzon: Cagayan prov., near San Vicente, forest at sea level, 8-Mar-1909. LUZON: Ilocos Norte, Cagayan. Low elevation forests.
  15. Discocalyx maculata Merr., PJS 7 c (1912) Bot. 326; EPFP 3 (1923) 268; --Type: BS 13967 Ramos, Luzon: Cagayan prov., Abulug River, in forest, Feb-1912. LUZON: Cagayan. Low elevation forests.
  16. Discocalyx merrillii Mez, PJS 1 (1906) Suppl. 274; --Merr., EPFP 3 (1923) 268; --Type: Merrill 3745 (US, syn), Luzon: Bataan prov., Mt Mariveles, Jan-1904; Whitford 213 (syn); FB 139 Barnes (syn); FB 1348 Borden (syn), Luzon: Bataan prov., Mt Mariveles along Lamao River. LUZON: Cagayan, Pangasinan, Zambales, Bataan, Bulacan, Rizal, Quezon. Medium elevation primary forests, ascending to 1000m.
  17. Discocalyx micrantha Merr., PJS 12 c (1917) Bot. 146; EPFP 3 (1923) 268; --Type: BS 27132 Ramos (US, iso), Luzon: Ilocos Norte prov., Burgos. Low elevation forests along stream, 12-Mar-1917. LUZON: Ilocos Norte, Isabela (Palanan), MINDANAO: Surigao. Low elevation primary forests. Photos
  18. Discocalyx mindanaensis Elmer, LPB 2 (1910) 674; --Merr.,  EPFP 3 (1923) 269; --Type: Elmer 11294 (PNH, holo, lost; A, GH, MO, US, iso), Mindanao: Davao, Mt Apo, Todaya, Aug-1909. MINDANAO: Lanao del Sur (Camp Keithley), Bukidnon, Davao (Mt Apo), Agusan del Norte. Medium elevation primary forests, ascending to 1200m.
  19. Discocalyx minor Mez in Engl. Pflanzenreich 9 (1902) 213; --Merr., EPFP 3 (1923) 269; --Type: Comision de la Flora Forestal de Filipinas 427 (L, iso), Luzon: Bulacan prov., Angat.
  20. Discocalyx montana Elmer, LPB 2 (1908) 443; --Merr.,  EPFP 3 (1923) 269. LUZON: Ilocos Norte, Apayao, Cagayan, Mountain Province, Benguet, Nueva Vizcaya, Quezon, Laguna, MINDORO. Stream gullies and forested slopes, 1000-2000m.
  21. Discocalyx nom. nov., ined.; --Amblyanthopsis philippinensis Mez, PJS 1 (1906) Suppl. 274; --Merr., EPFP 3 (1923) 266; --Type: Merrill 3881, Luzon: Bataan prov., Mt Mariveles. LUZON: Bataan. Primary forests along small streams, c. 800m.
  22. Discocalyx pachyphylla Merr., PJS 12 c (1917) Bot. 146; EPFP 3 (1923) 269. LUZON: Ilocos Norte, Quezon (Guinayanagan). Damp forested slopes at medium elevation, ascending to 1000m.
  23. Discocalyx palawanensis Elmer ex Merr., EPFP 3 (1923) 269; --Discocalyx merrillii auct. non Mez; Elmer, LPB 5 (1913) 1821; --Type: Elmer 12825 (PNH, holo, lost; A, MO, US, iso), Palawan: Mt Pulgar (Thumb Peak), Mar-1911. Endemic to the Philippines. MINDORO, PALAWAN, BANCALAN. Low elevation primary forests. Conservation status: Other Threatened Species (DAO 2017-11).
  24. Discocalyx phanerophlebia Merr., PJS 20 (1922) 425; EPFP 3 (1923) 269; --Type: BS 38690. MINDANAO: Bukidnon. Forests along streams, 1200-1400m.
  25. Discocalyx philippinensis (A.DC.) Mez in Engl. Pflanzenreich 9 (1902) 212; --Merr., EPFP 3 (1923) 269; --Badula philippinensis A.DC., Prodr. 8 (1844) 108. LUZON: Ilocos Norte, Mountain Province, Benguet, Nueva Ecija, Pampanga, Bataan, Rizal, Laguna, Batangas, PANAY. Damp ravines and primary forests, extending into the mossy forest, 600-1400m.
  26. Discocalyx psychotrioides Elmer, LPB 2 (1908) 442; --Merr., EPFP 3 (1923) 270. LUZON: Isabela, Rizal, Laguna, Quezon. Primary forests, 300-800m. Photos
  27. Discocalyx samarensis Merr., PJS 12 c (1917) Bot. 147; EPFP 3 (1923) 270; --Type: FB 24004 Lasquety, Samar: Northern Samar prov., San Jose de Buan, along streams, 300m, 7-May-1915. SAMAR: Northern Samar (San Jose de Buan). Lowland forest, 300m.
  28. Discocalyx sessilifolia Merr., PJS 12 c (1917) Bot. 149; EPFP 3 (1923) 270; --Type: FB 20735 Rafael & Ponce (US, iso), Mindanao: Agusan del Norte prov. (indicated in label as Butuan Subprovince), near Butuan (according to protologue). Swamp forest at low elevation, 12-Sep-1913. MINDANAO: Agusan del Norte. Swamp forest at low elevation.
  29. Discocalyx stenophylla Merr., PJS 12 c (1917) Bot. 148; EPFP 3 (1923) 270; --Type: FB 24815 Pascual (US, iso), Luzon: Pangasinan prov., Mangatarem, among undergrowth in forests, c. 300m, 23-Nov-1915. LUZON: Ilocos Norte, Pangasinan, Rizal. Primary forests, 300-700m.
  30. Discocalyx suluensis Merr., PJS 30 (1926) 420; --Type: BS 44293 Ramos & Edaño (US, iso), Tawi-tawi, damp forests at low elevation, Jul-1924. TAWI-TAWI. Lowland forests.
  31. Discocalyx tecsonii Merr., PJS 12 c (1917) Bot. 150; EPFP 3 (1923) 270; --Type: FB 24683 Tecson (US, iso), Basilan: Tumakis, in forest, 20m, locally common, teste Tecson, 4-Nov-1915. BASILAN (Tumakis; Mt Singal), MINDANAO: Misamis Occidental (Mt Malindang). Low elevation primary forests, 10-100m.
  32. Discocalyx vidalii Mez in Engl. Pflanzenreich 9 (1902) 213; --Merr., EPFP 3 (1923) 270; --Type: Vidal 423, Luzon: Quezon prov., Mt Banahaw. LUZON: Quezon (Mt Banahaw). Perhaps also represented by BS 33312 Ramos, 29288 Ramos & Edaño; from primary forests, c. 300m.
  33. Discocalyx xiphophylla Quisumb. & Merr., PJS 37 (1928) 185; --Type: BS 45678 Ramos & Edaño, Luzon: Aurora prov., Mt Alzapan, in forests, c. 1800m, 6-Jun-1925. LUZON: Aurora (Mt Alzapan). Montane forests, c. 1800m.  Comments from the original protologue: Belonging to the group of Discocalyx linearifolia Elmer, but totally different in its vegetative and inflorescence characters. Notably differs from Discocalyx angustissima Merr. in its entire leaves.

Embelia Burm.f.

Last edited by Pieter B. Pelser, 22 September 2017

Photos
  1. Embelia bataanensis Merr., Govt. Lab. Publ. (Philipp.) 29 (1905) 44; EPFP 3 (1923) 271. LUZON: Mountain Province, Benguet, Bataan. Ravines and forested ridges, 600-1300m.
  2. Embelia coriacea Wall. ex A.DC., Trans. Linn. Soc. 17 (1834) 135; --Wall., Cat. (1830) No. 2314, nom. nud.; --Merr., EPFP 3 (1923) 271. Sumatra, Peninsular Malaysia, Java, Borneo, Moluccas, Singapore, Thailand, Philippines. MINDANAO: Lanao del Sur, Bukidnon, Davao, Agusan del Norte. Low and medium elevation primary forests.
  3. Embelia elliptica Merr., PJS 20 (1922) 426; EPFP 3 (1923) 271; --Type: BS 39069 Ramos & Edaño. MINDANAO: Bukidnon. Streamside thickets, 900m.
  4. Embelia loheri Merr., PJS 27 (1925) 45; EPFP 4 (1925) 253; --Type: Loher 13845 (PNH, holo, ?lost; UC, iso), Luzon: Rizal prov., Mt Oriud, Mar-1914. LUZON: Rizal (Mt Paningtingan; Mt Oriud).
  5. Embelia luzoniensis Merr., PJS 20 (1922) 427; EPFP 3 (1923) 271; --Type: BS 20098 McGregor. LUZON: Nueva Vizcaya. Medium elevation thickets or forests.
  6. Embelia nigro-punctata Merr., PJS 7 c (1912) Bot. 326; EPFP 3 (1923) 271. LUZON: Bataan, Rizal, MINDORO, PALAWAN. Low and medium elevation thickets and forests.
  7. Embelia obtusiuscula Mez, Fedde Repert. 16 (1920) 422; --Merr., EPFP 3 (1923) 271; --Type: Elmer 5881. LUZON: Benguet. Steep slopes in thickets, c. 1200m.
  8. Embelia ovatifolia Merr., PJS 20 (1922) 428; EPFP 3 (1923) 272; --Type: BS 39001 Ramos & Edaño. MINDANAO: Bukidnon. Forested slopes, c. 1200m.
  9. Embelia philippinensis A.DC., Prodr. 8 (1844) 83; --Merr., EPFP 3 (1923) 272. --Rhamnus lando Llanos, Frag. Fl. Philip. (1851) 57; --Fern.-Vill. & Naves in Blanco, Fl. Filip. ed. 3, 41 (1880) 41. --Samara philippinensis S.Vidal, Sinopsis Atlas (1883) 30. Borneo, Philippines. LUZON: Benguet, Pangasinan, Nueva Vizcaya, Nueva Ecija, Bulacan, Rizal. Laguna, Sorsogon. LEYTE, MINDANAO: Lanao, Bukidnon, Davao. Medium elevation forests, often common. Photos
  10. Embelia whitfordii Merr., PJS 1 (1906) Suppl. 113; EPFP 3 (1923) 272; --Type: Whitford 453. --Embelia urdanetensis Elmer, LPB 8 (1915) 2783; --Merr., EPFP 3 (1923) 272; --Type: Elmer 13745. LUZON: Mountain Province, Benguet, Bataan, Laguna, MINDANAO: Agusan del Norte.

Labisia Lindl.

Last edited by Pieter B. Pelser, 22 September 2017

Photos
  1. Labisia pumila (Blume) Fern.-Vill., Novis. App. (1880) 123; --Merr., EPFP 3 (1923) 266; --Ardisia pumila Blume, Catalogus (1823) 44. Indochina, Sumatra, Peninsular Malaysia, Java, Borneo, Philippines. MINDANAO, MINDORO, LEYTE, JOLO. Damp primary forests up to 500m. Very rare and local in the Philippines. Photos

Loheria Merr.

Last edited by Pieter B. Pelser, 22 September 2017

Photos
  1. Loheria bracteata Merr., PJS 5 c (1910) Bot. 374; EPFP 3 (1923) 270; --Type: Merrill 4783. LUZON: Mountain Province, Benguet. Mossy forests, 1900-2400m. Photos
  2. Loheria crassifolia (Merr.) Stone, Micronesica 24 (1991) 72; --Amblyanthopsis crassifolia Merr., PJS 14 (1919) 445; EPFP 3 (1923) 266. LUZON: Ilocos Norte, Aurora. Forested slopes, c. 1000m. Photos
  3. Loheria jubilaria Stone, Micronesica 24 (1991) 74; --Loheria radlkoferi (Mez) Merr., EPFP 3 (1923) 271, nom. illeg.; --Jubilaria radlkoferi Mez, Fedde Repert. 16 (1920) 421; --Type: Merrill 5743 (US, iso), Mindoro: Mindoro Oriental prov., Alag River, Nov-1906. MINDORO. Low elevation damp primary forests.
  4. Loheria porteana (Mez) Merr., EPFP 3 (1923) 271; --Embelia porteana Mez, Pflanzenr. 4, 236 (1902) 302.  LUZON: Quezon, Rizal, POLILLO. Low elevation damp primary forests.

 Rapanea Aubl. = Myrsine

Lysimachia L.

Last edited by Pieter B. Pelser, 22 September 2017

Photos
  1. Lysimachia capillipes Hemsl., J. Linn. Soc. Bot. 26 (1889) 48; --Merr., EPFP 3 (1923) 274; --Bentvelzen, Fl. Males. ser. 1, 6 (1962) 179. China (Yunnan; Guizhou; Sichuan; Hubei), Taiwan, N Philippines. N LUZON. Damp thickets along streams, 1300-1500m.
  2. Lysimachia decurrens G.Forst., Prodr. (1786) 12; --Merr., EPFP 3 (1923) 274; --Bentvelzen, Fl. Males. ser. 1, 6 (1962) 185. E Himalaya, Myanmar, Thailand, Indochina, S China, Japan, Taiwan, W Java, Lesser Sunda Isls (Timor), N Philippines, Vanuatu, New Caledonia. LUZON: Benguet.
  3. Lysimachia japonica Thunb., Fl. Jap. (1784) 83; --Merr., EPFP 3 (1923) 275; --Bentvelzen, Fl. Males. ser. 1, 6 (1962) 182.
  4. Lysimachia mauritiana Lam., Encycl. 3 (1789) 592; --Merr., EPFP 3 (1923) 275; --Bentvelzen, Fl. Males. ser. 1, 6 (1962) 183. Mauritius, Réunion, E China, Korean islands, Japan, Ryukyu Isls, Taiwan, N Philippines, Bonin Isls, Marianas, Hawaiian Isls, New Caledonia, Loyalty Isls. BATANES. Coastal coral rocks and beaches. Photos
  5. Lysimachia sikokiana Miq., Ann. Mus. Bot. Lugd.-Bat. 3 (1867) 121; --Bentvelzen, Fl. Males. ser. 1, 6 (1962) 178. --Lysimachia ramosa Wall. ex Duby in DC., Prodr. 8 (1844) 65; --Merr., EPFP 3 (1923) 275; --Wall., Cat. (1828) No. 1490, nom. nud. S Japan, Ryukyu Isls, Taiwan, S China (Yunnan; Guangdong), Hainan, N Thailand, Philippines. LUZON: Mountain Province, MINDORO, NEGROS, CAMIGUIN. Mossy forests, damp ravines, 1200-2300m. Photos

Maesa Forssk.

Edited by Pieter B. Pelser, 11 August 2012

Photos
  1. Maesa cumingii Mez in Engl., Pflanzenreich  9 (1902) 49; --Merr., EPFP 3 (1923) 253; --Utteridge & Saunders, Bot. J. Linn. Soc. 145 (2004) 20; --Type: Cuming 1474 (K, lecto; BM, K, W, iso), Luzon: Camarines Sur prov., 1836-1839. --Maesa pachyphylla Merr., PJS 7 c (1912) 328; EPFP 3 (1923) 255; --Type: BS 11030 Ramos (PNH, holo, lost; BM, lecto; A (fragment), isolecto), Cebu: Hills behind the City of Cebu (Merrill 1912), Mar-1912. --Maesa undulata Merr., PJS 20 (1922) 422; EPFP 3 (1923) 255; --Type: BS 26872 Edaño (PNH, holo, lost; K, lecto), Luzon: Quezon Prov., near Pagbilao, Mar-1917. Endemic to the Philippines, but widespread throughout the archipelago. Along clearings and streams, secondary growth; 50-900m. Fl. Oct-May; fr. throughout the year. Conservation status: Not Threatened (Utteridge & Saunders 2004). Photos
  2. Maesa davaensis Quisumb., PJS 41 (1930) 341; --Utteridge & Saunders, Bot. J. Linn. Soc. 145 (2004) 22; --Type: BS 49123 Ramos & Edaño (PNH, holo, lost; UC, lecto; A, NY, P, iso), Mindanao: Davao Oriental prov., Mati, Dawau Streams, 9-Apr-1927. MINDANAO: Davao Oriental. Low elevation forests. Fl. & fr. Apr.  This species is only known from a single locality, from a single collection made in 1927. Conservation status: Critically Endangered B1a+b(iii) (Utteridge & Saunders 2004).
  3. Maesa denticulata Mez in Engl., Pflanzenreich 9 (1902) 48; --Merr., EPFP 3 (1953) 253; --Utteridge & Saunders, Bot. J. Linn. Soc. 145 (2004) 23; --Type: Warburg 15025 (E, lecto; B, lost, isolecto), Sulu Archipelago: 1888. --Maesa vulcanica Elmer in Merr., EPFP 3 (1923) 253 (1923), nom. nud.; --Voucher: Elmer 14966 (GH, L, PNH, UC), Luzon: Sorsogon prov., Irosin (Mt Bulusan), Nov-1915. Borneo (Sabah), Sulawesi, Philippines. BATAN and N LUZON to PALAWAN and MINDANAO. Forest edges, disturbed forests, secondary forests, lowland, submontane and mossy forests; 100-1550m. Fl. Dec-Aug; fr. throughout the year. Conservation status: Least Concern (Utteridge & Saunders 2004). Photos
  4. Maesa ferruginea Merr., PJS 7 c (1912) 327; EPFP 3 (1923) 254; --Utteridge & Saunders, Bot. J. Linn. Soc. 145 (2004) 24; --Type: BS 7109 Ramos (PNH, holo, lost; NY, lecto), Luzon: Abra prov., Mt Paraga, 7-Feb-1909. LUZON (Abra). In forest, c. 3000m. Only known from two collections. Fl. Feb, May; fr. May. Maesa ferruginea is only known from Abra Province (c. 4000km2). The narrow distribution range of the species, combined with the forest habitat degradation in the Philippines, suggests that Maesa ferruginea faces a very high risk of extinction in the wild. Conservation status: Endangered B1a+b(iii) (Utteridge & Saunders 2004).
  5. Maesa gaudichaudii A.DC., Ann. Sci. Nat. Bot. Sér. II, 16 (1841) 80; --Merr., EPFP 3 (1923) 254; --Utteridge & Saunders, Bot. J. Linn. Soc. 145 (2004) 25; --Type: Gaudichaud s.n. (G- DEL, holo), Luzon: Manila, Dec-1836. --Maesa piscatorum Mez, Repert. Spec. Nov Regni Veg. 16 (1920) 309; --Type: BS 19 Foxworthy (PNH, holo, lost; K, lecto; BO, GH, NSW, NY, PNH lost, isolecto), Luzon: Rizal prov., Jan-1906. LUZON, MINDORO, MINDANAO, NEGROS, PALAWAN. Forest edges, secondary growth; 100-300m. Fl. & fr. throughout the year. Conservation status: Not Threatened (Utteridge & Saunders 2004).
  6. Maesa haenkeana Mez in Engl., Pflanzenreich 9 (1902) 32; --Merr., EPFP 3 (1923) 254; --Utteridge & Saunders, Bot. J. Linn. Soc. 145 (2004) 26; --Type: Haenke 471 (PR, lecto; NY, isolecto), Luzon: Sorsogon prov., 1792. --Maesa embelioides Elmer, LPB 2 (1908) 444; --Type: Elmer 7459 (B, holo, lost; K, lecto; A, E, L, LE, isolecto), Luzon: Quezon, Lucban, May-1906. --Maesa longipetiolata Merr., PJS 11 c (1916) 28; EPFP 3 (1923) 254; --Type: BS 23358 Ramos (PNH, holo, lost; L, lecto; A, BO, K, NY, P, isolecto), Luzon: Sorsogon prov., Mt Pacdal, 5-Aug-1915. --Maesa brunnea Merr., PJS 13 c (1918) Bot. 321; Merr., EPFP 3 (1923) 253; --Type: BS 30500 Ramos (PNH, holo, lost; K, lecto; A, P, isolecto), Catanduanes: Mt Mariguidon, 14-Nov to 11-Dec-1917. --Maesa cuprifolia Elmer in Merr., EPFP 3 (1923) 254, nom. nud.; --Voucher: Elmer 16521 (A, K, L, PNH, US), Luzon: Sorsogon prov., Irosin (Mt Bulusan), Jul-1916. --Maesa platyphylla var. minor Elmer in Merr., EPFP 3 (1923) 254, nom. nud.; --Voucher: Elmer 15558 (A, BO, GH, L), Luzon: Sorsogon prov., Irosin (Mt Bulusan), Nov-1915. --Maesa irosinensis Elmer, LPB 10 (1939) 3765, nom. inval., no Latin descr.; --Utteridge & Saunders, Bot. J. Linn. Soc. 145 (2004) 37; --Type: Elmer 15558 (PNH, holo, lost; US, lecto; A 2 sheets, BO, C, GH, L, MO, NY 2 sheets, isolecto), Luzon: Sorsogon prov., Mt Bulusan, Nov-1915. LUZON: Quezon, Camarines, Albay, Sorsogon, CATANDUANES, SAMAR. Lowland forests; 150-760m. F. Mar-Jul; fr. throughout the year. Conservation status: Not Threatened (Utteridge & Saunders 2004).
  7. Maesa indica (Roxb.) A.DC., Trans. Linn. Soc. 17 (1834) 133; --Utteridge & Saunders, Bot. J. Linn. Soc. 145 (2004) 29; --Baeobotrys indica Roxb., Fl. Ind. 2 (1824) 230. --Maesa laxa Mez in Engler, Pflanzenreich 9 (1902) 32; --Merr., EPFP 3 (1923) 254; --Type: Cuming 983 (P, lecto; BM, E, K 2 sheets: Herb. Benth. & Herb. Hook., LE, MO, W 2 sheets, isolecto), Luzon: Pangasinan prov., 1836-1839. --Maesa elmeri Mez, PJS 1 (1906) Suppl. 271; --Merr., EPFP 3 (1923) 253; --Type: Elmer 5925 (US, iso), Luzon: Benguet prov., Baguio, Mar-1904. --Maesa grossedentata Mez, Repert. Spec. Nov. Regni Veg. 16 (1920) 309; --Merr., EPFP 3 (1923) 255; --Type: BS 1769 Ramos (PNH, holo, lost; NY, lecto), Luzon: Rizal prov., Jan-1907. --Maesa ramosii Quisumb. & Merr., PJS 37 (1928) 186; --Type: BS 47272 Ramos & Edaño (PNH, holo, lost; A, lecto; K, NY, isolecto), Luzon: Isabela prov., Mt Moises, 4-Mar-1926. Widespread in SE Asia, occurring throughout the Philippines. Secondary forests, mossy forests, and upper pine forests; 100-2000m. Fl. & fr. throughout the year. Conservation status: Least Concern (Utteridge & Saunders 2004). Photos
  8. Maesa manillensis Mez in Engl., Pflanzenreich 9 (1902) 34; --Utteridge & Saunders, Bot. J. Linn. Soc. 145 (2004) 31; --Type: Calléry 2 (P, holo), Luzon: Mountain Province (?), Mt Igorrotes, May-1840. N LUZON: Ilocos Norte, Mountain Province. Forests; 150-1500m. Fl. Feb, Mar, Nov; fr. Aug, Oct, Nov. Conservation status: Not Threatened (Utteridge & Saunders 2004).
  9. Maesa megalobotrya Merr., PJS 20 (1922) 422; Merr., EPFP 3 (1923) 255; --Utteridge & Saunders, Bot. J. Linn. Soc. 145 (2004) 32; --Type: Merrill 9176 (PNH, holo, lost; L, lecto; BO, K, NSW, NY, P, isolecto), Palawan: Taytay, Apr-1913. PALAWAN. Closed broad-leaved forests over ultrabasic rock; 490-590m. Fl. Apr; fr. Apr-May. This species has only been collected twice. Conservation status: Vulnerable B1ab(iii) (Utteridge & Saunders 2004).
  10. Maesa megaphylla Merr., PJS 12 c (1917) Bot. 158; Merr., EPFP 3 (1923) 255; --Utteridge & Saunders, Bot. J. Linn. Soc. 145 (2004) 33; --Type: Wenzel 1510 (PNH, holo, lost; A 2 sheets, lecto; BM, GH 2 sheets, NSW, PNH, isolecto), Leyte: Tigbao near Tacloban, 3-Aug-1915. BOHOL, LEYTE and MINDANAO. Dry forests; sea level to 600m. Fl. Jun, Aug; fr. Feb, Apr, Aug, Oct. This species is only known from seven collections, the last of which was made in 1923. Conservation status: Vulnerable B1ab(iii) (Utteridge & Saunders 2004).
  11. Maesa platyphylla Elmer, LPB 2 (1910) 667; --Utteridge & Saunders, Bot. J. Linn. Soc. 145 (2004) 34; --Type: Elmer 10570 (PNH, holo, lost; NY, lecto; E, GH 2 sheets, K, L, LE, MO, isolecto), Mindanao: Davao, Mt Apo, Todaya, May-1909. MINDANAO: Davao (Mt Apo). Primary forests; 910-1200m. Only known from two collections. Fl. May; fr. May, July and August. Conservation status: Not Threatened (Utteridge & Saunders 2004).
  12. Maesa reflexa Utteridge & R.M.K.Saunders, Kew Bull. 55 (2000) 981; Bot. J. Linn. Soc. 145 (2004) 34; --Type: BS 38973 Ramos & Edaño (A, holo; K, iso), Mindanao: Bukidnon prov., Mt Dumalucpihan, June-July 1920.  MINDANAO: Bukidnon. Habitat information not available but presumably from montane forests; c. 2350m. Fl. Jun-Jul; fr. Apr, Jun-Jul. Maesa reflexa is known from only two localities (three collections made in 1920 and 1949) from Bukidnon province (c. 9000 km2). The lack of collections from other areas on MINDANAO (especially Mt Apo, which has been relatively well collected) implies that this species has a very restricted distribution in specific mountain ranges. Conservation status: Vulnerable B1ab(iii) (Utteridge & Saunders 2004).
  13. Maesa stonei Utteridge & R.M.K.Saunders, Bot. J. Linn. Soc. 145 (2004) 37; --Type: PNH 109009 Gutierrez, Yen & Reynoso (PNH, holo; BISH, iso), Mindanao: South Cotabato prov., Tasaday forest, Jul/Aug-1972. MINDANAO: South Cotabato. Dipterocarp forests, c. 1220m. Fl. & fr. Jul-Aug. Maesa stonei is only known from a single locality, in an area which has not been extensively collected; the species is furthermore only known from one collection (dating from 1972). Conservation status: Data Deficient (Utteridge & Saunders 2004).
  14. Maesa tenera Mez in Engl. Pflanzenreich 9 (1902) 28; --Hatusima, Mem. Fac. Agr. Kagoshima Univ. 5 (1966) 43. S Japan, Ryukyus, Taiwan, S China to Indochina, Philippines. BATAN (Mt Iraya). In thickets on the ridge of Mt Iraya, 400m. Locally rare.

Doubtful and excluded names:

  1. Maesa lobuligera Mez, Repert. Spec. Nov. Regni Veg. 16 (1920) 310; Merr., PJS 20 (1922) 421; --Utteridge & Saunders, Bot. J. Linn. Soc. 145 (2004) 39; --Type: BS 727 Foxworthy (PNH, holo, lost), Palawan. See Utteridge & Saunders (2004).
  2. Maesa membranacea A.DC., Ann. Sci. Nat. Bot. Sér. II, 16 (1841) 80. --Utteridge & Saunders, Bot. J. Linn. Soc. 145 (2004) 39; --Type: from Vietnam, 1836-1837, Gaudichaud, number not cited (G-DEL 2 sheets, syn). See Utteridge & Saunders (2004).
  3. Maesa philippinensis Gaudich., Bull. Soc. Bot. France 60 (1913) 24. --Utteridge & Saunders, Bot. J. Linn. Soc. 145 (2004) 39; --Type: from Peninsular Malaysia. Malacca, 1839-1840, Cuming 2286 (L, iso). This is Maesa ramentacea (Roxb.) A.DC., see Utteridge & Saunders (2004).

Myrsine L.

Photos
  1. Myrsine amorosoana Pipoly, Sida 17 (1996) 126; --Rapanea retusa auct. non Aiton; --Merr., PJS 2 c (1907) Bot. 297; EPFP 3 (1923) 274; --Type: Merrill 5734 (US, lecto; K, NY, isolecto). LUZON: Benguet (Mt Pulag), Quirino (Mt Alzapan), Aurora (Mt Dingalan), Quezon (Mt Binuang; Quezon National Forest Park, Atimonan), Rizal (Angilog), Camarines Norte, Camarines Sur (Bicol National Park; Mt Isarog), Sorsogon, MINDORO: Mindoro Oriental (N coast, Subaan River headwaters, inland from San Teodoro; Mt Halcon), SIBUYAN (Mt Giting-giting), NEGROS: Negros Occidental (Mt Kanlaon), MINDANAO: Bukidnon (Mt Candoon; Mt Lipa; Kitanglad Range at Mt Kinasalapi and Mt Dulang-dulang). Mid- to high-montane mossy forests, 1400-2825m. Photos
  2. Myrsine apoensis (Elmer) Pipoly, Sida 17 (1996) 125; --Rapanea apoensis Elmer, LPB 2 (1910) 669; --Merr., EPFP 3 (1923) 273; --Type: Elmer 10629 (A, lecto; BISH, GH 2 sheets, K, L, NY, isolecto), Mindanao: Davao, Mt Apo, Todaya, May-1909. MINDANAO: Davao (Mt Apo, Todaya; Mt McKinley), North Cotabato (Mt Apo, Kidapawan, Geothermal Project Site). High-montane mossy forest, 1666-2900m. 
  3. Myrsine aralioides (Philipson) Pipoly, Sida 17 (1996) 152; --Rapanea aralioides Philipson, J. Bot. (London) 78 (1939) 104; --Type: from Sabah. Borneo (Sabah), Philippines. MINDANAO: Davao Oriental (Mt Hamiguitan; Mati, South Basiao; along Biga Creek), Surigao (del Sur prov.?), PALAWAN (Mt Bloomfield). 150-200m in Palawan. Seen at c. 1200m on Mt Hamiguitan. On ultramafics.
  4. Myrsine avenis (Blume) A.DC., Trans. Linn. Soc. 17 (1834) 107; --Pipoly, Sida 17 (1996) 144; --Rapanea avenis (Blume) Mez in Engl., Pflanzenreich 9 (1902) 357; --Merr., EPFP 3 (1923) 273; --Ardisia avenis Blume, Bijdr. (1825) 691. --Type: from Java. Sumatra, Java, Buru, Borneo (Sarawak; Brunei; Sabah; E & W Kalimantan), Philippines, Sulawesi. LUZON: Ilocos Norte (Mt Palimlim), Abra, Mountain Province, Ifugao (Banawe, Bayninan), Benguet (Mt Pulogloco; Mt Sto Tomas; Baguio and vicinity), Bataan (Mt Mariveles, Lamao River), Rizal (Mt Susong Dalaga; Sumag; Montalban), Laguna (Mt Makiling); Laguna-Quezon (Mt Banahaw), Quezon (Lucban), Camarines Norte (Paracale), Camarines Sur (Mt Isarog), MINDORO: Mindoro Oriental (Mt Halcon; Mansalay, Mt Yagaw), NEGROS: Negros Occidental (Mt Kanlaon), MINDANAO: Bukidnon (Mt Kitanglad), Davao (Mt Apo). Hill to mossy forests and wind-swept ridges, (300-)1400-2400m.
  5. Myrsine cruciata (Philipson) Pipoly, Sida 17 (1996) 130; --Rapanea cruciata Philipson, J. Bot. (London) 77 (1939) 105; --Type: from Mt Kinabalu, Sabah. Philippines, Borneo (Sabah: Mt Kinabalu). LUZON: Abra, Zambales, PALAWAN (Mt Beaufort). At least recorded between 490-815m in Palawan, on presumably ultramafic habitat.
  6. Myrsine densiflora Scheffer, Comm. Myrs. Archipelago Ind. (1876) 50; --Pipoly, Sida 17 (1996) 132. Philippines, Sulawesi, Moluccas, Papua New Guinea. BATAN, LUZON: Pangasinan, Zambales, PALAWAN, PANAY: Capiz, NEGROS, SAMAR, DINAGAT, MINDANAO: Surigao. Photos
  7. Myrsine fastigiata (Elmer) Pipoly, Sida 17 (1996) 149; --Rapanea fastigiata Elmer, LPB 2 (1910) 670; --Merr., EPFP 3 (1923) 273; --Type: Elmer 11447 (GH, lecto; A, K, L, NY, isolecto), Mindanao: Davao, Mt Apo, Todaya, 2000m, Aug-1909. LUZON: Ifugao (Mt Polis), Mountain Province (Bontoc; Mt Caua), Benguet (Atok, Pauai and vicinity; Mt Tabayoc; Mt Pulag; Mt Singakalsa; Mt Nangaoto; Baguio; Mt Sto Tomas), Zambales (Sta Cruz),  Bataan (Mt Mariveles, upper Lamao River), Camarines Norte (Paracale), BILIRAN (Mt Suiro), MINDANAO: Bukidnon (Mt Lipa), Davao (Mt Apo). Low to high montane sites, but most conspicuous at 2000-2500m.
  8. Myrsine glandulosa (Elmer) Pipoly, Sida 17 (1996) 154; --Rapanea glandulosa Elmer, LPB 8 (1915) 2784; --Merr., EPFP 3 (1923) 273; --Type: Elmer 14148 (A, lecto; BISH, GH, L, K, NY, isolecto), Mindanao: Agusan del Norte prov., Cabadbaran, Mt Urdaneta, 1524m, Oct-1912. NEGROS: Negros Occidental (Mt Kanlaon, SW slopes), Negros Oriental (Cuernos Mtns), LEYTE, MINDANAO: Bukidnon (Malaybalay, Kaatoan), Davao del Sur (Mt McKinley), South Cotabato (Mt Matutum), Agusan del Norte. Montane forests, 1350-2242m.
  9. Myrsine mediciloae Pipoly, Sida 17 (1996) 122; --Rapanea angustifolia Merr., PJS 20  (1922) 429, auct. non Myrsine angustifolia (A.DC.) D.Dietr., nec Myrsine angustifolia E.Meyer ex Schimper (nom. nud.), nec Myrsine angustifolia Heller, nec Myrsine angustifolia (Mez) Hosaka. --Suttonia angustifolia Mez, Pflanzenreich 9 (1902) 337; --Type: BS 33243 Ramos (A, lecto), Luzon: Ilocos Norte prov., Mt Nagapatan, Aug-1918. LUZON: Ilocos Norte (Mt Nagapatan; Burgos; Sicapoo Range, en-route from miner’s camp to Mt Burnay), Zambales (Mt Tapulao), Bataan (Mt Mariveles, Lamao River). Submontane mossy forests, 1000-1800m.
  10. Myrsine mindanaensis (Elmer) Pipoly, Sida 17 (1996) 140; --Rapanea mindanaensis Elmer, LPB 2 (1910) 671; --Merr., EPFP 3 (1923) 273; --Type: Elmer 11332 (K, lecto; A, BISH, GH, L, NY, isolecto), Mindanao: Davao, Mt Apo, Todaya, Aug-1909. LUZON: Camarines Sur (Mt Isarog), BILIRAN: Biliran (Naval, Libtong), LEYTE: Leyte (Cabalian), MINDANAO: Davao del Sur; Davao-North Cotabato (Mt Apo). Low to high montane forests, 895-2200m. Photos
  11. Myrsine oblongibacca (Merr.) Pipoly, Sida 17 (1996) 129; --Rapanea oblongibacca Merr., PJS 20 (1922) 429; EPFP 3 (1923) 273; --Type: BS 33256 Ramos (A, lecto; K, US, isolecto), Luzon: Ilocos Norte prov., Mt Palimlim, Aug-1918. LUZON: Ilocos Norte (Mt Palimlim), Cagayan (Mt Cagua; Mt Dos Cuernos; Mt Tabuan, Bauan district; Mt Balatongan), Nueva Ecija (Mt Umingan = Mingan), Aurora (Mt Dingalan), Rizal (Mt Susong Dalaga), DINAGAT. Possibly a submontane species at perhaps 800-1500m elevation. In Ilocos Norte and Dinagat possibly growing on ultramafics.
  12. Myrsine penibukana (Philipson) Pipoly, Sida 17 (1996) 147; --Rapanea penibukana Philipson, London J. Bot. 78 (1939) 105; --Type: from Mt Kinabalu, Sabah. Borneo (Mt Kinabalu), Philippines. LUZON: Abra (Mt Posuey), Aurora (Mt Dingalan), Rizal. 1000-1200m.
  13. Myrsine peregrina (Mez) Pipoly, Sida 17 (1996) 136; --Rapanea peregrina Mez, Repert. Spec. Nov. Regni Veg. 16 (1920) 424; --Type: M.S. Clemens 917 (US, lecto; A, isolecto), Mindanao: Lanao del Sur prov., Camp Keithley, Lake Lanao, Jun-1907. --Rapanea venosa Elmer, LPB 2 (1910) 672; --Merr., EPFP 3 (1923) 274, auct. non (A.DC.) Mez, nec Myrsine venosa A.DC.; --Type: Elmer 11445 (GH, lecto; A, BISH, K, L, NY, isolecto), Mindanao: Davao, Mt Apo, Todaya, 2000m, Aug-1909. BATAN (Mt Iraya), LUZON: Mountain Province (Mt Caua; Mt Data), Nueva Ecija (Mt Umingan = Mingan), MINDORO: Mindoro Oriental (Mt Halcon), PALAWAN: Palawan (Lipuun Peak), PANAY: Capiz (Mt Bulilao), Iloilo (Iwa Ilaya = Iwaya) to Coyaoyao (Koya), MINDANAO: Misamis Occidental (Mt Malindang, Lake Duminagat), Lanao del Sur (Camp Keithley), Bukidnon (Mt Kitanglad; Mt Kinasalapi; Kaatoan), Davao-North Cotabato (Mt Apo, Todaya and Kidapawan approaches, respectively), Davao del Sur (Mt McKinley), Davao Oriental (Mt Kampalili), South Cotabato (Mt Matutum). Submontane and montane mossy forests, up to 2800m.
  14. Myrsine philippinensis A.DC., Prodr. 8 (1844) 94; --Pipoly, Sida 17 (1996) 141; --Rapanea philippinensis (A.DC.) Mez, in Engl., Pflanzenreich 9 (1902) 364; --Merr., EPFP 3 (1923) 273; --Type: Cuming 869 (G, holo; fragm. in A; K 2 sheets, L, iso), Luzon: Albay prov., 1841. LUZON: Batangas, Quezon (Infanta; Tagkawayan; Calauag Bay), Camarines Norte (Paracale), Albay, POLILLO, MINDORO: Mindoro Oriental (Baco), SIBUYAN (Mt Giting-giting), CUYO ISL GROUP (PAMALICAN), GUIMARAS, LEYTE, SAMAR: Northern Samar, JOLO, MINDANAO: Surigao del Norte. Coastal forests and secondary vegetation, on sands and mangrove swamps, sea level to 305m.  Photos

Primula L.

Photos
  1. Primula umbellata (Lour.) Bentvelzen, Fl. Males. ser. 1, 6 (1962) 191; --Androsace umbellata (Lour.) Merr., PJS 15 (1919) 237; EPFP 3 (1923) 274; --Drosera umbellata Lour., Fl. Cochinch. (1790) 186. India, Bangladesh, Myanmar, N Vietnam, China (Yunnan; Sichuan; Guizhou; Guangdong; NE China), Korea, Japan, Ryukyu Isls, Taiwan, N Philippines, E New Guinea. LUZON: Mountain Province. Montane forests in shaded moist places, 1300-1500m.

Literature

Bentvelzen, PAJ. 1962. Primulaceae. Flora Malesiana series 1, 6: 173-192

DENR Administrative Order. 2017. Updated national list of threatened Philippine plants and their categories. DAO 2017-11

Dubéarnès, A, A Julius & TA Utteridge. 2015. A synopsis of the genus Embelia in Peninsular Malaysia and Singapore. Studies in Malaysian Myrsinaceae III. Kew Bulletin 70: 1-33.

Hu Chi Ming, 1999. New synonyms and combinations in Asiatic Ardisia. Blumea 44: 391-406

Pipoly III, J. J. 1996. Contributions towards a new flora of the Philippines: I. A synopsis of the genus Myrsine (Myrsinaceae). Sida 17: 115-162.


Utteridge, TMA & RMK Saunders. 2004. The genus Maesa (Maesaceae) in the Philippines. Botanical Journal of the Linnean Society 145: 17-43.


Acknowledgements                             

Derek Cabactulan and Racquel Tan Chua-Barcelo kindly provided some photographs for this webpage.

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Last updated 16 November 2017