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Urticaceae

including Cecropiaceae

Astrothalamus C.B.Rob.

No photos
  1. Astrothalamus reticulatus (Wedd.) C.B.Rob., PJS 6 c (1911) Bot. 19, pl. 3; --Merr., EPFP 2 (1923) 98. --Maoutia (?) planitora C.B.Rob., PJS 3c (1908) 180; --Type: RS Williams 2079 (cited by Robinson, loc. cit.), Mindanao: Zamboanga, Sax River, 31 Jan 1905. Marianas, Borneo (Sarawak), MINDANAO: Zamboanga (San Ramon; Sax River). Low elevation thickets.

Boehmeria Jacq.

Edited by Pieter B. Pelser, 2 February 2015

Photos
  1. Boehmeria beyeri C.B.Rob., PJS 6 c (1911) Bot. 309; --Merr., EPFP 2 (1923) 89; --Type: BS 13540 Beyer (K, NY, iso), Luzon: Ifugao prov., Bila, May-1911. Endemic to Philippines (Luzon). LUZON: Ifugao, Benguet (Mt Sto Tomas). Mossy forests, up to 1900m. Critically Endangered (CR).
  2. Boehmeria densiflora Hook. & Arn., Bot. Beechy’s Voy. (1841) 217; --Merr., EPFP 2 (1923) 90; --Yang, et al., Fl. Taiwan 2nd ed., 2 (1996) 199; --Type from Japan. S China, Hong Kong, Ryukyu and Bonin Isls., Taiwan, Philippines. LUZON: Ilocos Norte, Ifugao, Mountain Province, Benguet. Open slopes, up to 1650m.
  3. Boehmeria depauperata Wedd., Ann. Sci. Nat., Bot. sér. 4, 1 (1854) 202. --Pipturus mindanaensis Elmer, LPB 3 (1910) 898; --Merr., EPFP 2 (1923) 95; --Type: Elmer 10520 (PNH, holo, lost; K, LE, MO, US, iso), Mindanao: Davao, Mt Apo, Todaya, 1125m, May-1909. --Boehmeria cypholophoides Merr., PJS 8 c (1913) Bot. 368; EPFP 2 (1923) 90; --Type: Wenzel 35 (A, BM, F, G, L, US, iso), Leyte: Dagami, forested foothills, c. 60m, 20-May-1913. From Pakistan, India and Bangladesh to Bhutan, Sri Lanka, Myanmar, SW China, Thailand, Vietnam, Malaysia, Indonesia (Sumatra, Java, Borneo, Papua), Papua New Guinea, Philippines. LEYTE (low elevation forests), MINDANAO: Davao (Mt Apo, open slopes, c. 1125m). 100-1700m. Least Concern (LC).
  4. Boehmeria heterophylla Wedd. in Arch. Mus. Paris 9 (1856) 351; --Merr., EPFP 2 (1923) 90; --Type: Cuming 1561 (BM, G, K, M, P, iso), Mindoro. --Boehmeria blumei Wedd., in DC Prodr. 161 (1869) 204; --Merr., EPFP 2 (1923) 89; --Type: Cuming 731 (cited by Robinson, PJS 6 c (1911) 2), Luzon: Pampanga or Bulacan prov. --Boehmeria villosa C.B.Rob., PJS 6 c (1911) Bot. 3; --Merr., EPFP 2 (1923) 91; --Type: BS 3574 Fenix, erroneously cited ‘3754’ by Robinson & Merrill and Wilmot-Dear & Friis (2013) (NY, iso), Batan: Sto Domingo de Basco, 27 May 1907. SABTAN, BATAN. Habitat not noted.Indonesia (Sulawesi), Papua New Guinea, Philippines. LUZON, MINDORO, PANAY, MINDANAO. Thickets, damp ravines, and forests, sea-level up to c. 1100m. Near Threatened (NT). Photos
  5. Boehmeria multiflora C.B.Rob., PJS 3 c (1908) Bot. 179, --Merr., EPFP 2 (1923) 90; --Type: RS Williams 1088 (NY, US, iso), Luzon: Benguet prov., Baguio, 10-Jun-1904. Indonesia (Bouton), Philippines. LUZON: Ilocos Norte, Kalinga, Mountain Province, Benguet, Nueva Vizcaya, Nueva Ecija, BOHOL, MINDANAO. Open slopes and thickets, 600-2600m. Vulnerable (VU). Photos
  6. Boehmeria nivea (L.) Gaudich., Bot. Freyc. Voy. (1826) 499; --Merr., EPFP 2 (1923) 90. Not native to the Philippines.
  7. Boehmeria pilosiuscula (Blume) Hassk., Cat. Hort. Bot. Bogor. (Hasskarl) (1844) 79; --Urtica pilosiuscula Blume, Bijdr. Fl. Ned. Ind. 10 (1825) 491; --Type from Indonesia. India, Sri Lanka, Myanmar, S China, Taiwan, Thailand, Indonesia (Sumatra, Java, Borneo, Lesser Sunda Isls.), Philippines. LUZON. Up to 1650m. Near Threatened (NT) or Vulnerable (VU).
  8. Boehmeria rugosissima (Blume) Miq., Pl. Jungh. 1 (1851) 32; --Urtica rugosissima Blume, Bijdr. (1825) 490; --Type from Indonesia. Indonesia (Sumatra, Java, Sulawesi), Philippines. MINDORO, MINDANAO. 450-1600m. Near Threatened (NT) or Vulnerable (VU).
  9. Boehmeria virgata (G.Forst.) Guill., Ann. Sci. Nat. Ser. II, vii. (1837) 182; --Urtica virgata G.Forst., Florulae Insularum Australium Prodromus (1786) 66, no. 345. Widespread in Tropical Africa, Madagascar, Oceania, SE Asia, Australia.
  10. Boehmeria zollingeriana Wedd. in Ann. Sc. Nat. Bot. iv, 1 (1854) 201; --Merr., PJS 14 (1919) 381; EPFP 2 (1923) 91; --Type from Java. SE India, Myanmar to SW China, Taiwan, Thailand, Laos, Vietnam, Indonesia (Java), Philippines. LUZON: Ilocos Norte (Mt Nagapatan; Bangui), Pangasinan (Umingan). Low and medium elevation ravines and open slopes, ascending to 14000m. Least Concern (LC).
Possible occurence in the Philippines:
  1. Boehmeria sieboldiana Blume, Museum Botanicum Lugduno Batavum 2 (1857) 220.
Name of uncertain identity (Wilmot-Dear & Friis, 2013):
  1. Boehmeria ruprestris C.B.Rob., PJS 6 c (1911) Bot. 5; --Merr., EPFP 2 (1923) 91; --Type: BS 5542 Ramos (cited by Robinson, loc. cit.), Luzon: Benguet prov., La Trinidad, Rio Trinidad (=Balili River). LUZON: Benguet. Rocks along streams, c. 1300m. Probably B. pilosiuscula.

Cecropia Loefl. [Not native]

No photos
  1. Cecropia peltata L. Native of tropical America. Recently introduced in the Philippines. Very prolific. (Pancho, Fl. Mt Makiling 1 (1983) 121. Photos

Chamabainia Wight

No photos
  1. Chamabainia cuspidata Wight, Icon. Pl. Ind. Or. 6 (1853) 11, t. 1981; --Merr., EPFP 2 (1923) 91; --Yang, et al., Fl. Taiwan 2nd ed., 2 (1996) 202. Sri Lanka, SW China, Taiwan. SE Asia. LUZON: Mountain Province (Mt Data), Benguet (Mt Pulag), MINDANAO: Davao (Mt Apo). Mossy forests, grassy montane summits, cliffs, etc. 1700m to 2800m.

Conocephalus Blume =Poikilospermum

Cypholophus Wedd.

Photos
  1. Cypholophus brunneolus Elmer, LPB 3 (1910) 896; --Merr., EPFP 2 (1923) 93; --Type: Elmer 11641 (MO, NY, iso), Mindanao: Davao, Mt Apo, Todaya, dense woods at 4000ft, Sep-1909. Philippines, Borneo. LUZON: Mountain Province, Benguet, PANAY, NEGROS: Negros Occidental (Mt Kanlaon), MINDANAO: Davao (Mt Apo). Shaded ravines, mossy forests, etc., 600-2400m. Photos 
  2. Cypholophus microphyllus Elmer, LPB 3 (1910) 895; --Merr., EPFP 2 (1923) 93; --Type: Elmer 11588 (PNH, holo, lost; MO, NY, 2 sheets, US, iso), Mindanao: Davao, Mt Apo, Todaya, seepage ledges along Cati Creek at 6000ft, Aug-1909. MINDANAO: Davao (Mt Apo). c. 1800m.
  3. Cypholophus moluccanus (Blume) Miq., Ann. Mus. Bot. Lugd.-Bat. 4 (1869) 305; --Merr., EPFP 2 (1923) 93; --Yang, et al., Fl. Taiwan 2nd ed., 2 (1996) 202. Taiwan, Lanyu, Micronesia, Hawaii, Malesia: Sumatra and the Philippines. LUZON: Mountain Province to Sorsogon, MINDORO, PANAY, NEGROS, LEYTE, MINDANAO. Damp forested ravines, low elevation to 1500m. Photos 

Debregeasia Gaudich.

Photos
  1. Debregeasia longifolia (Burm.f.) Wedd. in DC Prodr. 161 (1869) 23524; --Merr., EPFP 2 (1923) 95; --Urtica longifolia Burm.f., Fl. Ind. (1768) 297; --Type from India. --Debregeasia angustifolia C.B.Rob., PJS 6 c (1911) Bot. 18; --Type: RS Williams 1551 (US, iso), Luzon: Benguet prov., Baguio. --Debregeasia luteocarpa Elmer, LPB 8 (1915) 2843; --Type: Elmer 14068 (L, MO, NA, NY, U, US, iso), Mindanao: Agusan del Norte prov., Mt Urdaneta, Oct-1912. India to China, Sumatra, Java, Borneo, Philippines. LUZON: Abra (Mt Paraga), Kalinga (Balbalasang), Mountain Province (Bauco), Ifugao, Benguet (Mt Pulag; Mt Sto Tomas; Baguio; Imugan-Nozo Trail), MINDANAO: Agusan del Norte (Mt Urdaneta). Common in thickets, ravines, and in damp forests, 1000-2000m. Note: Iso sheets at L & MO annotated by Wilmot-Dear as D. velutina Gaudich. in 1987. Photos 

Dendrocnide Miq.

Photos
  1. Dendrocnide carriana Chew, Gard. Bull. Sing. 25 (1969) 45, Fig. 17; --Type from Papua New Guinea. NE New Guinea (Papua), Moluccas, Lesser Sunda Isls (Bali, Lombok, Timor), MINDORO: Mindoro Oriental (Mt Yagaw, PNH 18933 Conklin, A, L, PNH).
  2. Dendrocnide crassifolia (C.B.Rob.) Chew, Gard. Bull. Sing. 21 (1965) 203; ibid. 25 (1969) 46, Fig. 5; --Laportea crassifolia C.B.Rob., PJS 5 c (1910) Bot. 480; --Merr., EPFP 2 (1923) 73; --Type: FB 3925 Hutchinson (US, holo; K, NY, iso), Mindanao: Misamis Occidental prov., Mataline Falls Talaud Isls(?), and the Philippines. MINDANAO: Misamis, Lanao, Davao Oriental (Mt Mayo). In thickets, 400-800m.
  3. Dendrocnide densiflora (C.B.Rob.) Chew, Gard. Bull. Sing. 21 (1965) 203; ibid. 25 (1969) 24, Fig. 7; --Laportea densiflora C.B.Rob., PJS 5 c (1910) Bot. 479; --Merr., EPFP 2 (1923) 73; --Type: Hallier 7414 (L, NY, iso), Mindanao: Zamboanga prov., San Ramon. --Laportea subpeltata C.B.Rob., PJS 5 c (1910) Bot. 485; --Merr., EPFP 2 (1923) 75; --Type: FB 4586 Hutchinson (NY, US, iso), Mindanao: Misamis Occidental prov., Mt Malindang. NEGROS: Negros Oriental (Lake Balinsasayao), CAMIGUIN, MINDANAO: Zamboanga (San Ramon), Misamis Occidental (Mt Malindang). Thickets and forests, low elevation to c. 1000m.
  4. Dendrocnide elliptica (Merr.) Chew, Gard. Bull. Sing. 21 (1965) 203; ibid. 25 (1969) 16, Fig. 4; --Laportea elliptica Merr., J. Arn. Arb. 35 (1954) 135; --Type (lectotype) from Sabah. Borneo (Sabah, Sarawak, Kalimantan) and the Philippines. LUZON: Ifugao prov. (Banawe, PNH 80485 Conklin & Buwaya, L). 
  5. Dendrocnide luzonensis (Wedd.) Chew, Gard. Bull. Sing. 21 (1965) 203; ibid. 25 (1969) 40, Fig. 15; --Laportea luzonensis (Wedd.) Warb., Perkins Frag. Fl. Philip. (1905) 168; --Merr., EPFP 2 (1923) 73; --Laportea crenulata (Roxb.) Wedd. var. luzonensis Wedd., Arch. Mus. Hist. Nat. Paris 9 (1856) 133; --Type: Cuming 522 (CGE, G, G-DC, K, L, M, MEL, MO, UPS, iso), locality not known. --Laportea agusanensis Elmer, LPB 8 (1915) 2860; --Merr., EPFP 2 (1923) 72; --Type: Elmer 14019 (BO, E, F, L, MO, US, iso), Mindanao: Agusan del Norte prov., Mt Urdaneta. --Laportea latilanceolata Elmer, LPB 9 (1934) 3238, not validly published; no Latin description; --Type: Elmer 15206 (A, BO, C, F, FI, K, L, MO, S, U, UC, US, Z, iso).
  6. Dendrocnide meyeniana (Walp.) Chew, Gard. Bull. Sing. 21 (1965) 204; ibid. 25 (1969) 21, Fig. 6; --Yang, et al., Fl. Taiwan 2nd ed., 2 (1996) 204; --Laportea meyeniana (Walp.) Warb., Perkins Frag. Fl. Philip. (1905) 168; --Merr., Fl. Manila (1912) 181; EPFP 2 (1923) 73; --Urtica meyeniana Walp., Nov Act. Acad. Nat. Cur. 19 (1843) Suppl. 1: 422. --Dendrocnide diffusa (C.B.Rob.) Chew, Gard. Bull. Sing. 21 (1965) 203; --Laportea diffusa C.B.Rob., PJS 5 c (1910) Bot. 482; --Merr., EPFP 2 (1923) 73; --Type: FB 5149 Curran (NY, US, iso), Luzon: Tarlac prov. --Laportea gaudichaudiana Wedd., Arch. Mus. Hist. Nat. Paris 9 (1856) 137; --Type: Cuming 567 (CGE, K), without locality. --Laportea leytensis C.B.Rob., PJS 5 c (1910) Bot. 484; --Merr., EPFP 2 (1923) 73; --Type: Elmer 7351 (A, BO, E, FI, K, NY, iso), Leyte: Palo. Photos
  7. Dendrocnide rigidifolia (C.B.Rob.) Chew, Gard. Bull. Sing. 21 (1965) 205; ibid. 25 (1969) 26, Fig. 8; --Laportea rigidifolia C.B.Rob., PJS 5 c (1910) Bot. 483; --Merr., EPFP 2 (1923) 74. – Type: RS Williams 991 (BO, E, F, L, NY, 2 sheets, US, iso), Luzon: Benguet prov., Mt Sto Tomas. LUZON: Benguet (Sablan; Mt Sto Tomas). Mossy forest, c. 2000m on Mt Sto Tomas,elevation medium  in Sablan.
  8. Dendrocnide stimulans (L.f.) Chew, Gard. Bull. Sing. 21 (1965) 206; ibid. 25 (1969) 11, Fig. 2. -- Laportea gracilipes Elmer, LPB 3 (1910) 876; --Merr., EPFP 2 (1923) 73; --Type: Elmer 10499 (PNH, holo, lost; A, BO, E, F, G, K, L, MO, NY (2sheets), WRSL, US, Z, iso), Mindanao: Davao del Sur prov., Mt Apo. --Laportea brunnea Merr., PJS 10 c (1915) Bot. 298; --Merr., EPFP 2 (1923) 72; --Type: BS 17559 Ramos (K, US). --Laportea lanaensis C.B.Rob., PJS 5 c (1910) Bot. 483; --Type: Clemens 462 (F, M), Mindanao: Lanao del Sur prov., Lake Lanao, Camp Keithley. --Dendrocnide stenophylla (Quisumb.) Chew, Gard. Bull. Sing. 21 (1965) 206; --Laportea stenophylla Quisumb., PJS 41 (1930) 317, t. 1; --Type: BS 49413 Ramos & Edaño (B, NY, UC, iso), Mindanao: Davao Oriental prov., Mt Mayo. Hainan, Taiwan, Thailand, Laos, Vietnam, Sumatra, Peninsular Malaysia, Java, Lesser Sunda Isls (Bali, Lombok), Anambas, Borneo (Sabah, Sarawak, Kalimantan), Philippines, Sulawesi, Moluccas (Buru). LUZON: Bataan, Laguna, Sorsogon, SAMAR (Cauayan Valley), MINDANAO: Lanao del Sur (Camp Keithley), Davao del Sur (Mt Apo), Davao Oriental (Mt Mayo), Agusan. Primary and secondary forests, thickets, near sea level to 1200m.
  9. Dendrocnide subclausa (C.B.Rob.) Chew, Gard. Bull. Sing. 21 (1965) 206; ibid. 25 (1969) 28, Fig. 9; --Laportea subclausa C.B.Rob., PJS 5 c (1910) Bot. 486; --Merr., EPFP 2 (1923) 74; --Type: Merrill 6290 (K, NY, US, iso), Luzon: Laguna prov., Mt Makiling. --Laportea pendula Merr., PJS 13 c (1918) Bot. 270; --Merr., EPFP 2 (1923) 74; --Type: BS 30229 Ramos (K, US, iso), Catanduanes: Mt Mariguidon, along small streams in damp forests, c. 200m, 21-Nov-1917. LUZON: Isabela (San Mariano; Palanan), Laguna (Mt Makiling), Camarines, Sorsogon (Mt Bulusan), CATANDUANES (Mt Mariguidon), PANAY: Capiz (Jamindan), MINDANAO: Agusan del Norte (Tungao). Lowland forests, usually along streams, ascending to 800m. Photos 
  10. Dendrocnide venosa (Elmer) Chew, Gard. Bull. Sing. 21 (1965) 207; ibid. 25 (1969) 30, Fig. 10; --Laportea venosa Elmer, LPB 3 (1910) 878; --Merr., EPFP 2 (1923) 75; --Type: Elmer 11948 (A, BO, E, F, FI, K, L, MO, NY (2 sheets), US, WRSL, Z), Mindanao: Davao del Sur prov., Mt Apo. --Laportea monticola Merr., PJS 10 c (1915) Bot. 299; --Merr., EPFP 2 (1923) 74; --Type: BS 19765 RC McGregor (US, iso), Luzon: Ifugao prov., Mt Polis. --Laportea triplinervia Merr., PJS 10 c (1915) Bot. 300; --Merr., EPFP 2 (1923) 75; --Type: BS 19462 Ramos (L, US, iso), Luzon: Quezon prov., Mt Pular. --Laportea conduplicata Elmer, LPB 9 (1934) 3237, not validly published; no Latin description; --Type: Elmer 17152 (A, BO, C, F, FI, K, L, MO, NY, S, U, UC, US, Z), Luzon: Sorsogon prov., Mt Bulusan. LUZON: Ifugao (Mt Polis), Quezon (Mt Pular), Sorsogon (Mt Bulusan), NEGROS: Negros Oriental (Lake Balinsasayao), MINDANAO: Davao del Sur (Mt Apo). Low montane to montane mossy forests, 400-2000m.

 Distemon Wedd. 1857 non Bouche 1844 =Neodistemon

Elatostema J.R.Forst. & G.Forst.

Photos
  1. Elatostema acrophilum C.B.Rob., PJS 5 c (1910) Bot. 523; --Merr., EPFP 2 (1923) 78; --Type: BS 6577 Robinson (US, iso), Luzon: Laguna prov., Mt Banahaw, summit area, Jan-1909. LUZON: Laguna, Quezon, Camarines, CATANDUANES (BS 75410 Ramos & Edaño, SING). Lower montane to high-montane mossy forests, 1000-2100m.
  2. Elatostema acuminatissimum Merr., PJS 14 (1919) 375; EPFP 2 (1923) 79; --Type: BS 29015 Ramos & Edaño (US, iso), Luzon: Quezon prov., Umiray, 2-Jun-1917. LUZON: Quezon (Umiray). Low elevation damp forest along streams.
  3. Elatostema agusanense Elmer, LPB 8 (1908) 2845; --Merr., EPFP 2 (1923) 79; --Type: Elmer 14250 (GH, US, iso), Mindanao: Agusan del Norte prov., Mt Urdaneta, Aug-1912. LUZON: Laguna, Sorsogon, MINDANAO: Agusan del Norte (Mt Urdaneta). Forested ravines along small streams at medium elevation.
  4. Elatostema angustatum C.B.Rob., PJS 5 c (1910) Bot. 533, erroneously as ‘523’ in --Merr., EPFP 2 (1923) 79; --Type: FB 3266 Ahern’s collector (cited by Robinson, loc. cit.), Luzon: Rizal prov., Bosoboso. --Elatostema camiguinense Elmer, LPB 8 (1915) 2849; --Type: Elmer 14227 (GH, US, iso), Camiguin: Mambajao, Nov-1912. LUZON: Rizal, Laguna, CAMIGUIN, MINDANAO. Along streams in shaded ravines at low and medium elevation. 
  5. Elatostema antonii Elmer, LPB 8 (1908) 2847; --Merr., EPFP 2 (1923) 79; --Type: Elmer 13894 (MO, US, iso), Mindanao: Agusan del Norte prov., Mt Urdaneta, Sep-1912. MINDANAO: Agusan del Norte (Mt Urdaneta). Boulders in densely shaded ravines along streams, c. 900m,
  6. Elatostema apoense Elmer, LPB 3 (1910) 885; --Merr., EPFP 2 (1923) 79; --Type: Elmer 11793 (MO, US, iso), Mindanao: Davao, Mt Apo, Todaya, Sep-1909. MINDANAO: Davao (Mt Apo). Dense forests, c. 1200m. According to Merrill, near E hookerianum Wedd.
  7. Elatostema appendiculatum Merr., PJS 14 (1919) 379; EPFP 2 (1923) 79; --Type: BS 29129 Ramos & Edaño (US, iso), Luzon: Quezon prov., Mt Tulaog, along small streams in damp forests at low elevation, 25-May-1917. LUZON: Quezon (Mt Tulaog). Low elevation forests along streams.
  8. Elatostema banahaense C.B.Rob., PJS 5 c (1910) Bot. 526, --Merr., EPFP 2 (1923) 79; --Type: BS 9856 Robinson (US, iso), Luzon: Laguna prov., Mt Banahaw, 1500m, Mar-1910. --Elatostema scriptum C.B.Rob., PJS 5 c (1910) Bot. 529; --Merr., EPFP 2 (1923) 85; --Type: BS 9766 Robinson (cited by Robinson, loc. cit.), Luzon: Laguna (Mt Banahaw), on rocks in forests, c. 800m. --Elatostema carinoi WR Shaw ex C.B.Rob., PJS 5 c (1910) Bot. 532; ibid. 6 c (1911) Bot. 307; --Type: Phil. Normal Sch. 428 Cariño (cited by Robinson, loc. cit.), Luzon: Benguet: Baguio. --Elatostema bulusanense Elmer, LPB 9 (1934) 3227; --Syntypes: Elmer 16172 (A, MO, US), 15575 (MO, US), 16399 (GH, MO, US), Luzon: Sorsogon prov., Mt Bulusan. LUZON: Ifugao (Kiangan), Mountain Province, Benguet, Cagayan (Gonzaga, BS 78627 Edano, SING), Laguna (Mt Makiling; Mt Banahaw), Sorsogon (Mt Bulusan), NEGROS: Negros Occidental (Katugasan, PNH 21834, SING), SIQUIJOR, MINDANAO: Zamboanga. Damp forests at medium and higher elevation, ascending to 2100m in the mossy forest on high mountains. Photos 
  9. Elatostema baruringense Elmer, LPB 3 (1910) 890; --Merr., EPFP 2 (1923) 79; --Type: Elmer 10916 (PNH, holo, lost; GH, MO, US, iso), Mindanao: Davao, Mt Apo, Todaya, Jun-1909. MINDANAO: Davao (Mt Apo). Dense forests, c. 1200m.
  10. Elatostema benguetense C.B.Rob., PJS 5 c (1910) Bot. 539; --Merr., EPFP 2 (1923) 80; --Type: FB 5006 Curran, Luzon: Benguet prov., Mt Tonglon (=Sto. Tomas). LUZON: Ifugao, Mountain Province (Mt Data), Benguet (Pauai; Mt Pulag; Mt Sto Tomas), MINDORO: Mindoro Oriental (Mt Halcon, PNH 3572 Edano), NEGROS: Negros Occidental (Mt Kanlaon, PNH 21963 Edano). Mossy forests, 1800-2300m. Photos 
  11. Elatostema bontocense Merr., PJS 20 (1922) 371; --Merr., EPFP 2 (1923) 80; --Type: BS 37660 Ramos & Edaño (US, iso), Luzon: Mountain Province, Mt Polis, on tree trunks in the mossy forest, 25-Feb-1920. LUZON: Mountain Province. Mossy forests, 1800m.
  12. Elatostema brongniartianum Wedd., Ann. Sc. Nat. Bot. iv 1 (1854) 190; --Merr., EPFP 2 (1923) 80; --Type: Cuming 629, Luzon: Laguna prov., Calauan. --Elatostema rupicolum Elmer, LPB 9 (1934) 3233; --Type: Elmer 18000 (GH, MO, US, iso), Luzon: Laguna prov., Mt Makiling, Jun/Jul-1917. N LUZON to MINDANAO. On rocks, cliffs, and banks of shaded ravines at medium elevation, in Luzon ascending to 1400m. 
  13. Elatostema brunneolus Elmer, LPB 9 (1934) 3219; --Type: Elmer 22102 (B, GH, MO, US, iso), Luzon: Pampanga prov., Mt Pinatubo, Camp Stotsenburg, May-1927. LUZON: Pampanga (Mt Pinatubo), NEGROS: Negros Oriental (Cuernos Mtns, PNH 7332 Edano)
  14. Elatostema brunonianum Quisumb., PJS 41 (1930) 318, t. 2; --Type: BS 49401 Ramos & Edaño (PNH, holo, possibly lost; UC?, iso), Mindanao: Davao Oriental prov., Mt Mayo, trunk epiphyte in mossy forest, c. 1300m. MINDANAO: Davao Oriental (Mt Mayo), trunk epiphyte in mossy forest, 1300m. Known only from the type.
  15. Elatostema buderi H Schröter, Fedde, Repert. Beih. 83 i (1935) 6; --ibid. 103. MINDORO.
  16. Elatostema capizense Merr., PJS 20 (1922) 372; EPFP 2 (1923) 80; --Type: BS 30748 Ramos & Edaño (US, iso), Panay: Aklan prov., Mt Macosolon (=Magosolon), boulders along small streams in damp forests at low elevation, 19-Apr-1918. PANAY: Aklan (Mt Magosolon). Low elevation forests along stream.
  17. Elatostema catalonanum H Schröter, Fedde, Repert. Beih. 83 i (1935) 28, in obs., nomen; --ibid. 113. --Elatostematoides mindanaense C.B.Rob., PJS 5 c (1910) Bot. 502; --Merr., EPFP 2 (1923) 88; --Type: Copeland 935 (staminate), 936 (pistillate), cited by Robinson, loc. cit., Mindanao: Davao del Sur prov., Catalunan. MINDANAO (Davao). In forests at medium elevation.
  18. Elatostema catanduanense Merr., PJS 13 c (1918) Bot. 271; EPFP 2 (1923) 80; --Type: BS 30555 Ramos (US, iso), Catanduanes: Sto Domingo River, on stones along small streams in forests at low elevation, 5-Dec-1917. CATANDUANES. Rupicolous in low elevation forested streams.
  19. Elatostema caudatum Hallier f. Ann. Jard. Bot. Buitenzorg xiii. 307. Photos
  20. Elatostema cheirophyllum C.B.Rob., PJS 5 c (1910) Bot. 518; --Merr., EPFP 2 (1923) 80; --Type: Whitford 1595 (US, iso), Negros: Negros Occidental prov., Himugaan River, May-1906. NEGROS: Negros Occidental, MINDANAO: Zamboanga. Banks and boulders along streams in shaded ravines at medium elevation. 
  21. Elatostema contiguum C.B.Rob., PJS 5 c (1910) Bot. 536; --Merr., EPFP 2 (1923) 80; --Type: BS 9479 Robinson, Luzon: Quezon prov., Mahabang Sugsugan River. LUZON: Laguna, Quezon, Camarines. Shaded stream embankments at medium elevation.
  22. Elatostema delicatulum Wedd., Ann. Sc. Nat. Bot. iv 1 (1854) 190; --Merr., EPFP 2 (1923) 80. --Elatostema delicatum Elmer, LPB 2 (1908) 467; --Type: Elmer 10343 (PNH, holo, lost; GH, MO, US, iso), Negros: Negros Oriental prov., Cuernos Mtns, Jun-1908. LUZON: Cagayan, Isabela, Rizal, Laguna, PANAY, NEGROS. On rocks and ledges in shaded stream beds at low and medium elevation. 
  23. Elatostema discolor C.B.Rob., PJS 6 c (1911) Bot. 301; --Merr., EPFP 2 (1923) 81; --Type: BS 13475 Ramos (US, iso), Luzon: Benguet prov., Baguio, in ravines, c. 1400m, May-1911. LUZON: Ifugao, Benguet. Forests at or above 1400m.
  24. Elatostema diversilimbum Merr., PJS 14 (1919) 378; --Merr., EPFP 2 (1923) 81; --Type: Merrill 9127 (US, iso), Mindanao: Zamboanga: Mtns back of Zamboanga, damp shaded ravines along small streams on banks and ledges, 200-400m, 6-Dec-1911. MINDANAO: Zamboanga. On banks and ledges in damp, shaded ravines, 200-400m. 
  25. Elatostema edanoi Merr., PJS 20 (1922) 374; --Merr., EPFP 2 (1923) 81 (‘edañoii’); --Type: BS 37603 Ramos & Edaño (US, iso), Luzon: Kalinga prov., Mt Masingit near Lubuagan, on tree trunks in forests along streams, c. 1300m, 17-Feb-1920. LUZON: Kalinga. In forests, c. 1300m.
  26. Elatostema edule C.B.Rob., PJS 5 c (1910) Bot. 531; --Merr., EPFP 2 (1923) 81; --Yang, et al., Fl. Taiwan 2nd ed., 2 (1996) 206, pl. 103. Lanyu, Lutao, N Philippines. BATAN. Along streams, ravines and moist forest floor.
  27. Elatostema elmeri Merr., EPFP 2 (1923) 81; --Elatostema pictum Elmer, LPB 8 (1915) 2850, non Hallier f; --Type: Elmer 13484. LUZON: Quezon, MINDANAO: Agusan del Norte. About boulders in forests, c. 1200m. 
  28. Elatostema euphlebium Merr., PJS 5 c (1910) Bot. 531; --Merr., EPFP 2 (1923) 81; --Type: BS 37765 Ramos & Edaño (US, iso), Luzon: Mountain Province, Mt Pukis, forested slopes, c. 1800m, 10-Mar-1920. LUZON: Mountain Province. Forested slopes, c. 1800m.
  29. Elatostema falcifolium H Schröter, Fedde, Repert. Beih. 83 i (1935) 28; --ibid. 102. PANAY.
  30. Elatostema ferrugineum (Merr.) Co, ined.; --Polychroa ferruginea Merr., PJS 20 (1922) 381; --Merr., EPFP 2 (1923) 77; --Type: BS 38781 Ramos & Edaño (US, iso), Mindanao: Bukidnon prov., Mt Candoon, in forests, c. 1000m, 27-Jul-1920. MINDANAO: Bukidnon (Mt Candoon). Forests, c. 1000m.
  31. Elatostema filicaule C.B.Rob., PJS 5 c (1910) Bot. 516; --Merr., EPFP 2 (1923) 81; --Type: Merrill 6621 (US, iso), Luzon: Benguet prov., Pauai, mossy forest, c. 2100m, May-1909. LUZON: Ifugao (Mt Polis), Benguet (Pauai; Mt Sto Tomas). Mossy forests, 1900-2300m.
  32. Elatostema gitingense Elmer, LPB 8 (1908) 2852; --Merr., EPFP 2 (1923) 81; --Type: Elmer 12421 (US, iso), Sibuyan: Mt Giting-giting, May-1910. SIBUYAN (Mt Giting-giting), PANAY (Antique). On cliffs and in wet soil along small streams in shaded ravines, 250-500m.
  33. Elatostema glaucescens Wedd., Arch. Mus. Paris 9 (1856) 325; --Merr., EPFP 2 (1923) 81. LUZON: Benguet, Bataan (Mt Mariveles), Bulacan (Norzagaray), Rizal (Jalajala), Laguna (Calauan; Los Baños; Mt Makiling), PANAY. Chiefly on mossy boulders and cliffs in shaded stream beds at low and medium elevation; in places ascending to 1200m.
  34. Elatostema glomeratum C.B.Rob., PJS 6 c (1911) Bot. 302; --Merr., EPFP 2 (1923) 82; --Type: Merrill 7642 (TAIF, iso), Luzon: Benguet prov., Baguio, on wet ledges in ravine. LUZON: Benguet (Baguio; Mt Sto Tomas). Wet ledges, shaded ravines, and in the mossy forest, 1400-1900m.
  35. Elatostema gracilifolium Merr., PJS 13 c (1908) Bot. 8; EPFP 2 (1923) 82; --Type: BS 26428 Ramos & Edaño, Luzon: Nueva Ecija prov., Mt Umingan, on rocks in forests, 18-Aug-1916. LUZON: Nueva Ecija (Mt Umingan). On rocks in forests.
  36. Elatostema gracilipes (C.B.Rob.) H Schröter, Fedde, Repert. Beih. 83 i (1935) 3, 18; --ibid. 114.b; --Elatostematoides gracilipes C.B.Rob., PJS 5 c (1910) Bot. 503; --Merr., EPFP 2 (1923) 87; --Type: Merrill 5269 (NY, US, iso), Bucas Grande, 4-Oct-1906. BUCAS GRANDE. Cliffs in shaded ravines at low elevation.
  37. Elatostema halconense C.B.Rob., PJS 5 c (1910) Bot. 540; --Merr., EPFP 2 (1923) 82; --Type: Merrill 5786 (US, iso), Mindoro: Mindoro Oriental prov., Mt Halcon, ridge forest, 750m. MINDORO: Mindoro Oriental (Mt Halcon). Forested ridges, c. 750m.
  38. Elatostema hastatum Elmer, LPB 2 (1908) 466; --Merr., EPFP 2 (1923) 82; --Type: Elmer 9829 (GH, MO, US, iso), Negros: Negros Oriental prov., Cuernos Mtns, Apr-1908. PANAY, NEGROS: Negros Oriental (Cuernos Mtns), MINDANAO: Agusan. Damp forests, c. 1200m. 
  39. Elatostema heterophyllum C.B.Rob., PJS 5 c (1910) Bot. 517; --Merr., EPFP 2 (1923) 82; --Type: Merrill 6911 (US, iso), Negros: Negros Occidental prov., Mt Kanlaon, Apr-1910. NEGROS: Negros Occidental (Mt Kanlaon). In forests on cliffs and in damp shaded ravines, 900-1320m.
  40. Elatostema holophyllum Merr., PJS 11 c (1910) Bot. 5; EPFP 2 (1923) 82; --Type: BS 23329 Ramos (US, iso), Luzon: Sorsogon prov., Mt Bagacaua, growing on trees in the mossy forest, 20-Aug-1915. LUZON: Nueva Ecija, Sorsogon, CATANDUANES. On boulders in shaded ravines at medium elevation, epiphytic in the mossy forests. 
  41. Elatostema inaequifolium Elmer, LPB 3 (1910) 877; --Merr., EPFP 2 (1923) 82; --Type: Elmer 11545 (US, iso), Mindanao: Davao, Mt Apo, Todaya, Aug-1909. MINDANAO: Davao (Mt Apo). On wet rocks along streams, c. 1800m.
  42. Elatostema integrifolium (Don) Wedd. in DC Prod. 161 (1869) 179; --Merr., EPFP 2 (1923) 82. India to Java. PALAWAN, LEYTE (Ormoc, Mt Lamban, PNH 12145 Edano, SING), MINDANAO: Bukidnon (Pagantocan, PNH 13488 Añonuevo, SING). Damp forests at medium elevation.
  43. Elatostema iridense Quisumb., PJS 41 (1930) 320, t. 3; --Type: BS 48515 Ramos & Edaño (PNH, holo, possibly lost; UC?, iso), Luzon: Rizal prov., Mt Irid. On rocks in damp forest, c. 700m. LUZON: Rizal (Mt Irid). Only known from the type collection.
  44. Elatostema kalingaense Merr., PJS 20 (1922) 376; EPFP 2 (1923) 83; --Type: BS 37493 Ramos & Edaño (US, iso), Luzon: Kalinga prov., Mt Masingit, near Lubuagan, on boulders or ledges along streams, c. 1200m, 16-Feb-1920. LUZON: Kalinga. Boulders along streams, c. 1200m.
  45. Elatostema lagunense Merr. ex C.B.Rob., PJS 5 c (1910) Bot. 527; --Merr., EPFP 2 (1923) 83; --Type: Merrill 5118 (US, iso), Luzon: Laguna prov., Los Baños, 12-Mar-1906. LUZON: Ifugao (Kiangan), Bataan (Mt Mariveles), Laguna (Los Baños; Mt Makiling), Quezon (Mt Binuang; Lucban), MINDANAO: Lanao del Sur (Camp Keithley). Damp shaded ravines along small streams at low and medium elevation, ascending to 1000m in wet forests. Photos 
  46. Elatostema lanaense C.B.Rob., PJS 5 c (1910) Bot. 528; --Merr., EPFP 2 (1923) 83; --Type: MS Clemens 405, Mindanao: Lanao del Sur prov., Camp Keithley. Mindanao: Lanao del Sur (Camp Keithley). Shaded ravines, c. 700m.
  47. Elatostema lignosum Merr., PJS 20 (1922) 377; EPFP 2 (1923) 83; --Type: Merrill 9697 (US, iso), Luzon: Benguet prov., Baguio. In shaded places about limestone boulders, c. 1400m. LUZON: Benguet (Baguio), on limestone boulders, c. 1400m.
  48. Elatostema lingua H Schröter, Fedde, Repert. Beih. 83 i (1935) 28; --ibid. 112. SAMAR, MINDANAO.
  49. Elatostema longifolium Wedd., Ann. Sc. Nat. Bot. iv, 1 (1854) 189; --Merr., EPFP 2 (1923) 83; --Type: Cuming 456, Luzon: Laguna prov., Calauan. LUZON: Nueva Vizcaya (Mt Umuguen), Laguna (Los Baños; Calauan), Quezon, MINDANAO. Forests at medium elevation.
  50. Elatostema longipedunculatum Elmer, LPB 3 (1910) 886; --Merr., EPFP 2 (1923) 83; --Type: Elmer 11593 (PNH, holo, lost; MO, US, iso), Mindanao: Davao, Mt Apo, Todaya, Aug-1909. MINDANAO: Davao. On seepage slopes and ledges, c. 1800m.
  51. Elatostema lutescens C.B.Rob., PJS 6 c (1911) Bot. 304; --Merr., EPFP 2 (1923) 83; --Type: Merrill 7643 (US, iso), Luzon: Benguet prov., Baguio and vicinity, in ravines, May-1911. LUZON: Ilocos Norte, Benguet (Baguio; Mt Sto Tomas, Sablan). Shaded cliffs, thickets, and in the mossy forests, 1000-1900m.
  52. Elatostema luzonense C.B.Rob., PJS 5 c (1910) Bot. 512; --Merr., EPFP 2 (1923) 83; --Type: BS 6135 Robinson (US, iso), Luzon: Rizal prov., Montalban, 1-Aug-1908. LUZON: Mountain Province, Benguet, Pampanga (Mt Arayat), Rizal, Laguna, Camarines, PALAWAN (Mt Mantalingahan), PANAY. Common on wet cliffs, banks of ravines, etc., from sea level to at least 1200m. 
  53. Elatostema macgregorii Merr., PJS 14 (1919) 373; EPFP 2 (1923) 83; --Type: BS 32396 RC McGregor (US, iso), Panay: Antique prov., Culasi, in damp forests, 24-May-1918. PANAY: Antique (Culasi, 4 collections cited in the original protologue). Damp forests, 800-1300m.
  54. Elatostema manillense Wedd., Ann. Sc. Nat. Bot. iv, 1 (1854) 189; --H Schröter, Fedde Repert. Beih. 83 ii (1936) 100; --Elatostematoides manillense (Wedd.) C.B.Rob., PJS 5 c (1910) Bot. 501; --Merr., EPFP 2 (1923) 87; --Type: Cuming 786. BATAN, LUZON: Cagayan, Quezon, POLILLO, PANAY: Antique, Capiz, LEYTE, MINDANAO: Misamis, Bukidnon, Agusan. Low and medium elevation forests.
  55. Elatostema merrillii C.B.Rob., PJS 6 c (1911) Bot. 305; --Merr., EPFP 2 (1923) 84; --Type: Merrill 7647 (MO, US, iso), Luzon: Benguet prov., Mt Tonglon (=Mt Sto Tomas), in very shaded places along trail at c. 1800m, May-1911. LUZON: Benguet. Forests, 1800m.
  56. Elatostema microphyllum Elmer, LPB 1 (1908) 286; --Merr., EPFP 2 (1923) 84; --Type: Elmer 9149 (PNH, holo, lost; MO, US, iso), Luzon: Quezon prov., Lucban, May-1907. --Elatostema maquilingense Elmer, LPB 9 (1934) 3229; --Syntypes: Elmer 17636 (GH, MO, US), 18361 (MO), Luzon: Laguna prov., Mt Makiling, Jun/Jul-1917. LUZON: Laguna, Quezon, PANAY. On boulders in damp shaded ravines alomg streams at medium elevation.
  57. Elatostema mindanaense (C.B.Rob.) H Schröter, Fedde, Repert. Beih. 83 i (1935) 18; --ibid. 40; --Polychroa mindanaensis (C.B.Rob.) Merr., EPFP 2 (1923) 78; --Pellionia mindanaensis C.B.Rob., PJS 5 c (1910) Bot. 496; --Type: Copeland 900 (US, iso), Mindanao: Davao, Apr-1904. MINDORO: Mindoro Oriental (Mt Halcon), MINDANAO: Lanao del Sur (Camp Keithley), Bukidnon, Davao, Davao del Sur (Mt Apo).
  58. Elatostema multinervium (Merr.) H Schröter, Fedde, Repert. Beih. 83 i (1935) 18; --ibid. 75; --Polychroa multinervia Merr., PJS 20 (1922) 380; EPFP 2 (1923) 78; --Type: BS 39137 Ramos & Edaño (US, iso), Mindanao: Bukidnon prov., near Tangculan, on trees in damp forests at c. 900m, 5-Jul-1920. MINDANAO: Bukidnon. On trees in damp forests, c. 900m.
  59. Elatostema oblanceolatum C.B.Rob., PJS 5 c (1910) Bot. 524; --Merr., EPFP 2 (1923) 84; --Type: Merrill 6909 (US, iso), Negros: Negros Occidental prov., Mt Kanlaon, on steep moss- covered shaded banks of stream, at 900m, Apr-1910. NEGROS: Negros Occidental (Mt Kanlaon). Shaded mossy stream banks, 900m.
  60. Elatostema obovatum Wedd., Ann. Sc. Nat. Bot. iv, 1 (1854) 190; --Merr., EPFP 2 (1923) 84; --Type: Cuming 628, Luzon: Laguna prov., Calauan. LUZON: Laguna. Shaded ravines at low elevation.
  61. Elatostema obtusiusculum C.B.Rob., PJS 5 c (1910) Bot. 537; --Merr., EPFP 2 (1923) 84; --Type: BS 9840 Robinson, Luzon: Quezon prov., Mahabang Sugsugan River, growing on rocks at an elevation of 100-150m. LUZON: Cagayan, Benguet (Sablan) to Quezon, Camarines Sur (Maagnas), LEYTE, MINDANAO. Low and medium elevation, on rocks and banks in shaded ravines.
  62. Elatostema palawanense C.B.Rob., PJS 5 c (1910) Bot. 526; --Merr., EPFP 2 (1923) 84; --Type: BS 677 Foxworthy (US, iso), Palawan: Mt Victoria, 1100m, Mar/Apr-1906. PALAWAN (Mt Victoria). Damp forested slopes, c, 1100m.
  63. Elatostema panayense Merr., PJS 14 (1919) 372; EPFP 2 (1923) 84; --Type: BS 32396 Ramos & Edaño, Panay: Antique prov., Culasi, damp forests along streams, May-1918. PANAY: Antique. Damp forests along small streams at medium elevation.
  64. Elatostema perlongifolium Elmer, LPB 9 (1934) 3231; --Type: Elmer 16408 (GH, MO, US), Luzon: Sorsogon prov., Mt Bulusan, Jun-1916. LUZON: Sorsogon. Ecol. Photos 
  65. Elatostema philippinense Elmer, LPB 3 (1910) 888; --Merr., EPFP 2 (1923) 84; --Syntypes: Elmer 11746 (MO, US), Elmer 11751 (GH, MO, US), Mindanao: Davao, Mt Apo, Todaya, Sep-1909. --Elatostema glabratum Elmer, LPB 9 (1934) 3228; --Syntypes: Elmer 16325 (GH, MO, US), Luzon: Sorsogon prov., Mt Bulusan, Jun-1916; --Elmer 15576 (GH, MO, US). Luzon: Sorsogon prov., Mt Bulusan, Nov-1915. LUZON: Cagayan, Rizal, Sorsogon (Mt Bulusan), POLILLO, NEGROS, MINDANAO
  66. Elatostema pilosum Merr., PJS 14 (1919) 376; --Merr., EPFP 2 (1923) 84; --Type: BS 20060 RC McGregor, Luzon: Nueva Vizcaya prov., near Imugan, 29-Dec-1912. LUZON: Nueva Vizcaya. In forests at medium elevation.
  67. Elatostema pinnatinervium Elmer, LPB 1 (1908) 286; --Merr., EPFP 2 (1923) 85; --Type: Elmer 9193 (PNH, holo, lost; MO, US, iso), Luzon: Quezon prov., Lucban, May-1907. LUZON: Quezon (Infanta, Anoling River; Lucban), POLILLO (Mt Malulud). Mossy boulders in shaded ravines, 30-750m. 
  68. Elatostema plumbeum C.B.Rob., PJS 5 c (1910) Bot. 535; --Merr., EPFP 2 (1923) 85; --Type: BS 3495 Mearns, Luzon: Benguet prov. LUZON: Benguet. Shaded ravines on cliffs and in the mossy forest, 1000-1900m. 
  69. Elatostema podophyllum Wedd., Ann. Sc. Nat. Bot. iv, 1 (1854) 189; --Merr., EPFP 2 (1923) 85; --Type: Cuming 789 (MO, iso), Luzon. Quezon prov., Mt Banahaw. LUZON: Ifugao, Benguet (Pauai), Quezon (Mt Binuang; Mt Banahaw; Mt Banahaw de Lucban) to Sorsogon (Mt Bulusan), MINDORO: Mindoro Oriental (Mt Halcon). Mossy forests on the higher mountains, up to 2300m.
  70. Elatostema pulchellum C.B.Rob., PJS 5 c (1910) Bot. 522; --Merr., EPFP 2 (1923) 85; --Type: BS 9824 Robinson (US, iso), Luzon: Laguna prov., Mt Banahaw, 5/7-Mar-1910. LUZON: Laguna (Mt Banahaw), Quezon, MINDANAO: Misamis Occidental (Mt Malindang), Davao (Mt Apo). Mossy forests on the higher mountains, 1800-2475m. Photos
  71. Elatostema purpureum C.B.Rob., PJS 6 c (1911) Bot. 306; --Merr., EPFP 2 (1923) 85; --Syntypes: Merrill Phil. Plants 812 (US), Luzon: Benguet prov., Mt Tonglon (=Sto Tomas), mossy forest, c. 1900m, May-1911; --Merrill 7644: also from the same mountain but at a more shaded place at a slightly lower elevation. LUZON: Ifugao, Mountain Province, Benguet, Mossy forests, 1800-1900m.
  72. Elatostema rigidum Wedd., Arch. Mus. Paris 9 (1856) 320; --Type: Cuming 1674 (cited by Robinson in PJS 5 c (1910) 503), Samar. --Elatostematoides rigidum (Wedd.) C.B.Rob., PJS 5 c (1910) Bot. 503; --Merr., EPFP 2 (1923) 88. LUZON: Cagayan, CATANDUANES, PANAY, NEGROS: Negros Occidental (Himugaan River), BOHOL (BS 42872 Ramos, SING), LEYTE, SAMAR, MINDANAO: Zamboanga, Agusan del Norte (Mt Urdaneta).
  73. Elatostema robinsonii Merr., PJS 14 (1919) 376; EPFP 2 (1923) 85; --Type: BS 19727 RC McGregor (US, iso), Luzon: Ifugao prov., Mt Polis, 30-Jan-1913. LUZON: Ifugao (Mt Polis), apparently in mossy forests at or above 1500m.
  74. Elatostema samarense (Merr.) H Schröter, Fedde, Repert. Beih. 83 i (1935) 28; --ibid. 108; --Elatostematoides samarense Merr., PJS 14 (1919) 380; EPFP 2 (1923) 88; non Elatostema samarense Merr., PJS 20 (1922) 378; --Type: BS 17468 Ramos (US, iso), Samar: Western Samar prov., Yabong (=Yabon), damp forests near small streams at low elevation, 16-Mar-1914. --Elatostema trichophlebium Merr. ex H. Schröter, Fedde, Repert. Beih. 83 i (1935) 28; --ibid. 104; --Type: BS 42378 Edaño (A, SING, iso), Panay: Aklan (previously in Capiz) prov., Mt Timbaban, May-1923. PANAY: Aklan, SAMAR. Lowland primary forest along streams.
  75. Elatostema scapigerum C.B.Rob., PJS 5 c (1910) Bot. 542; --Merr., EPFP 2 (1923) 85; --Type: Vanoverbergh 587. LUZON: Mountain Province, Benguet (Baguio; Mt Sto Tomas). Damp cliffs and in the mossy forest, 1100-2000m.
  76. Elatostema serratifolium Elmer, LPB 9 (1934) 3234; --Type: Elmer 14712 (PNH, holo, lost; GH, MO, US, iso), Luzon: Sorsogon prov., Mt Bulusan, Aug-1916. LUZON: Sorsogon. Ecol.
  77. Elatostema simulans C.B.Rob., PJS 5 c (1910) Bot. 519; --Merr., EPFP 2 (1923) 86; --Type: BS 8145 Ramos (US, iso), Luzon: Nueva Vizcaya prov., Bayombong, May-1909. --Elatostema pacificum Elmer, LPB 9 (1934) 3230; --Syntypes: Elmer 14613 (MO, US), Luzon: Sorsogon prov., Mt Bulusan, Oct-1915; --Elmer 16173 (MO, US). Also from Mt Bulusan, May-1916; --Elmer 16968 (GH, MO, US). Also from Mt Bulusan, Aug-1916. LUZON: Benguet to Sorsogon. Damp cliffs and in forests at low and medium elevation.
  78. Elatostema sinuatum (Blume) Hassk., Cat. Hort. Bogor. (1844) 79; --Polychroa sinuata (Blume) Merr., PJS 20 (1922) 382; --Merr., EPFP 2 (1923) 78; --H. Schröter, Fedde, Repert. Beih. 83 i (1935) 29, descr. ampl.; --Pellionia sinuata (Blume) Boerl., Handl. Kenn. Fl. Ned. Ind. 3 (1900) 375; --C.B.Rob., PJS 5 c (1910) Bot. 497; --Procris sinuata Blume, Bijdr. (1825) 511; --Type from Java. --Elatostema laciniatum Elmer, LPB 1 (1908) 287; --H Schröter, Fedde, Repert. Beih. 83 ii (1936) 36; --Type: Elmer 9196 (US, iso), Luzon: Quezon prov., Lucban, May-1907. --Polychroa urdanetensis Elmer, LPB 9 (1934) 3241; --Type: Elmer 13491 (US, iso), Mindanao: Agusan del Norte prov., Mt Urdaneta, Aug-1912. Java, Philippines, Sulawesi, Moluccas. C LUZON to MINDANAO. Damp forests at low and medium elevation. Exceedingly polymorphic. Photos 
  79. Elatostema sorsogonense Elmer, LPB 9 (1934) 3235; --Syntypes: Elmer 16322 (GH, MO, US), Luzon: Sorsogon prov., Mt Bulusan, Jun-1916; --Elmer 14614 (MO), Luzon: Sorsogon prov., Mt Bulusan, Oct-1916. LUZON: Sorsogon.
  80. Elatostema spinulosum Elmer, LPB 2 (1908) 468; --Merr., EPFP 2 (1923) 86; --Type: Elmer 9776 (PNH, holo, lost; MO, US, iso), Negros: Negros Oriental prov., Cuernos Mtns, Apr-1908. PANAY, NEGROS. Damp forests, 600-1000m. Photos 
  81. Elatostema sublaxum (Elmer) H Schröter, Fedde, Repert. Beih. 83 i (1935) 28; --ibid. 105; --Elatostematoides sublaxum Elmer, LPB 8 (1915) 2858; --Type: Elmer 13493 (A, MO, NA, NY, US, iso), Mindanao: Agusan del Norte prov., Mt Urdaneta, Aug-1912. --Elatostematoides laxum (Elmer) C.B.Rob., PJS 5 c (1910) Bot. 502; --Merr., EPFP 2 (1923) 87; -- Elatostema laxum Elmer, LPB 2 (1908) 465; --Type: Elmer 10337 (PNH, holo, lost; A, MO, US, iso), Negros: Negros Oriental prov., Cuernos Mtns, Jun-1908. LUZON: Quezon, Sorsogon, PANAY, NEGROS, LEYTE, SAMAR, MINDANAO. Low and medium elevation damp forests. Photos
  82. Elatostema sublignosum C.B.Rob., PJS 5 c (1910) Bot. 541; --Merr., EPFP 2 (1923) 86; --Type: BS 9754 Robinson (L, US, iso), Luzon: Laguna prov., Mt Banahaw, 5/7-Mar-1910. LUZON: Laguna (Mt Banahaw; Mabalucbaluc), Forests, 600-800m.
  83. Elatostema suborbiculare Merr., PJS 14 (1919) 377; EPFP 2 (1923) 86; --Type: BS 19729 RC McGregor (US, iso), Luzon: Ifugao prov., Mt Polis, apparently in the mossy forest. Feb-1913. LUZON: Ifugao (Mt Polis). Apparently in the mossy forest.
  84. Elatostema surigaoense Elmer, LPB 8 (1915) 2854; --Merr., EPFP 2 (1923) 86; --Type: Elmer 13637 (GH, MO, US, iso), Mindanao: Agusan del Norte prov., Mt Urdaneta, Aug-1912. MINDANAO: Agusan del Norte. Forests, c. 1000m.
  85. Elatostema umbrinum Elmer, LPB 8 (1915) 2855; --Merr., EPFP 2 (1923) 86; --Type: Elmer 13831, Mindanao: Agusan del Norte prov., Mt Urdaneta. --Elatostema urdanetense Elmer, LPB 8 (1915) 2857; --Type: Elmer 14249 (MO, iso), Mindanao: Agusan del Norte prov., Mt Urdaneta, Sep-1912. MINDANAO: Agusan del Norte. Dense forests, c. 1600m.
  86. Elatostema variabile C.B.Rob., PJS 5 c (1910) Bot. 514; ibid. 6 c (1911) Bot. 307, t. 1; --Merr., EPFP 2 (1923) 86; --Type: Vanoverbergh 528, Luzon: Mountain Province, Bauco, in forests, 1650m. LUZON: Mountain Province. Forests, c. 1650m. Photos 
  87. Elatostema variegatum C.B.Rob., PJS 5 c (1910) Bot. 538; ibid. 6 c (1911) Bot. 31; --Merr., EPFP 2 (1923) 86; --Type: BS 9767 Robinson, Luzon: Laguna prov., base of Mt Banahaw, on rocks, 800m. LUZON: Benguet (Sablan), Laguna (base of Mt Banahaw). Lithophytic in shaded stream beds, c. 800m.
  88. Elatostema viridescens Elmer, LPB 1 (1908) 285; --Merr., EPFP 2 (1923) 87; --Type: Elmer 8076. Luzon: Laguna prov., Los Baños. LUZON: Pampanga (Mt Pinatubo), Bataan, Laguna, Cavite, Quezon, Sorsogon, PANAY. Low and medium elevation along streams and damp forests.
  89. Elatostema volubile (Elmer) H. Schröter, Fedde, Repert. Beih. 83 i (1935) 18; --ibid. 74; --Polychroa volubilis (Elmer) Merr., EPFP 2 (1923) 78; --Procris volubilis Elmer, LPB 8 (1915) 2867; --Synypes: Elmer 13898 (MO, NA, NY), Mindanao: Agusan del Norte prov., Mt Urdaneta, Sep-1912; Elmer 13856 (not seen). Same area as preceding. POLILLO, LEYTE, SAMAR, CAMIGUIN, MINDANAO: Zamboanga, Bukidnon, Agusan del Norte. Low and medium elevation damp forests, ascending to 1400m. 
  90. Elatostema wenzelii H Schröter, Fedde, Repert. Beih. 83 i (1935) 28; --ibid. 104. MINDANAO.
  91. Elatostema whitfordii Merr., PJS 1 suppl. (1906) 48; --Merr., EPFP 2 (1923) 87; --Syntypes: Whitford 254 (US), Luzon: Bataan prov., Mt Mariveles, May-1904; FB 1234 Borden (US). Luzon: Bataan prov., Mt Mariveles, Jun-1904, wet shaded banks subject to constant drip of water in canon of the Lamao River, 550-800m. LUZON: Bataan, MINDANAO: Davao (Mt McKinley, PNH 1045 Edano, SING). Densely shaded ravines along streams, at and above 600m. 
  92. Elatostema zamboangense Merr., PJS 14 (1919) 374; --Merr., EPFP 2 (1923) 87; --Type: Merrill 8228 (US, iso), Mindanao: Zamboanga, Sax River, back of San Ramon, damp shaded ravine, c. 200m, 27-Nov-1911. MINDANAO: Zamboanga. Damp shaded ravines, c. 200m.
  93. Elatostema catubigense ined. --Elatostema samarense Merr., PJS 20 (1922) 378, non Elatostematoides samarense Merr., 1919, which is now =Elatostema samarense (Merr.) Schroter; --Type: BS 34117 Ramos, Samar: Catubig River, in damp forest at low elevation, Feb-1916. SAMAR. Damp forests at low elevation.

Non-naturalized species

  1. Elatostema repens (Lour.) Hallier f. in H Schröter, Fedde, Repert. Beih. 83 i (1935) 25; --Polychroa repens Lour., Fl. Cochin. (1790) 559; --Merr., EPFP 2 (1923) 78. Native of Myanmar and Indochina. 

Doubtful taxon

  1. Elatostema tomentosum Wedd., Ann. Sc. Nat. Bot. iv, 1 (1854) 189; --Merr., EPFP 2 (1923) 86; --Type: Callery s.n. (P, fide Merrill), Luzon: probably Benguet. --Elatostema sessile Forst. var. tomentosum Wedd. in DC, Prodr. 161 (1869) 173. --Elatostema subincanum Wedd., Arch. Mus. Paris 9 (1856) 295, A species of doubtful status.

Possible new species (ined.) materials at PNH

  1. Elatostema lepantoense Merr. 

 Elatostematoides C.B.Rob. = Elatostema

 Fleurya Gaudich. = Laportea

 Gonostegia Turcz. = Pouzolzia

Laportea Gaudich.

Photos
  1. Laportea interrupta (L.) Chew, Gard. Bull. Sing. 25 (1969) 200; --Yang, et al., Fl. Taiwan 2nd ed., 2 (1996) 226; --Fleurya interrupta (L.) Wight, Icon. Pl. Ind. Or. 6 (1853) 10, t. 1975; --Merr., Fl. Manila (1912) 182; EPFP 2 (1923) 75. Tropical and subtropical Africa and Asia and the Pacific Islands. Photos 
  2. Laportea ruderalis (G.Forst.) Chew, Gard. Bull. Sing. 21 (1965) 201; --Fleurya ruderalis (G Forst.) Wedd., Ann. Sc. Nat. ser. 4, 1 (1854) 182; --Merr., EPFP 2 (1923) 75. Christmas Is., Philippines, Sulawesi, Lesser Sunda Isls., Moluccas, New Guinea, Bismarck Archipelago to Melanesia, Micronesia and Polynesia. In the Philippines recorded from CAVILLI IS., LUZON: Bulacan, Angat.

Lecanthus Wedd.

Photos
  1. Lecanthus peduncularis (Wall. ex Royle) Wedd. in DC. Prod. 161 (1869) 164, pro parte; --Merr., EPFP 2 (1923) 77; --Yang, et al., Fl. Taiwan 2nd ed., 2 (1996) 226, pl. 113. Africa, S Asia, China, Taiwan, Philippines, Java, and the Pacific islands. LUZON: Ifugao (Mt Polis), Benguet (Pauai; Mt Pulag), MINDANAO: Davao (Mt Apo). On the higher mountains above 1900m. Photos 

Leucosyke Zoll. & Moritzi

Photos
  1. Leucosyke arcuatovenosa Unruh in Engl., Bot. Jahrb. 73 (1943) 246. LUZON.
  2. Leucosyke aspera C.B.Rob., PJS 6 c (1911) Bot. 23; --Merr., EPFP 2 (1923) 96; --Type: BS 2915 Mearns, Luzon: Albay prov., Mt Mayon. LUZON: Camarines, Albay, SIBUYAN, PANAY. Forests, c. 1000m. Photos
  3. Leucosyke benguetensis Unruh, in Engl., Bot. Jahrb. 73 (1943) 247. LUZON: Ilocos Norte (Mt Pico de Loro, PNH 17997, SING), Apayao (Mt Duraragan, PNH 19884 Edano, SING). Photos
  4. Leucosyke brunnescens C.B.Rob., PJS 6 c (1911) Bot. 25; --Merr., EPFP 2 (1923) 96; --Unruh, Bot. Jahrb. 73 (1943) 243; --Type: BS 7821 Ramos, Luzon: Cagayan prov., Maunan. LUZON: Ilocos Sur (Sugpon), Cagayan, Isabela (San Mariano, BS 47163, SING), Nueva Vizcaya (Dupax, BS 11388 McGregor, SING), Pangasinan, Bataan (Mt Mariveles, Lamao River, Merrill 2544, SING). Low and medium elevation thickets. Photos
  5. Leucosyke buderi Unruh in Engl., Bot. Jahrb. 73 (1943) 245. LUZON
  6. Leucosyke capitellata (Poir.) Wedd. in DC Prodr. 161 (1869) 235 27; --Merr., EPFP 2 (1923) 96. --Leucosyke capitellata var. celtidifolia (Gaudich.) Wedd. in DC Prodr. 161 (1869) 235 26; --Merr., EPFP 2 (1923) 97. --Leucosyke palawanensis Unruh in Engl., Bot. Jahrb. 73 (1943) 229; --Type: Elmer 13247 (GH, isosyn), Palawan: Puerto Princesa, Mt Pulgar (=Thumb Peak), May 1911. --Leucosyke leytensis Merr., PJS 10 c (1915) Bot. 269; EPFP 2 (1923) 97; --Type: Wenzel 1061 (GH, US, iso), Leyte: Buenavista, Jaro, in forests, c. 500m, 20-Aug-1914. --Leucosyke angusta Unruh, PJS 84 (1955) 167; --Type: PNH 6480 Castro & Añonuevo (A, holo), Polillo: along Anibawan River, sea level, 28-Nov-1948. --Leucosyke samarensis Unruh, PJS 84 (1955) 172; --Type: PNH 6334 Sulit and Coñese (A, holo), Samar: Western Samar prov., Bagacay, Concord, open place near road km 58.5 near bridge, 2nd growth forest, 200m, 20-Jun-1950. Taiwan, Philippines, Java to New Guinea. Throughout the Philippines: LUZON: Ilocos Norte, Mountain Province, Benguet, Cagayan, Isabela, Nueva Vizcaya, Pangasinan, Bataan, Pampanga, Rizal, Laguna, Cavite, Batangas, Quezon, Camarines, Albay, Sorsogon, POLILLO, MINDORO, PALAWAN, MASBATE, PANAY, NEGROS, CEBU, BOHOL, LEYTE, SAMAR, BASILAN, MINDANAO: Zamboanga, Lanao del Sur, Davao. In thickets, second-growth forests, etc. at low and medium elevation. In the Cordillera Highlands ascending to at least 2000m. Common. Photos
  7. Leucosyke elmeri Unruh in Engl., Bot. Jahrb. 73 (1943) 246. MINDANAO.
  8. Leucosyke hispidissima (Wedd.) Miq. Fl. Ind. Bat.2 (1858-59) 265; --Merr., EPFP 2 (1923) 97; --Missiessya hispidissima Wedd., Ann. Sci. Nat. Bot. IV, 1 (1854) 195; --Type: Callery s.n., Luzon: ‘in insulae Manillae montibus Igorrotes’ (=possibly Benguet prov. not far from Baguio). --Leucosyke magallanensis Elmer, LPB 2 (1910) 677; --Merr., EPFP 2 (1923) 97; --Type: Elmer 12282 (PNH, holo, lost; GH, MO, US, iso), Sibuyan: Mt Giting-giting, Magallanes (=Magdiwang), Apr-1910. LUZON: Mountain Province, Benguet, Nueva Ecija, SIBUYAN, PANAY: Antique (BS 32516 McGregor, SING). Thickets, forested river banks up to c. 1400m. Photos
  9. Leucosyke mindorensis C.B.Rob., PJS 6 c (1911) Bot. 22; --Merr., EPFP 2 (1923) 98; --Type: FB 11461 Merritt (US, iso), Mindoro: Mindoro Occidental prov., Amnay River, 600m, Apr/May-1908. MINDORO: Mindoro Occidental. Forests, c. 600m.
  10. Leucosyke negrosensis C.B.Rob., PJS 6 c (1911) Bot. 27; --Merr., EPFP 2 (1923) 98; --Type: FB 4316 Everett (US, iso), Negros: Negros Occidental prov., Pioc River, in dense forest. PALAWAN, PANAY, NEGROS, MINDANAO: Agusan. Low and medium elevation forests. Photos 
  11. Leucosyke nivea C.B.Rob., PJS 6 c (1911) Bot. 21; --Merr., EPFP 2 (1923) 98; --Type: MS Clemens sn, Mindanao: Lanao del Sur prov., Camp Keithley, Sep-1907. LEYTE, MINDANAO. Forests, 500-800m. 
  12. Leucosyke ovalifolia C.B.Rob., PJS 6 c (1911) Bot. 26; --Merr., EPFP 2 (1923) 98; --Type: Merrill 481, Culion. --Leucosyke merrillii Unruh in Engl., Bot. Jahrb. 73 (1943) 225; --Type: Merrill 706 (GH, iso), Palawan: E- wi-ig (=Iwahig) River. LUZON: Zambales, CULION, PALAWAN (Iwahig), PANAY: Capiz (Mt Timbaban, BS 42489 Edaño, SING). Low elevation dry thickets and open forests. Photos
  13. Leucosyke quadrinervia C.B.Rob., PJS 6 c (1911) Bot. 28; --Merr., EPFP 2 (1923) 98; --Yang, et al., Fl. Taiwan 2nd ed., 2 (1996) 228. Lanyu, Lutao and N Philippines. BATANES (SABTAN), BABUYAN ISLS (CAMIGUIN). Along streams at low elevation. Closely allied to L. capitellata (Poir.) Wedd. Photos
  14. Leucosyke rizalensis Unruh in Engl., Bot. Jahrb. 73 (1943) 247. LUZON: Rizal
  15. Leucosyke weddelii Unruh in Engl., Bot. Jahrb. 73 (1943) 226.

Maoutia Wedd.

Photos
  1. Maoutia setosa Wedd. in Ann. Sc. Nat. Bot. iv, 1 (1854) 194; --Merr., EPFP 2 (1923) 98; --Yang, et al., Fl. Taiwan 2nd ed., 2 (1996) 229. Ryukyus, Lanyu, Lutao and the Philippines. N LUZON to MINDANAO. Forests and thickets, in some regions at low and medium elevation; ascending to 2200m in Benguet. Photos

Musanga C.Sm. ex R.Br. [Not native]

Photos
  1. Musanga cecropioides R Br. Native of Congo, now pantropic in cultivation. Recently introduced into forest plantations in the Philippines (Pancho, Fl. Mt Makiling 1 (1983) 121. Photos

Neodistemon Babu & A.N.Henry

No photos
  1. Neodistemon indicum (Wedd.) Babu & AN Henry, Taxon 19 (1970) 651; --Distemon indicus Wedd., Arch. Mus. Paris 9 (1857) 551, t. 20, A; --Merr., EPFP 2 (1923) 93 India (Assam) to Myanmar, Java, Philippines. LUZON: Rizal (Antipolo; Morong). Open slopes at low elevation.

Nothocnide Blume

No photos
  1. Nothocnide discolor (C.B.Rob.) Chew, Gard. Bull. Sing. 24 (1969) 368; --Pipturus discolor C.B.Rob., PJS 6 c (1911) Bot. 15; --Merr., EPFP 2 (1923) 95; --Type: MS Clemens 676 (NY, iso), Mindanao: Lanao del Sur prov., Camp Keithley, Sep/Oct-1906. New Guinea and Philippines. PANAY, MINDANAO: Lanao del Sur (along Lake Lanao), Bukidnon, Agusan, Davao. Forests, 700m and above.
  2. Nothocnide repanda (Blume) Blume, Ann. Mus. Bot. Lugd.-Bat. 2 (1856) 137, t. 14; --Pipturus repandus (Blume) Wedd. in Arch. Mus. Paris 9 (1856) 448; --Merr., EPFP 2 (1923) 95; --Urtica repanda Blume, Bijdr. (1825) 501; --Type from Java. --Pipturus subalpinus Elmer, LPB 9 (1934) 3221; --Type: Elmer 22016 (MO, US, iso), Luzon: Pampanga prov., Mt Pinatubo, Camp Stotsenburg, May-1927. Sumatra, Borneo, Philippines, Sulawesi, Java, Bali, Moluccas, New Guinea, Bismarck Archipelago and Solomon Islands. LUZON: Mountain Province to Sorsogon, MINDORO, PANAY, LEYTE, MINDANAO: Davao (Mt Apo). Damp forested ravines near streams, at low and medium elevation. In Mountain Province sprawling over ledges and boulders up to 1600m. 

Oreocnide Miq.

Edited by Pieter B. Pelser, 22 June 2013

Photos
  1. Oreocnide rubescens (Blume) Miq., Ann. Mus. Bot. Lugd.-Bat. 4 (1869) 306; --Merr., EPFP 2 (1923) 96; --Urtica rubescens Blume, Bijdr. (1825) 506; --Villebrunea rubescens (Blume) Blume, Ann. Mus. Bot. Lugd.-Bat. 2 (1856) 167. Java, Philippines, Sulawesi, Moluccas. LUZON: Laguna, Quezon to MINDANAO. Damp shaded ravines, near streams at low and medium elevation. Photos
  2. Oreocnide trinervis (Wedd.) Miq. Fl. Ind. Bat. 12 (1858-59) 270; --Merr., EPFP 2 (1923) 96; --Yang, et al., Fl. Taiwan 2nd ed., 2 (1996) 230; --Villebrunea trinervis Wedd., Ann. Sc. Nat. Bot. iv, 1 (1854) 196; --Type: Cuming 777 (MO, NY, iso), Luzon: Quezon prov. Lanyu, Philippines, Borneo and Java. BATAN ISLS to MINDANAO. Thickets and forests, especially along streams at low and medium elevation. Photos

Pellionia Gaudich. = Elatostema

Pilea Lindl.

Photos
  1. Pilea apoensis Elmer, LPB 3 (1910) 882; --Merr., EPFP 2 (1923) 76; --Type: Elmer 11551 (L, NY, iso), Mindanao: Davao, Mt Apo, Todaya, on moss-covered wet ledges along Cati Creek at 5750ft, Aug-1909. MINDANAO: Bukidnon, Davao (Mt Apo). Forests, c. 1700m.
  2. Pilea benguetensis C.B.Rob., PJS 5 c (1910) Bot. 488; --Merr., EPFP 2 (1923) 76; --Type: Elmer 6304 (MO, NY, US, iso), Luzon: Benguet prov., Baguio, May-1904. LUZON: Mountain Province, Benguet (Baguio; Mt Pulag), Nueva Vizcaya. On limestone boulders, damp shaded ravines, cliffs, etc., 1300-2000m.
  3. Pilea calcicola C.B.Rob., PJS 5 c (1910) Bot. 493; --Merr., EPFP 2 (1923) 76; --Type: BS 9529 CB Robinson (L, NY, US), Luzon: Rizal prov., Montalban, 2-Nov-1909. LUZON: Bulacan, Rizal. Low and medium elevation forests on limestone, on limestone cliffs and gorges. 
  4. Pilea dataensis C.B.Rob., PJS 5 c (1910) Bot. 494; --Merr., EPFP 2 (1923) 76; --Type: Merrill 4602 (US, iso), Luzon: Mountain Province, Mt Data, mossy forest, 2100m, Nov-1905. LUZON: Mountain Province (Mt Data), Ifugao (Mt Polis). Mossy forests, c. 2100m. 
  5. Pilea humilis C.B.Rob., PJS 5 c (1910) Bot. 488; --Merr., EPFP 2 (1923) 76; --Type: Merrill 6912 (L, NY, US, iso), Negros: Negros Occidental prov., Mt Kanlaon, Apr-1910. LUZON: Mountain Province, Benguet, NEGROS: Negros Occidental (Mt Kanlaon). In ravines, in the mossy forest, and on open peaks of the higher mountains, 1200-2800m.
  6. Pilea intumescens C.B.Rob., PJS 5 c (1910) Bot. 492; --Merr., EPFP 2 (1923) 76; --Type: MS Clemens 1223 (NY, US, iso), Mindanao: Lanao del Sur prov., along Lake Lanao, Camp Keithley, Sep/Oct-1907. PANAY, MINDANAO: Lanao del Sur. Damp shaded ravines, c. 700m.
  7. Pilea loheri Merr., PJS 27 (1925) 23; EPFP 4 (1925) 252; --Type: Loher 12797, Luzon: Rizal prov., Balintingan, Jul-1909. LUZON: Rizal (Balintingan).
  8. Pilea luzonensis Merr., PJS 1 suppl. (1906) 48; --Merr., EPFP 2 (1923) 76; --Syntypes: Whitford 279 (US), Luzon: Bataan prov., Mt Mariveles, Lamao River, May-1904; Whitford 1129 (US), Luzon: Bataan prov., Lamao River, Mar-1905. LUZON: Benguet (Baguio; Mt Sto. Tomas), Nueva Vizcaya (Caraballo Sur), Bataan (Mt Mariveles), Rizal, MINDANAO: Agusan del Norte (Mt Urdaneta). Damp shaded ravines along streams at medium elevation, ascending to 1400m.
  9. Pilea melastomoides (Poir.) Wedd., Ann. Sc. Nat. Bot. ser. 4, 1 (1854) 186; --Merr., EPFP 2 (1923) 76; --Yang, et al., Fl. Taiwan 2nd ed., 2 (1996) 242, pl. 122; --Urtica melastomoides Poir. in Lam., Encycl. Suppl. 4 (1816) 223. India, Sri Lanka, Vietnam, China, Taiwan, Philippines, Java. LUZON: Kalinga, Ifugao, Mountain Province (Mt Data), Benguet (Mt Pulag; Pauai; Mt Sto Tomas), Laguna (Mt Banahaw). Mossy forests, 1800-2300m. Photos
  10. Pilea microphylla (L.) Liebm., Vidensk. Selsk. Skr. 52 (1851) 302; --Merr., Fl. Manila (1912) 76; EPFP 2 (1923) 76. Native to S America, widely naturalized in tropical and subtropical areas of the world. Throughout the Philippines in and about towns, on damp walls, etc. Photos
  11. Pilea monticola C.B.Rob., PJS 5 c (1910) Bot. 489; --Merr., EPFP 2 (1923) 77; --Type: BS 5533 Ramos, Luzon: Benguet prov., Rio Trinidad (=Balili River); Paratype: BS 5566 Ramos (US). Luzon: Benguet prov., Rio Trinidad (=Balili River), Dec 1908. LUZON: Mountain Province, Benguet (Mt Pulag; Mt Sto. Tomas; La Trinidad). Mossy forests, damp shaded ravines, etc., 1200-2300m.
  12. Pilea peploides (Gaudich.) Hook. & Arn., Bot. Beechey Voy. (1832) 96; --Hatusima, Mem. Fac. Agr. Kagoshima Univ. 5 (1966) 27.
  13. Pilea rigida C.B.Rob., PJS 5 c (1910) 491; --Merr., EPFP 2 (1923) 77; --Type: BS 8181 Ramos (NY, US, iso), Luzon: Nueva Vizcaya prov., Mt Dalemdim, 1500m, 19-May-1909. LUZON: Ifugao, Nueva Vizcaya. c. 1500m.
  14. Pilea robinsonii Elmer, LPB 3 (1910) 880; --Merr., EPFP 2 (1923) 77; --Syntypes: Elmer 10487 (MO, NY 2 sheets, US), Mindanao: Davao, Mt Apo, Todaya, black humus-covered soil of moist dense woods at 3750ft, May-1909; --Elmer 11758 (MO, NY 2 sheets, US), Mindanao: Davao, Mt Apo, Todaya, among boulders below the Colan Creek Falls and at its junction with the Baruring River at 3750ft, Sep-1909. MINDANAO: Bukidnon, Davao (Mt Apo). Damp forests, c. 1000m.
  15. Pilea sylvatica Elmer, LPB 3 (1910) 879; --Merr., EPFP 2 (1923) 77; --Syntypes: Elmer 11698 (MO, NY 2 sheets, US), Mindanao: Davao, Mt Apo, Todaya, on moist, fertile humus-covered soil of densely wooded flats at 4000ft, Sep-1909; --Elmer 11547 (MO, NY, US), Mindanao: Davao, Mt Apo, Todaya, on moss-covered seepage ledges along the Cati Creek at 5750ft, above its falls of Mt Apo, Aug-1909. MINDANAO: Bukidnon, Davao (Mt Apo). Damp forests and on wet cliffs, 1200-1800m. Photos

Pipturus Wedd.

Edited by Pieter B. Pelser, 22 October 2013

Photos
  1. Pipturus angustifolius Merr., PJS 20 (1922) 379; EPFP 2 (1923) 94; --Type: BS 15320 Ramos (US, iso), Leyte: Dagami, thickets along streams at low elevation, 6-Aug-1912. LEYTE. Thickets along streams at low elevation.
  2. Pipturus arborescens (Link) C.B.Rob., PJS 6 c (1911) Bot. 13; --Merr., EPFP 2 (1923) 94; --Yang, et al., Fl. Taiwan 2nd ed., 2 (1996) 253; --Urtica arborescens Link, Enum. Hort. Berol. 2 (1822) 386; --Blanco Fl. Filip. ed. 2 (1845) 483, ed. 3, 3 (1879) 100, t. 371. --Pipturus asper Wedd., Ann. Sci. Nat. ser. 4, 1 (1854) 197. Ryukyus, Taiwan (Lanyu and Lutao), the Philippines and Borneo. Very common and widely distributed in all islands and provinces. Chiefly in low and medium elevation thickets and second-growth forests, ascending to 2000m in Benguet. Photos
  3. Pipturus argenteus (G Forst.) Wedd. in DC, Prodr. 161 (1869) 23519; --Merr., EPFP 2 (1923) 94; --Urtica argentea G Forst., Prodr. (1784) 65. Sumatra to Australia and Polynesia. BALABAC, LUMBUCAN (near Balabac), MINDANAO: Surigao. On sandy soil, open forests near the sea.
  4. Pipturus dentatus (C.B.Rob.) C.B.Rob., PJS 6 c (1911) Bot. 311; --Merr., EPFP 2 (1923) 95; --Pouzolzia dentata C.B.Rob., PJS 6 c (1911) 7; --Type: Merrill 4741 (NY, US, iso), Luzon: Benguet prov., Pauai, Oct/Nov-1905. LUZON: Ifugao (Mt Polis), Benguet (Mt Pulag; Pauai; Mt Sto Tomas). Open slopes, thin pine forests, etc., 1900-2300m.
  5. Pipturus succulentus Elmer, LPB 3 (1910) 897; --Merr., EPFP 2 (1923) 95; --Type: Elmer 10739 (PNH, holo, lost; MO, NA, US, iso), Mindanao: Davao, Mt Apo, Todaya, May-1909. MINDANAO: Davao (Mt Apo). Thickets along Baruring River, c. 1000m.

Poikilospermum Zipp. ex Miq.

Photos
  1. Poikilospermum acuminatum (Trec.) Merr., Contr. Arnold Arb. 8 (1934) 49; --Conocephalus acuminatus Trec., Ann. Sci. Nat. Bot. 3, 8 (1847) 91; --Merr., EPFP 2 (1923) 70. --Poikilospermum diffusum (Merr.) Merr., Contr. Arnold Arb. 8 (1934) 49; --Conocephalus diffusus Merr., PJS 9 c (1914) Bot. 355; EPFP 2 (1923) 70. --Conocephalus diffusus var. obtusus Merr., EPFP 2 (1923) 70. --Poikilospermum grande (Wedd.) Merr., Contr. Arnold Arb. 8 (1934) 50; --Conocephalus grandis (Wedd.) Merr., EPFP 2 (1923) 70; --Procris grandis Wedd., Arch. Mus. Paris 8 (1855-56) 337; --Vidal, Rev. Pl. Vasc. Filip. (1886) 256; Phan. Cuming. Philip. (1885) 146; --Type: Cuming 1730 (K). --Poikilospermum grandifolium (Warb.) Merr., Contr. Arnold Arb. 8 (1934) 50; --Conocephalus grandifolius Warb. in Perkins, Fragm. Fl. Philip. (1904) 167; --Merr., EPFP 2 (1923) 71. --Poikilospermum molle (Merr.) Merr., Contr. Arnold Arb. 8 (1934) 51; --Conocephalus mollis Merr., PJS 18 (1921) 51; EPFP 2 (1923) 71; --Type: BS 17582 Ramos. --Poikilospermum warburgii (Elmer) Merr., Contr. Arnold Arb. 8 (1934) 52; --Conocephalus warburgii Elmer, LPB 1 (1908) 278; --Merr., EPFP 2 (1923) 72; --Type: Elmer 8973. BABUYAN ISLS, LUZON (widespread), CATANDUANES, PANAY, LEYTE, SAMAR, MINDANAO. Lowland to montane forests (including mossy) and thickets. Photos
  2. Poikilospermum erectum (Blanco) Merr., Contr. Arnold Arb. 8 (1934) 49; --Conocephalus erectus (Blanco) Fern.-Villar, Novis. App. (1880) 203; --Merr., Govt. Lab. Publ. (Philip.) 27 (1905) 80; Sp. Blancoanae (1918) 129, excl. syn. C. grandifolius Warb.; EPFP 2 (1923) 70; --Procris erecta Blanco, Fl. Filip. (1837) 707; ed. 2 (1845) 490; ed. 3, 3 (1879) 111; --Type: Not extant.  “Illustrative specimen”: Merrill Sp. Blancoanae 871, 960. LUZON: Cagayan, Isabela, Rizal, Cavite, Laguna, Quezon. Lowland and medium elevation forests.
  3. Poikilospermum oblongifolium (Barg.-Petr.) Merr., Contr. Arnold Arb. 8 (1934) 51. Borneo, S Philippines. BASILAN, MINDANAO: Bukidnon (Mt Kitanglad). Lowland riverine forests to as high as 1800m.
  4. Poikilospermum suaveolens (Blume) Merr., Contr. Arnold Arb. 8 (1934) 47; --Conocephalus suaveolens Blume, Bijdr. (1825) 483; --Merr., EPFP 2 (1923) 71. --Poikilospermum piperi (Elmer) Merr., Contr. Arnold Arb. 8 (1934) 51; --Conocephalus piperi Elmer, LPB 8 (1915) 2770; --Merr., EPFP 2 (1923) 71. India to Malaya. N LUZON to MINDANAO. Lowland and medium elevation forests (Merr., 1923). Photos

 Polychroa Lour. = Pellionia = Elatostema

Pouzolzia Gaudich.

Photos
  1. Pouzolzia arachnoidea (Walp.) Wedd. in DC Prodr. 161 (1869) 228; --Merr., EPFP 2 (1923) 91. --Stachyocnide luzonica Blume, Mus. Bot. Lugd.-Bat. 2 (1856) 228, t. 55; --Type: Cuming 1492 (L, holo), Mindoro: Mindoro Oriental prov. LUZON: Ilocos Sur (San Quintin), Rizal (Montalban; Antipolo; Morong), Batangas (de la Paz), MINDORO. Boulders and cliffs and in thickets and thin forests at low and medium elevation. 
  2. Pouzolzia hirta (Blume) Hassk., Cat. Hort. Bogor. (1844) 80; --Gonostegia hirta (Blume) Miq., Ann. Mus. Bot. Lugd.-Bat. 4 (1869) 303; --Merr., EPFP 2 (1923) 92; --Yang, et al., Fl. Taiwan 2nd ed., 2 (1996) 223; --Urtica hirta Blume, Bijdr. (1825) 4(1825) 495. --Gonostegia integrifolia C.B.Rob., PJS 6 c (1911) Bot. 10, non Miq.
  3. Pouzolzia pentandra (Roxb.) Benn., Pl. Jav. Rar. (1832) 64, t. 14; --Gonostegia pentandra (Roxb.) Miq., Ann. Mus. Bot. Lugd.-Bat. 4 (1869) 302; --Merr., EPFP 2 (1923) 93; --Yang, et al., Fl. Taiwan 2nd ed., 2 (1996) 224; --Urtica pentandra Roxb., Fl. Ind. ed. 2, 3 (1832) 583. India, SE Asia, Taiwan. LUZON to MINDANAO. Cultivated lands along streams and in swampy places at low and medium elevation, ascending to 1400m.
  4. Pouzolzia rubricaulis (Blume) Wedd. in DC Prodr. 161 (1869) 229; --Merr., EPFP 2 (1923) 92. Philippines, Java. LUZON: Benguet (Ambuklao to Daklan, Merrill 4390). Damp shaded banks, c. 1400m.
  5. Pouzolzia zeylanica (L.) Benn., Pl. Jav. Rar. (1838) 67; --Merr., EPFP 2 (1923) 92; --Yang, et al., Fl. Taiwan 2nd ed., 2 (1996) 254. Taiwan, SE Asia. Throughout the Philippines at low and medium elevation, in open, usually damp grasslands. BATAN, BABUYAN ISLS (CAMIGUIN), LUZON: Mountain Province (Aluling), Ifugao (Kiangan), Aurora (Casiguran), Zambales (Subic), Pampanga (Bacolor), Rizal (Bosoboso), Manila, Laguna (Calauan; Los Baños), Camarines, POLILLO, MINDORO: Mindoro Oriental (Pola; Baco), MINDANAO: Zamboanga, Zamboanga del Sur (Port Banga), Lanao del Sur (Camp Keithley), Davao (Todaya). Photos

Procris Comm. ex Juss.

Photos
  1. Procris brunnea Merr., PJS 11 c (1916) Bot. 5; EPFP 2 (1923) 88; --Type: BS 23705 Ramos (US, iso), Luzon: Sorsogon prov., Mt Pocdal, epiphytic in mossy forest, 7-Aug-1915. LUZON: Sorsogon. Epiphytic in the mossy forest. Perhaps not sufficiently distinct from P. frutescens Blume.
  2. Procris dolichophylla Merr., PJS 8 c (1913) Bot. 369; EPFP 2 (1923) 88; --Type: Wenzel 98 (US, iso), Leyte: Dagami, climbing on trees in forested foothills, c. 60m, 29-Mar-1923. PANAY, NEGROS, LEYTE, MINDANAO: Agusan. Epiphytic in low and medium elevation damp forests. 
  3. Procris frutescens Blume, Bijdr. (1825) 510; --Merr., EPFP 2 (1923) 88. --Procris frutescens var. nervosa Elmer, LPB 8 (1915) 2863; --Type: Elmer 13602 (MO, US, iso), Mindanao: Agusan del Norte prov., Mt Urdaneta, Aug-1912. Sumatra, Peninsular Malaysia, Java, Philippines. LUZON: Laguna (San Antonio), Quezon (Lucban), NEGROS: Negros Occidental (Mt Kanlaon), MINDANAO: Lanao del Sur (Camp Keithley), Davao (Mt Apo), Agusan del Norte (Waloe; Mt Urdaneta). Damp forests at low and medium elevation. Photos
  4. Procris laevigata Blume, Bijdr. (1825) 508; --Merr., EPFP 2 (1923) 88; --Yang, et al., Fl. Taiwan 2nd ed., 2 (1996) 254, pl. 128. Africa and tropical Asia. Taiwan. N LUZON to MINDANAO. In forests, usually epiphytic, sometimes on ledges, sea-level to 1000m or more, widely scattered.
  5. Procris mindanaensis H Schröter, Fedde, Repert. Beih. 45 (1938) 189, in key, p. 281.
  6. Procris pedunculata (JR & G Forst.) Wedd., xxxx --H. Schröter, Fedde, Repert. Beih. 45 (1938) 259. Mascarenes, Malayan Region; Philippines, Polynesia.
  7. Procris urdanetensis Elmer, LPB 8 (1915) 2865; --Merr., EPFP 2 (1923) 89; --Syntypes: Elmer 13792 (NA, NY, US), Mindanao: Agusan del Norte prov., Mt Urdaneta, Sep-1912; --Elmer 13708 (MO, US), Mindanao: Agusan del Norte prov., Mt Urdaneta, Sept 1912. MINDANAO: Agusan del Norte (Mt Urdaneta). Mossy forest, 1600-1800m. Closely allied to P. frutescens Blume. Photos
  8. Procris wightiana (Wedd.) Wall., Cat. (1831) no 4638, nom. nud., Wedd. Monogr. (1856) 336; -- Merr., EPFP 2 (1923) 89; --H. Schröter, Fedde, Repert. Beih. 83 i (1935) 191. Procris crenata C.B.Rob., PJS 5 c (1910) Bot. 507; --Type: Vanoverbergh 635 (US, iso), Luzon: Mountain Province, Bauco, 1400m, Nov/Dec-1910. India to China and Taiwan. LUZON: Mountain Province. Epiphytic, c. 1400m. Reduced by some authors to P. laevigata Blume.

Urtica L.

Photos
  1. Urtica bullata Blume, Ann. Mus. Bot. Lugd.-Bat. 2 (1856) 145; --Merr., EPFP 2 (1923) 72; --Type from Java. Java, Philippines. LUZON: Mountain Province, Benguet (Mt Pulag), MINDANAO: Davao (Mt Apo). Open pine or cogon slopes, 1900-2400m on Mt Pulag.

 Villebrunea = Oreocnide


Literature 

Babu, CR & AN Henry. 1970. A new generic name in Urticaceae. Taxon 19: 651.

Chew, WL. 1963. Poikilospermum. Gardens’ Bulletin Singapore 20: 1-103.

Chew, WL. 1965. Laportea and allied genera (Urticaceae). Gardens’ Bulletin Singapore 21: 198-208.

Chew, WL. 1965. Nothocnide (Urticaceae) in Malesia. Gardens Bulletin Singapore 24: 361-373.

Chew Wee Lek, 1969. A monograph of Dendrocnide (Urticaceae). Gardens’ Bulletin Singapore 25: 1-104.

Chew, WL. 1969. A monograph of Laportea (Urticaceae). Gardens’ Bulletin Singapore 25: 111-178.

Johns, RJ. 1988. Urticaceae. The Flowering Plants of Papuasia, Dicotyledons, 2: 7-57.

Robinson, CB. 1911. Philippine Urticaceae. Philippine Journal of Science 6, no. 1, sect. C (Botany) 1-33, 3 fig.

Robinson, CB. 1911. Philippine Urticaceae. Philippine Journal of Science 6, no. 4, sect. C (Botany) 299-314.

Robinson, CB. 1911. Philippine Urticaceae. Philippine Journal of Science 5, no. 6, sect. C (Botany) 445-543.

Schröter, H. 1935-36. Feddes Repert. Beih. 83, 1-2 (1935-36).

Shih, B-L, Y-P Yang & H-Y Liu. 1995. A revision on Pilea, Lecanthus, and Procris (Urticaceae) of Taiwan. Taiwaniana 40: 256-290.

Unruh, M. 1943. Monographie der Gattung Leucosyke Zoll. et Mor. Botanische Jahrbücher 73: 191-258.

Wilmot-Dear, CM. 2009. Urticaceae for the non-specialist: identification in the Flora Malesiana region, Indochina and Thailand. Blumea 54: 233-241

Wilmot-Dear, CM & I Friis. 2013. The old world species of Boehmeria (Urticaceae, tribus Boehmerieae). A taxonomic revision. Blumea 58: 85-216

Wilmot-Dear, CM, I Friis & RHA Govaerts. 2014. Nomenclatural corrections to the taxonomic revision of the old world species of Boehmeria (Urticaceae, tribus Boehmerieae) by Wilmot-Dear & Friis (2013). Blumea 59: 95-97

Yang, Y-P, B-L Shih & H-Y Liu. 1996. Urticaceae. Flora of Taiwan ed. 2, 2: 197-257.


Acknowledgements 

Racquel Tan Chua-Barcelo and Danilo Tandang provided photographs for this Urticaceae page.


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Last updated 24 December 2016