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Heliotropiaceae

Argusia Boehmer = Heliotropium

Euploca Nutt.

Last edited by Pieter B. Pelser, 23 August 2019

Photos
  1. Euploca strigosa (Willd.) Diane & Hilger, Bot. Jahrb. Syst. 125 (2003) 49; --Heliotropium strigosum Willd., Sp. Pl. 1 (1797) 743; --Merr., EPFP 3 (1923) 378. --Heliotropium brevifolium Wall. Fl. Ind., ed. 1820 2 (1824) 2; --Riedl, Fl. Males. ser. 1, 13 (1997) 102. Africa, Bangladesh, Bhutan, China, India, Laos, Middle East, Myanmar, Nepal, Pakistan, Sri Lanka, Thailand, Vietnam, Philippines. LUZON: Zambales. Open sandy places near the sea. Photos

Heliotropium L.

Last edited by Pieter B. Pelser, 23 August 2019

Photos
  1. Heliotropium biblianum Craven, Blumea 50 (2005) 379; --Tournefortia tetrandra Blume, Bijdr. (1826) 845; I.M.Johnst., J. Arnold Arb. 16 (1935) 150; --Riedl, Fl. Males. ser. 1, 13 (1997) 123. Nicobars, Sri Lanka, Anambas & Natuna Isls, Sumatra, Peninsular Malaysia, Borneo, Java, New Guinea, Philippines. Montane forests and scrubs.
  2. Heliotropium bracteatum R.Br., Prodr. (1810) 493; --Merr., EPFP 3 (1923) 377. Java to tropical Australia. LUZON: Ilocos Norte, MINDANAO: Davao. Open sites at low elevation. Unaccounted for in the FM revision.
  3. Heliotropium curassavicum L., Sp. Pl. (1753) 130; --C.B.Rob., PJS 4 c (1909) 695; --Riedl, Fl. Males. ser. 1, 13 (1997) 103. Native of the Americas from Patagonia to the US and the West Indies, Java (near Bogor), Philippines. Occasionally occurring as weed. Photos
  4. Heliotropium foertherianum Diane & Hilger, Bot. Jahrb. Syst. 125 (2003) 46. --Tournefortia argentea L.f., Suppl. (1781) 133; --Merr., EPFP 3 (1923) 376; --Riedl, Fl. Males. ser. 1, 13 (1997) 117; --Argusia argentea (L.f.) Heine, Fl. New Caled. & Depend. 7 (1976) 108; --Dayang Awa in Tree Fl. Sabah & Sarawak 2 (1996) 95; --Messerschmiedia argentea (L.f.) I.M.Johnst., J. Arnold Arb. 16 (1935) 164; J. Arnold Arb. 32 (1951) 121; --Type: from Sri Lanka. E African coasts, across the Indian Ocean, Ryukyus, Taiwan, Hainan, Vietnam, Sumatra, Borneo, Java, Philippines, Sulawesi, Lesser Sunda Isls (Bali, Lombok, Timor), Moluccas, New Guinea, N Australia to Polynesia, New Caledonia. BATANES and N LUZON to PALAWAN, MINDANAO, and the SULU ARCHIPELAGO. Along sandy seashores. Photos
  5. Heliotropium indicum L., Sp. Pl. (1753) 130; --Merr., Fl. Manila (1912) 395; EPFP 3 (1923) 377; --I.M.Johnst., J. Arnold Arb. 32 (1951) 111; --Riedl, Fl. Males. ser. 1, 13 (1997) 105. Probably a native of tropical America, now pantropical. In Malesia: Sumatra, Peninsular Malaysia, Singapore, Borneo, Java, Philippines, Sulawesi, Lesser Sunda Isls (Lombok, Sumbawa, Sumba, Flores, Timor), Moluccas and New Guinea. A common weed in the settled areas throughout the Philippines at low elevation. Photos
  6. Heliotropium luzonicum (I.M. Johnst.) Craven, Blumea 50 (2005) 379; --Tournefortia luzonica I.M.Johnst., J. Arnold Arb. 16 (1935) 156; --Riedl, Fl. Males. ser. 1, 13 (1997) 118; --Type: Adduru 237 (A, K), Luzon.
  7. Heliotropium muelleri (I.M. Johnst.) Craven, Blumea 50 (2005) 380; --Tournefortia muelleri I.M.Johnst., J. Arnold Arb. 16 (1935) 157; --Riedl, Fl. Males. ser. 1, 13  (1997) 120; --Type: from Australia. N Australia, ?Java, Lesser Sunda Isls (?Bali, Sumba), New Guinea & Philippines. Along rivers and in tidal areas. 
  8. Heliotropium sarmentosum (Lam.) Craven, Blumea 50 (2005) 380; --Tournefortia sarmentosa Lam., Tabl. Encycl. 1 (1797) 416; --Merr., Fl. Manila (1912) 394; EPFP 3 (1923) 377; --I.M.Johnst., J. Arnold Arb. 16 (1935) 147; J. Arnold Arb. 32 (1951) 115; --Riedl, Fl. Males. ser. 1, 13 (1997) 121. --Tournefortia horsfieldii Miq., Fl. Ind. Bat. 2 (1858) 927; --Merr., EPFP 3 (1923) 376. Australia, Taiwan, Sumatra, Malay Peninsula, Borneo, Java, Philippines, Celebes, Lesser Sunda Islands (Lombok, Flores, Timor, Alor, Wetar), Moluccas, New Guinea. Photos

Messerschmidia Hebenstreit = Heliotropium

Tournefortia L. = Heliotropium


Acknowledgements

Ernest Kurt Tan kindly provided a number of photographs found on this webpage.

Literature

Craven, LA. 2005. Malesian and Australian Tournefortia transferred to Heliotropium and notes on delimitation of Boraginaceae. Blumea 50: 375-381.

Hilger HH & N Diane. 2003. A systematic analysis of Heliotropiaceae (Boraginales) based on trnL and ITS1 sequence data. Botanische Jahrbücher 125: 19-51

Johnston, IM. 1951. Studies in the Boraginaceae. Journal of Arnold Arboretum 32: 1-26; 99-122.

Luebert, F, G Brokamp, J Wen, M Weigend & HH Hilger. 2011. Phylogenetic relationships and morphological diversity in Neotropical Heliotropium (Heliotropiaceae). Taxon 60: 663-680.

Riedl, H. 1997. Boraginaceae. Flora Malesiana series 1, 13: 43-144

Pelser, P.B., J.F. Barcelona & D.L. Nickrent (eds.). 2011 onwards. Co's Digital Flora of the Philippines. www.philippineplants.org

Copyright © 2011, Co's Digital Flora of the Philippines

Last updated 17 May 2020