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Primulaceae

Including Aegicerataceae, Maesaceae, Myrsinaceae

Aegiceras Gaertn.

Last edited by Pieter B. Pelser, 31 May 2018

Photos
  1. Aegiceras corniculatum (L.) Blanco, Fl. Filip. (1837) 79; --Merr., Fl. Manila (1912) 361; EPFP 3 (1923) 255; --Rhizophora corniculata L., Herb. Amb. (1754) 13.  Andaman Isls, Bangladesh, Cambodia, China, India, Java, Lesser Sunda Isls, Peninsular Malaysia, Moluccas, Myanmar, New Guinea, Nicobar Islsl, Norfolk Isl, Australia, Solomon Isl, Sri Lanka, Sulawesi, Sumatra, Vietnam, Philippines. Photos
  2. Aegiceras floridum Roem. & Schult., Syst. 4 (1819) 512; --Merr., EPFP 3 (1923) 256. Lesser Sunda Isls, Moluccas, New Guinea, Sulawesi, Philippines. Photos

Amblyanthopsis Mez

Last edited by Pieter B. Pelser, 19 July 2020

No photos
  1. Amblyanthopsis philippinensis Mez, PJS 1 (1906) Suppl. 274; --Merr., EPFP 3 (1923) 266; --Type: Merrill 3881, Luzon: Bataan Prov, Mt Mariveles. Endemic to the Philippines. LUZON: Bataan, Quezon, Rizal. Primary forests along small streams, c. 800m. Note by P.B. Pelser (19-Jul-20): According to Merrill (1923) perhaps better placed in Discocalyx.

Androsace L.

Last edited by Pieter B. Pelser, 4 August 2019

Photos
  1. Androsace umbellata (Lour.) Merr., PJS 15 (1919) 237; EPFP 3 (1923) 274; --Primula umbellata (Lour.) Bentvelzen, Fl. Males. ser. 1, 6 (1962) 191; --Drosera umbellata Lour., Fl. Cochinch. (1790) 186. Bangladesh, China, India, Japan, Myanmar, Nepal, New Guinea, North Korea, Pakistan, Russia, Ryukyu Isls, South Korea, Taiwan, Tibet, Vietnam, Philippines. LUZON: Mountain Province. Montane forests in shaded moist places, 1300-1500m.

Ardisia Sw.

Last edited by Pieter B. Pelser, 21 October 2020

Photos
  1. Ardisia alverezii Merr., PJS 26 (1925) 484; EPFP 4 (1925) 250. Endemic to the Philippines. BABUYAN ISLS (CALAYAN), LUZON: Apayao, Cagayan. Forests at low elevations.
  2. Ardisia basilanensis Merr., PJS 12 c (1917) Bot. 153; --Merr., EPFP 3 (1923) 256. Endemic to the Philippines. BASILAN, MINDANAO: Zamboanga. Primary forests at low and medium elevation.
  3. Ardisia brevipetiolata (Merr.) Merr., EPFP 3 (1923) 256; --Ardisia serrata var. brevipetiolata Merr., PJS 2 c (1907) Bot. 296. --Type: Merrill 4049, Mindoro: Mindoro Oriental Prov, Baco River near base of Mt Halcon. --Ardisia sibuyanensis Elmer, LPB 4 (1912) 1497; --Merr., EPFP 3 (1923) 263; --Molina & Buot, J. New Biol. Rep. 9 (2020) 295. Endemic to the Philippines. MINDORO: Mindoro Oriental, SIBUYAN. Primary forests, also reported along streams, 225-550m. Ardisia sibuyanensis is a form with entire leaves.
  4. Ardisia calavitensis Merr., PJS 20 (1922) 423; EPFP 3 (1923) 256. Endemic to the Philippines. MINDORO. Primary forests at low and medium elevation.
  5. Ardisia castaneifolia Mez in Engl., Pflanzenreich 9 (1902) 138; --Merr., EPFP 3 (1923) 257. Endemic to the Philippines. LUZON: Bulacan. Probably in forests at low and medium elevation.
  6. Ardisia confertiflora Merr., PJS 5 c (1910) Bot. 213; EPFP 3 (1923) 257; --Molina & Buot, J. New Biol. Rep. 9 (2020) 290. Endemic to the Philippines. BABUYAN, BATAN, FUGA, MINDORO. Low elevation thickets. Photos
  7. Ardisia copelandii Mez, PJS 1 (1906) Suppl. 272; --Merr., EPFP 3 (1923) 257; --Molina & Buot, J. New Biol. Rep. 9 (2020) 290. Borneo, Sulawesi, Philippines. MINDANAO: Bukidnon, Davao, Lanao. Primary forests, 600-1000m.
  8. Ardisia crispa (Thunb.) A.DC., Trans. Linn. Soc. 17 (1834) 124; --Merr., EPFP 3 (1923) 257; --Molina & Buot, J. New Biol. Rep. 9 (2020) 291; --Bladhia crispa Thunb., Syst. Veg. ed. 14 (1784) 237. Borneo, China, India, Japan, Java, Korea, Laos, Ryukyus, Sumatra, Taiwan, Vietnam, Philippines. LUZON: Benguet, Ilocos Norte, Mountain Province, Nueva Vizcaya, Pangasinan, Zambales, NEGROS, PANAY. Chiefly in the mossy forest, 800-2200m. Photos
  9. Ardisia cumingiana A.DC., Prod. 8 (1844) 136; --Merr., EPFP 3 (1923) 257. --Ardisia centenoi S.Vidal, Sinopsis Atlas (1883) 30; --Molina & Buot, J. New Biol. Rep. 9 (2020) 290. Endemic to the Philippines. LUZON: Batangas, Benguet, Cagayan, Ilocos Sur. Ravines, up to 1500m.
  10. Ardisia curranii Merr., PJS 5 c (1910) Bot. 214; EPFP 3 (1923) 258; --Molina & Buot, J. New Biol. Rep. 9 (2020) 291. Endemic to the Philippines. LUZON: Camarines Sur. Primary forests at low elevation.
  11. Ardisia curtiflora Elmer, LPB 8 (1915) 2778; --Merr., EPFP 3 (1923) 258. Endemic to the Philippines. MINDANAO: Agusan del Norte (Mt Urdaneta). Mossy forests, c. 1600m.
  12. Ardisia cymosa Blume, Bijdr. (1826) 689; --C.M.Hu, Blumea 44 (1999) 393. --Ardisia cagayanensis Merr., PJS 7 c (1912) Bot. 323; EPFP 3 (1923) 256. Borneo, China, Java, Peninsular Malaysia, Sumatra, Taiwan, Thailand, Vietnam, Philippines. CATANDUANES, LUZON: Cagayan. Primary forests at low elevation (as Ardisia cagayanensis Merr.).
  13. Ardisia darlingii Merr., PJS 5 c (1910) Bot. 215; EPFP 3 (1923) 258; --C.M.Hu, Blumea 44 (1999) 398; --Molina & Buot, J. New Biol. Rep. 9 (2020) 292. --Ardisia gitingensis Elmer, LPB 4 (1912) 1498; --Merr., EPFP 3 (1923) 258. Endemic to the Philippines. Photos
  14. Ardisia diffusa Merr., PJS 5 c (1910) Bot. 216; EPFP 3 (1923) 258; --Molina & Buot, J. New Biol. Rep. 9 (2020) 292. --Ardisia diffusa Mez, Fedde Repert. 16 (1920) 410; --Molina & Buot, J. New Biol. Rep. 9 (2020) 293. Endemic to the Philippines. MINDANAO: Misamis Occidental, Misamis Oriental, Zamboanga del Norte. Forests, probably at medium elevation.
  15. Ardisia elliptica Thunb., Nov. Gen. Pl. 8 (1798) 119; --Molina & Buot, J. New Biol. Rep. 9 (2020) 292. --Ardisia sorsogonensis Elmer ex Merr., EPFP 3 (1923) 264, nom. inval. --Ardisia squamulosa C.Presl, Rel. Haenk. 2 (1830) 65; --Merr., EPFP 3 (1923) 263. --Ardisia pirifolia Mez in Engl., Pflanzenreich 9 (1902) 129; --Merr., EPFP 3 (1923) 261; --Molina & Buot, J. New Biol. Rep. 9 (2020) 293. --Ardisia verrucosa C.Presl, Reliq. Haenk. 2 (1835) 66; --Merr., EPFP 3 (1923) 265; . --Ardisia negroensis Mez, Fedde Repert. 16 (1920) 411; --Molina & Buot, J. New Biol. Rep. 9 (2020) 293. Bangladesh, Borneo, Cambodia, India, Java, Laos, Peninsular Malaysia, Maldives, Nansei-shoto, New Guinea, Nicobar Isls, Sri Lanka, Sulawesi, Sumatra, Taiwan, Thailand, Vietnam, Philippines. LUZON: Aurora, Batangas, Cagayan, Camarines Sur, Cavite, Ilocos Norte, Isabela, Laguna, La Union, Quezon, Pangasinan, Sorsogon, Zambales, MINDANAO: Davao, Surigao del Norte, MINDORO, NEGROS, PALAWAN, PANAY. In most or all islands and provinces. Common in primary forests at low and medium elevation, ascending to 1000m. According to Merrill (1923), a tree rather than a shrub, not certainly distinct from Ardisia obovata Blume. Note by L.L.Co: Ardisia verrucosa is hardly distinguishable from Ardisia squamulosa. Conservation status: Vulnerable (DAO 2017-11; as Ardisia squamulosa).
  16. Ardisia elmeri Mez, PJS 1 (1906) Suppl. 273; --Merr., EPFP 3 (1923) 258; --Molina & Buot, J. New Biol. Rep. 9 (2020) 292; --Type: Elmer 6493 (MO, NY, US, iso), Luzon: Benguet Prov, Baguio, Jun-1904. --Ardisia fragrans Elmer, LPB 2 (1910) 664; --Merr., EPFP 3 (1923) 258; --Molina & Buot, J. New Biol. Rep. 9 (2020) 292; --Type: Elmer 11647 (US, lecto; BISH, GH, L, MO, NA, isolecto), Mindanao: Davao, Mt Apo, Todaya, Sep-1909. Endemic to the Philippines. LUZON: Albay, Benguet, Camarines Sur, Mountain Province, Rizal, Sorsogon, MINDANAO: Davao (Mt Apo). Humid lower and midmontane forests, damp ravines, 1200-1600m.
  17. Ardisia geissanthoides Mez, Fedde Repert. 16 (1920) 412; --Merr., EPFP 3 (1923) 258; --Molina & Buot, J. New Biol. Rep. 9 (2020) 292. --Ardisia glauca Mez, Fedde Repert. 16 (1920) 412; --Molina & Buot, J. New Biol. Rep. 9 (2020) 292. Endemic to the Philippines. LUZON: Laguna, MINDORO. Mossy forest, 1800-2000m.
  18. Ardisia ilocana Merr., PJS 14 (1919) 443; EPFP 3 (1923) 259. Endemic to the Philippines. LUZON: Ilocos Norte. Forests, c. 500m.
  19. Ardisia iwahigensis Elmer, LPB 5 (1913) 1817; --Merr., EPFP 3 (1923) 259. Endemic to the Philippines. PALAWAN: Puerto Princesa (Mt Pulgar = Thumb Peak). Damp primary forests, c. 225m. Conservation status: Vulnerable (DAO 2017-11).
  20. Ardisia keithleyi Merr., PJS 10 c (1915) Bot. 335; EPFP 3 (1923) 259; --Molina & Buot, J. New Biol. Rep. 9 (2020) 292. --Ardisia oblongifolia Merr., PJS 5 c (1910) Bot. 219, nom. illeg., auct. non Gamble. Endemic to the Philippines. MINDANAO: Agusan del Norte, Lanao del Sur, Surigao del Sur, Zamboanga, NEGROS. Primary forests, c. 650m.
  21. Ardisia laxiflora Merr., PJS 12 c (1917) Bot. 153; EPFP 3 (1923) 259; --Molina & Buot, J. New Biol. Rep. 9 (2020) 292. Endemic to the Philippines. LUZON: Quezon, MINDANAO: Lanao del Norte. Primary forests at low elevation.
  22. Ardisia leytensis Merr., PJS 8 c (1913) 386; --Type: Wenzel s.n., Leyte, Dagami, 5-Jun-1913. Endemic to the Philippines. LEYTE.
  23. Ardisia loheri Merr., PJS 12 c (1917) Bot. 152; EPFP 3 (1923) 259; --Molina & Buot, J. New Biol. Rep. 9 (2020) 293. Endemic to the Philippines. CEBU, LUZON: Laguna, Rizal, MINDANAO: Bukidnon, NEGROS. Primary forests at medium elevation, ascending to 1200m.
  24. Ardisia longipetiolata Merr., PJS 14 (1919) 444; EPFP 3 (1923) 259. Endemic to the Philippines. LUZON: Camarines Norte. Damp primary forests at low elevation.
  25. Ardisia macgregorii Merr., PJS 5 c (1910) Bot. 217; EPFP 3 (1923) 260; --Type: BS 1722 McGregor (US, lecto), Cebu: near Toledo. Endemic to the Philippines. CEBU, LUZON: Ilocos Norte, SAMAR. In thickets and forests at low and medium elevation.
  26. Ardisia marginata Blume, Bijdr. (1826) 688; --Molina & Buot, J. New Biol. Rep. 9 (2020) 293. --Ardisia apoensis Elmer, LPB 2 (1910) 663; --Merr., EPFP 3 (1923) 256. --Ardisia cincta Mez, Fedde Repert. 16 (1920) 312; --Molina & Buot, J. New Biol. Rep. 9 (2020) 290. Borneo, Java, Philippines. BILIRAN, CAMIGUIN DE MINDANAO, LEYTE, LUZON: Abra, Benguet, Cagayan, Camarines Norte, Laguna, Mountain Province, Nueva Ecija, Quezon, Rizal, MINDANAO: Davao, Lanao del Sur. Primary forests at low and medium elevation, also on forested ridges to 1200m.
  27. Ardisia mezii Elmer, LPB 2 (1908) 440; --Merr., EPFP 3 (1923) 260. Endemic to the Philippines. LUZON: Laguna, Quezon, Sorsogon, MINDANAO: Bukidnon. Primary forests, 700-1200m.
  28. Ardisia mindanaensis Mez in Engl., Pflanzenreich 9 (1902) 102; --Merr., EPFP 3 (1923) 260; --Molina & Buot, J. New Biol. Rep. 9 (2020) 293. Endemic to the Philippines. LEYTE, MINDANAO: Agusan del Norte, Bukidnon, Davao, Lanao. Primary forests, 500-1000m.
  29. Ardisia mirandae Merr., PJS 12 c (1917) Bot. 154; EPFP 3 (1923) 260; --Molina & Buot, J. New Biol. Rep. 9 (2020) 293. Endemic to the Philippines. LUZON: Camarines Sur. Exposed peaks and ridges in the mossy forest, c. 700m.
  30. Ardisia nigromaculata Merr., PJS 13 c (1918) Bot. 49; EPFP 3 (1923) 260; --Molina & Buot, J. New Biol. Rep. 9 (2020) 293. Endemic to the Philippines. LUZON: Camarines Sur, Quezon, NEGROS, PANAY. Mossy forests, c. 1000m.
  31. Ardisia oreophila C.M.Hu, Blumea 47 (2002) 496; --Type: Soejarto, Fernando & Sagcal 8828 (L, holo), Palawan: Brookes Point, Mt Mantalingahan. Ridges and slopes of Sindurapi Peak, 1100m. Endemic to the Philippines. PALAWAN. Known only from the type collection.
  32. Ardisia palawanensis Merr., PJS 5 c (1910) Bot. 220; EPFP 3 (1923) 261. Endemic to the Philippines. PALAWAN. Undershrub in lowland flat forest.
  33. Ardisia polysticta Miq., Fl. Ind. Bat. Suppl. 1 (1861) 576; --C.M.Hu, Blumea 44 (1999) 395; --Molina & Buot, J. New Biol. Rep. 9 (2020) 294. --Ardisia jagorii Mez in Engl., Pflanzenreich 9 (1902) 148; --Merr., EPFP 3 (1923) 259. --Ardisia pardalina Mez in Engl., Pflanzenreich 9 (1902) 148; --Merr., EPFP 3 (1923) 261. --Ardisia ramosii Merr., PJS 7 (1912) 148; --C.M.Hu, Blumea 44 (1999) 395. --Ardisia sinuato-crenata Mez in Engl., Pflanzenreich 9 (1902) 148; --Merr., EPFP 3 (1923) 263. Assam, Bangladesh, Borneo, Cambodia, China, Hainan, Laos, Peninsular Malaysia, Myanmar, Sumatra, Taiwan, Thailand, Vietnam, Philippines. BILIRAN, LEYTE, LUZON: Albay, Bataan, Benguet, Cagayan, Camarines, Ilocos Norte, Laguna, Mountain Province, Nueva Vizcaya, Rizal, Sorsogon, MINDANAO, MINDORO: Bukidnon, PANAY, SAMAR. Primary forests from low to high elevation, in mossy forest ascending to 2300m (as Ardisia pardalina and Ardisia ramosii).
  34. Ardisia pulchella Mez, Fedde Repert. 16 (1920) 410; --Merr., EPFP 3 (1923) 261. Endemic to the Philippines. LUZON: Benguet, Mountain Province. Mossy forests, 2000-2400m.
  35. Ardisia purpurea Reinw. ex Blume, Bijdr. Fl. Ned. Ind. (1826) 684; --Molina & Buot, J. New Biol. Rep. 9 (2020) 294. --Ardisia lanceolata Roxb., Fl. Ind. ed. Carey 2 (1824) 275, nom. illeg., auct. non C.F.Gaertn.; --Hort. Beng. (1814) 85, nom. nud.; --Merr., EPFP 3 (1923) 259; --C.M.Hu, Blumea 44 (1999) 401. --Ardisia clementis Elmer, LPB 2 (1910) 665; --Merr., EPFP 3 (1923) 257. --Ardisia lanaensis Mez, Fedde Repert. 16 (1920) 411. Borneo, Java, Lesser Sunda Isls, Peninsular Malaysia, Sulawesi, Sumatra, Thailand, Philippines. BASILAN, LUZON: Sorsogon, MINDANAO: Davao (Mt Apo), Lanao del Sur, Zamboanga. Primary forests, from the lowlands (sometimes near the seashore) to 1100m.
  36. Ardisia pusilla A.DC., Trans. Linn. Soc. 17 (1836) 126. --Ardisia reptans Merr., PJS 5 c (1910) Bot. 220; EPFP 3 (1923) 262; --Molina & Buot, J. New Biol. Rep. 9 (2020) 294. China, Japan, Korea, Ryukyus, Taiwan, Thailand, Philippines. LUZON: Pampanga. Forested ravines, c. 1350m.
  37. Ardisia pyramidalis (Cav.) Pers., Syn. 1 (1805) 233; --Merr., EPFP 3 (1923) 261; --Anguillaria pyramidalis Cav., Icon. 6 (1800) 1. --Ardisia perrottetiana A.DC., Ann. Sci. Nat., Bot. sér. 2, 16 (1841) 95. Borneo, Philippines. LUZON: Cagayan to Albay, PALAWAN, MINDANAO, in most islands and provinces. Primary forests at low and medium elevation, often common. Photos
  38. Ardisia racemosopaniculata Mez, PJS 1 (1906) Suppl. 273; --Merr., EPFP 3 (1923) 262; --Molina & Buot, J. New Biol. Rep. 9 (2020) 294. Endemic to the Philippines. MINDANAO: Davao and North Cotabato (Mt Apo), Surigao del Norte. Primary forests, c. 1600m.
  39. Ardisia reynosoi Stone, Sida 16 (1994) 263; --Type: Argent & Reynoso 89-163 (PNH, holo; AAU, E, iso), Sibuyan: Mt Giting-giting, above Magdiwang on ridge leading to Mayo’s Peak, grassy area above Camp 3, 1400m. Endemic to the Philippines. SIBUYAN.
  40. Ardisia rivularis Merr., PJS 13 c (1918) Bot. 50; EPFP 3 (1923) 262. Endemic to the Philippines. LUZON: Quezon, in forests at low and medium elevation. Photos
  41. Ardisia romanii Elmer, LPB 5 (1913) 1820; --Merr., EPFP 3 (1923) 262. --Ardisia magnifica Mez, Fedde Repert. 16 (1920) 413; --Molina & Buot, J. New Biol. Rep. 9 (2020) 293. Endemic to the Philippines. PALAWAN. Forests at low elevation. Conservation status: Vulnerable (DAO 2017-11). Photos
  42. Ardisia saligna Mez in Engl., Pflanzenreich 9 (1902) 143; --Merr., EPFP 3 (1923) 262; --Molina & Buot, J. New Biol. Rep. 9 (2020) 294. --Ardisia macropus Mez, Fedde Repert. 16 (1920) 415; EPFP 3 (1923) 260; --Molina & Buot, J. New Biol. Rep. 9 (2020) 293. Endemic to the Philippines. LUZON: Abra, Bataan, Batangas, Benguet, Bulacan, Cavite, Ilocos Norte, Laguna, Mountain Province, Nueva Ecija, Quezon, Rizal, NEGROS, POLILLO. Primary forests at low and medium elevation, ascending to 1000m.
  43. Ardisia samarensis Merr., PJS 12 c (1917) Bot. 151; EPFP 3 (1923) 263. Endemic to the Philippines. SAMAR. Damp primary forests at low elevation.
  44. Ardisia scabrida Mez in Engl., Pflanzenreich 9 (1902) 118; --Merr., EPFP 3 (1923) 263; --Molina & Buot, J. New Biol. Rep. 9 (2020) 294. --Ardisia punctata auct. non Lindl.; --Elmer, LPB 2 (1908) 439; --Type: Elmer 10385 (A, iso?), Negros: Negros Oriental Prov, Cuernos Mtns, Jun-1908. Endemic to the Philippines. CAMIGUIN, MINDANAO: Bukidnon, Davao (Mt Apo), Misamis, NEGROS, SULU. Forests, 600-1300m.
  45. Ardisia scalaris Mez in Engl., Pflanzenreich 9 (1902) 142; --Merr., EPFP 3 (1923) 263. Endemic to the Philippines. LUZON: Quezon, Rizal. Primary forests along streams at low elevation. According to Merrill (1923), very similar to Ardisia saligna Mez.
  46. Ardisia serrata (Cav.) Pers., Syn. 1 (1805) 233; --Merr., EPFP 3 (1923) 263; --C.M.Hu, Blumea 44 (1999) 398; --Molina & Buot, J. New Biol. Rep. 9 (2020) 294; --Anguillaria serrata Cav., Icon. 6 (1800) 2. --Ardisia luzonensis C.Presl, Reliq. Haenk. 2 (1835) 65; --Molina & Buot, J. New Biol. Rep. 9 (2020) 293. --Ardisia angustifolia A.DC., Prodr. 8 (1844) 134; --Merr., EPFP 3 (1923) 256. --Ardisia abanii Stone, Proc. Acad. Nat. Sci. Philadelphia 142 (1990) 53. --Ardisia mindorensis Merr., PJS 5 c (1910) Bot. 218; EPFP 3 (1923) 260. --Ardisia oligocarpa Merr., J. Arn. Arb. 35 (1954) 148. --Ardisia oligantha auct. non Baker; --Elmer, LPB 4 (1912) 146; --Merr., EPFP 3 (1923) 260. --Ardisia whitfordii Mez, PJS 1 (1906) Suppl. 271; --Merr., EPFP 3 (1923) 265. --Ardisia grandidens Mez in Engl., Pflanzenreich 9 (1902) 137; --Merr., EPFP 3 (1923) 259; --Molina & Buot, J. New Biol. Rep. 9 (2020) 292. --Ardisia curtipes Merr., PJS 5 c (1910) Bot. 372; EPFP 3 (1923) 258. --Ardisia dataensis Mez, Fedde Repert. 16 (1920) 414. Borneo, Java, Philippines. BILIRAN, LEYTE, LUZON: Apayao, Benguet, Cagayan, Ifugao, Laguna, Mountain Province, Nueva Ecija, Quezon, Rizal, Sorsogon, Zambales, MINDANAO, MINDORO, PALAWAN, PANAY, POLILLO, SAMAR, SIBUYAN. Primary forests from lowland to mossy, on ridges, to 2400m. Data compiled from distribution range of above-cited synonyms, excepting Ardisia abanii Stone and Ardisia oligocarpa Merr. Photos
  47. Ardisia stichantha B.C.Stone, Pacific Sci. 47 (1993) 290; --Type: BS 29315 Ramos & Edaño (US, holo), Luzon: Rizal Prov, Mt Susong-Dalaga. Aug-1917. Endemic to the Philippines. LUZON: Rizal.
  48. Ardisia sulcata Mez in Engl., Pflanzenreich 9 (1902) 112; --Merr., EPFP 3 (1923) 264; --Molina & Buot, J. New Biol. Rep. 9 (2020) 295; --Type: Warburg 12286. Endemic to the Philippines. BILIRAN, LEYTE, LUZON: Apayao, Cagayan, Camarines Sur, Tayabas, SAMAR. Damp primary forests at low elevation.
  49. Ardisia tayabensis Merr., PJS 12 c (1917) Bot. 156; EPFP 3 (1923) 264; --Molina & Buot, J. New Biol. Rep. 9 (2020) 295. Endemic to the Philippines. LUZON: Aurora, Quezon. Primary forests at or above 300m.
  50. Ardisia taytayensis Merr., J. Arnold Arbor. 35 (1954) 148. --Ardisia pachyphylla auct. non Dunn; --Merr., PJS 12 c (1917) Bot. 154; EPFP 3 (1923) 260. Endemic to the Philippines. BALABAC, PALAWAN. Dry thickets and borders of forests at low elevation. Conservation status: Vulnerable (DAO 2017-11).
  51. Ardisia tomentosa C.Presl, Reliq. Haenk. 2 (1835) 66; --Merr., EPFP 3 (1923) 265; --C.M.Hu, Blumea 44 (1999) 397; --Molina & Buot, J. New Biol. Rep. 9 (2020) 295. --Ardisia disticha A.DC., Prodr. 8 (1844) 129; --Pimelandra disticha (A.DC.) Fern.-Vill., Nov. App. (1880) 123; --Molina & Buot, J. New Biol. Rep. 9 (2020) 292. --Ardisia philippinensis A.DC., Prodr. 8 (1844) 128; --Merr., EPFP 3 (1923) 261; --Type: Cuming 1467. --Ardisia cuprea Elmer, LPB 8 (1915) 2777; --Merr., EPFP 3 (1923) 257; --Type: Elmer 13287 (C, L, NY 2 sheets, iso), Mindanao: Agusan del Norte Prov, Mt Urdaneta. --Ardisia peninsula Elmer, LPB 10 (1939) 3764, nom. inval., no Latin descr.; --Type: Elmer 16615 (C, NA, NY, iso), Luzon: Sorsogon Prov, Irosin, Mt Bulusan. Borneo, Philippines. LUZON: Albay, Batangas, Benguet, Bulacan, Camarines, Ilocos Norte, Isabela, Laguna, Nueva Ecija, Rizal, Sorsogon, Zambales, PANAY, MINDANAO: Agusan  del Norte, Surigao. Primary forests at low and medium elevation, ascending to 450m. Photos
  52. Ardisia villosa Roxb., Fl. Ind. ed. Carey 2 (1824) 272. Borneo, Cambodia, China, Hong Kong, Java, Laos, Myanmar, Peninsular Malaysia, Sumatra, Taiwan, Thailand, Vietnam, Philippines.
  53. Ardisia warburgiana Mez in Engl., Pflanzenreich 9 (1902) 143; --Merr., EPFP 3 (1923) 265; --C.M.Hu, Blumea 44 (1999) 396; --Molina & Buot, J. New Biol. Rep. 9 (2020) 295. --Ardisia proteifolia Mez in Engl., Pflanzenreich 9 (1902) 143; --Merr., EPFP 3 (1923) 261. --Ardisia sibulanensis Elmer, LPB 2 (1910) 662; --Molina & Buot, J. New Biol. Rep. 9 (2020) 295. --Ardisia membranifolia Mez, Fedde Repert. 16 (1920) 415; --Molina & Buot, J. New Biol. Rep. 9 (2020) 293. Endemic to the Philippines. LUZON: Apayao, Benguet, Cagayan, Isabela, Mountain Province, Pampanga, Sorsogon, Zambales, MINDANAO: Agusan, Bukidnon, Davao, Lanao, MINDORO, PANAY, SAMAR. Forests from low elevation to 1700m.
  54. Ardisia yatesii Merr., PJS 12 c (1917) Bot. 155; EPFP 3 (1923) 266. Endemic to the Philippines. LUZON: Quezon. Primary forests, 350-400m. Closely allied to Ardisia tomentosa C.Presl.
  55. Ardisia zambalensis Merr., PJS 10 c (1915) Bot. 334; EPFP 3 (1923) 266. Endemic to the Philippines. LUZON: Zambales. Exposed forested ridges, 1000-1400m.

 Discocalyx (DC.) Mez

Last edited by Pieter B. Pelser, 19 July 2020

Photos
  1. Discocalyx angustifolia Mez in Engl., Pflanzenreich 9 (1902) 214; --Merr., EPFP 3 (1923) 266; --Type: Gaudichaud 299 (P). Endemic to the Philippines. LUZON: Laguna, Quezon, ALABAT, PANAY. Low elevation damp primary forests.
  2. Discocalyx angustissima Merr., PJS 12 c (1917) Bot. 143; EPFP 3 (1923) 267. --Discocalyx longissima Merr., PJS 17 (1920) 302; EPFP 3 (1923) 268; --Type: BS 34456 Ramos & Pascasio. Endemic to the Philippines. LUZON: Cagayan, Nueva Ecija, Quezon, MINDANAO: Surigao. Low and medium elevation primary forests, rocky slopes. Discocalyx longissima is said to differ from Discocalyx angustissima by its entire leaf margin. The type of the former in US has minute teeth.
  3. Discocalyx brachybotrya Merr., PJS 20 (1922) 424; EPFP 3 (1923) 267; --Type: BS 37463 Ramos & Edaño. Endemic to the Philippines. MINDANAO: Zamboanga. Low elevation forests along streams.
  4. Discocalyx cybianthoides (A.DC.) Mez in Engl., Pflanzenreich 9 (1902) 213; --Merr., EPFP 3 (1923) 267; --Badula cybianthoides A.DC., Prodr. 8 (1844) 107. Borneo, Philippines. LUZON: Ilocos Norte, Abra, Cagayan, Nueva Vizcaya, Pangasinan, Nueva Ecija, Bataan, Bulacan, Rizal, Laguna, Batangas, Quezon, Sorsogon, MINDORO, MASBATE, LEYTE. Low and medium elevation primary forests, often common. Photos
  5. Discocalyx effusa Mez in Engl., Pflanzenreich 9 (1902) 213; --Merr., EPFP 3 (1923) 267. Endemic to the Philippines. LUZON: Nueva Ecija, Rizal, Laguna, Quezon, CATANDUANES, MINDORO, PANAY, NEGROS, MINDANAO: Davao. Low and medium elevation damp primary forests. Photos
  6. Discocalyx euphlebia Merr., PJS 12 c (1917) Bot. 144; EPFP 3 (1923) 267; --Type: BS 17646 Ramos. Endemic to the Philippines. SAMAR. Low elevation damp primary forests.
  7. Discocalyx filipes Mez, Fedde Repert. 16 (1920) 422; --Merr., EPFP 3 (1923) 267; --Type: BS 1857 Ramos. Endemic to the Philippines. LUZON: Rizal. Medium elevation primary forests.
  8. Discocalyx hymenandroides Mez, Fedde Repert. 16 (1920) 422; --Merr., EPFP 3 (1923) 267; --Type: FB 5096 Curran. Endemic to the Philippines. LUZON: Benguet, Cagayan, Rizal. Medium elevation damp forests, ascending to 1400m in Benguet.
  9. Discocalyx insignis Merr., PJS 5 c (1910) Bot. 221; EPFP 3 (1923) 268; --Type: FB 7616 Hutchinson, Mindanao: Agusan del Norte Prov, Agusan River valley near Amparo, in forests, c. 130m, 26-Aug-1907. --Discocalyx congestiflora Elmer, LPB 8 (1915) 2780; --Type: Elmer 13724 (BISH, iso), Mindanao: Agusan del Norte, Cabadbaran, Mt Urdaneta, Sep-1912. Endemic to the Philippines. LUZON: Tayabas, Camarines, NEGROS: Negros Occidental (Mt Marapara), LEYTE, MINDANAO: Agusan del Norte (Agusan River valley near Amparo). Low elevation primary forests. Photos
  10. Discocalyx leytensis Merr., PJS 26 (1925) 485; EPFP 4 (1925) 250; --Type: BS 41709 Edaño (US, iso), Leyte: Mt Abucayan. Feb-1923. Endemic to the Philippines. LEYTE. In primary forests.
  11. Discocalyx linearifolia Elmer, LPB 2 (1908) 441; --Merr., EPFP 3 (1923) 268; --Type: Elmer 7674 (A, iso), Luzon: Quezon Prov, Lucban, May-1907. Endemic to the Philippines. LUZON: Ilocos Norte, Abra, Kalinga, Mountain Province, Sierra Madre Range from Isabela to Quezon, Laguna, Sorsogon, POLILLO, CATANDUANES, SAMAR, SIARGAO. Photos
  12. Discocalyx longifolia Merr., PJS 7 c (1912) Bot. 325; EPFP 3 (1923) 268; --Type: BS 13181 Foxworthy & Ramos, Luzon: Quezon Prov, Quinatacutan, Mar-1911. Endemic to the Philippines. LUZON: Quezon. Low elevation forests.
  13. Discocalyx luzoniensis Merr., PJS 12 c (1917) Bot. 145; EPFP 3 (1923) 268; --Type: BS 26474 (US, iso), Luzon: Nueva Ecija Prov, Mt Umingan, forested slopes between 300-400m, 16-Aug-1916. Endemic to the Philippines. LUZON: Nueva Ecija (Mt Umingan). Primary forests, c. 350m.
  14. Discocalyx macrophylla Merr., PJS 5 c (1910) Bot. 222; EPFP 3 (1923) 268; --Type: FB 17237 Curran (US, iso), Luzon: Cagayan Prov, near San Vicente, forest at sea level, 8-Mar-1909. Endemic to the Philippines. LUZON: Ilocos Norte, Cagayan. Low elevation forests.
  15. Discocalyx maculata Merr., PJS 7 c (1912) Bot. 326; EPFP 3 (1923) 268; --Type: BS 13967 Ramos, Luzon: Cagayan Prov, Abulug River, in forest, Feb-1912. Endemic to the Philippines. LUZON: Cagayan. Low elevation forests.
  16. Discocalyx merrillii Mez, PJS 1 (1906) Suppl. 274; --Merr., EPFP 3 (1923) 268; --Type: Merrill 3745 (US, syn), Luzon: Bataan Prov, Mt Mariveles, Jan-1904; Whitford 213 (syn); FB 139 Barnes (syn); FB 1348 Borden (syn), Luzon: Bataan Prov, Mt Mariveles along Lamao River. Endemic to the Philippines. LUZON: Cagayan, Pangasinan, Zambales, Bataan, Bulacan, Rizal, Quezon. Medium elevation primary forests, ascending to 1000m.
  17. Discocalyx micrantha Merr., PJS 12 c (1917) Bot. 146; EPFP 3 (1923) 268; --Type: BS 27132 Ramos (US, iso), Luzon: Ilocos Norte Prov, Burgos. Low elevation forests along stream, 12-Mar-1917. Endemic to the Philippines. LUZON: Ilocos Norte, Isabela (Palanan), MINDANAO: Surigao. Low elevation primary forests. Photos
  18. Discocalyx mindanaensis Elmer, LPB 2 (1910) 674; --Merr., EPFP 3 (1923) 269; --Type: Elmer 11294 (PNH, holo, lost; A, GH, MO, US, iso), Mindanao: Davao, Mt Apo, Todaya, Aug-1909. Endemic to the Philippines. MINDANAO: Lanao del Sur (Camp Keithley), Bukidnon, Davao (Mt Apo), Agusan del Norte. Medium elevation primary forests, ascending to 1200m.
  19. Discocalyx minor Mez in Engl., Pflanzenreich 9 (1902) 213; --Merr., EPFP 3 (1923) 269; --Type: Comision de la Flora Forestal de Filipinas 427 (L, iso), Luzon: Bulacan Prov, Angat. Endemic to the Philippines.
  20. Discocalyx montana Elmer, LPB 2 (1908) 443; --Merr., EPFP 3 (1923) 269. Endemic to the Philippines. LUZON: Ilocos Norte, Apayao, Cagayan, Mountain Province, Benguet, Nueva Vizcaya, Quezon, Laguna, MINDORO. Stream gullies and forested slopes, 1000-2000m. Photos
  21. Discocalyx pachyphylla Merr., PJS 12 c (1917) Bot. 146; EPFP 3 (1923) 269. Endemic to the Philippines. LUZON: Ilocos Norte, Quezon (Guinayanagan). Damp forested slopes at medium elevation, ascending to 1000m.
  22. Discocalyx palawanensis Elmer ex Merr., EPFP 3 (1923) 269; --Discocalyx merrillii auct. non Mez; --Elmer, LPB 5 (1913) 1821; --Type: Elmer 12825 (PNH, holo, lost; A, MO, US, iso), Palawan: Mt Pulgar (Thumb Peak), Mar-1911. Endemic to the Philippines. MINDORO, PALAWAN, BANCALAN. Low elevation primary forests. Conservation status: Other Threatened Species (DAO 2017-11).
  23. Discocalyx phanerophlebia Merr., PJS 20 (1922) 425; EPFP 3 (1923) 269; --Type: BS 38690. Endemic to the Philippines. MINDANAO: Bukidnon. Forests along streams, 1200-1400m.
  24. Discocalyx philippinensis (A.DC.) Mez in Engl., Pflanzenreich 9 (1902) 212; --Merr., EPFP 3 (1923) 269; --Badula philippinensis A.DC., Prodr. 8 (1844) 108. Endemic to the Philippines. LUZON: Ilocos Norte, Mountain Province, Benguet, Nueva Ecija, Pampanga, Bataan, Rizal, Laguna, Batangas, PANAY. Damp ravines and primary forests, extending into the mossy forest, 600-1400m.
  25. Discocalyx psychotrioides Elmer, LPB 2 (1908) 442; --Merr., EPFP 3 (1923) 270. Endemic to the Philippines. LUZON: Isabela, Rizal, Laguna, Quezon. Primary forests, 300-800m. Photos
  26. Discocalyx samarensis Merr., PJS 12 c (1917) Bot. 147; EPFP 3 (1923) 270; --Type: FB 24004 Lasquety, Samar: Northern Samar Prov, San Jose de Buan, along streams, 300m, 7-May-1915. Endemic to the Philippines. SAMAR: Northern Samar (San Jose de Buan). Lowland forest, 300m.
  27. Discocalyx sessilifolia Merr., PJS 12 c (1917) Bot. 149; EPFP 3 (1923) 270; --Type: FB 20735 Rafael & Ponce (US, iso), Mindanao: Agusan del Norte Prov (indicated in label as Butuan Subprovince), near Butuan (according to protologue). Swamp forest at low elevation, 12-Sep-1913. Endemic to the Philippines. MINDANAO: Agusan del Norte. Swamp forest at low elevation.
  28. Discocalyx stenophylla Merr., PJS 12 c (1917) Bot. 148; EPFP 3 (1923) 270; --Type: FB 24815 Pascual (US, iso), Luzon: Pangasinan Prov, Mangatarem, among undergrowth in forests, c. 300m, 23-Nov-1915. Endemic to the Philippines. LUZON: Ilocos Norte, Pangasinan, Rizal. Primary forests, 300-700m.
  29. Discocalyx suluensis Merr., PJS 30 (1926) 420; --Type: BS 44293 Ramos & Edaño (US, iso), Tawi-tawi, damp forests at low elevation, Jul-1924. Endemic to the Philippines. TAWI-TAWI. Lowland forests.
  30. Discocalyx tecsonii Merr., PJS 12 c (1917) Bot. 150; EPFP 3 (1923) 270; --Type: FB 24683 Tecson (US, iso), Basilan: Tumakis, in forest, 20m, locally common, teste Tecson, 4-Nov-1915. Endemic to the Philippines. BASILAN (Tumakis; Mt Singal), MINDANAO: Misamis Occidental (Mt Malindang). Low elevation primary forests, 10-100m.
  31. Discocalyx vidalii Mez in Engl., Pflanzenreich 9 (1902) 213; --Merr., EPFP 3 (1923) 270; --Type: Vidal 423, Luzon: Quezon Prov, Mt Banahaw. Endemic to the Philippines. LUZON: Quezon (Mt Banahaw). Perhaps also represented by BS 33312 Ramos, 29288 Ramos & Edaño; from primary forests, c. 300m.
  32. Discocalyx xiphophylla Quisumb. & Merr., PJS 37 (1928) 185; --Type: BS 45678 Ramos & Edaño, Luzon: Aurora Prov, Mt Alzapan, in forests, c. 1800m, 6-Jun-1925. Endemic to the Philippines. LUZON: Aurora (Mt Alzapan). Montane forests, c. 1800m.  Comments from the original protologue: Belonging to the group of Discocalyx linearifolia Elmer, but totally different in its vegetative and inflorescence characters. Notably differs from Discocalyx angustissima Merr. in its entire leaves.

Embelia Burm.f.

Last edited by Pieter B. Pelser, 18 December 2019

Photos
  1. Embelia bataanensis Merr., Govt. Lab. Publ. (Philipp.) 29 (1905) 44; EPFP 3 (1923) 271; --Type: Merrill 3207 (US, iso), Luzon: Bataan Prov., Mt Mariveles, 21-Oct-1903. Endemic to the Philippines. LUZON: Mountain Province, Benguet, Bataan. Ravines and forested ridges, 600-1300m.
  2. Embelia coriacea Wall. ex A.DC., Trans. Linn. Soc. 17 (1834) 135; --Wall., Cat. (1830) No. 2314, nom. nud.; --Merr., EPFP 3 (1923) 271. Sumatra, Peninsular Malaysia, Java, Borneo, Moluccas, Singapore, Thailand, Philippines. MINDANAO: Lanao del Sur, Bukidnon, Davao, Agusan del Norte. Low and medium elevation primary forests.
  3. Embelia elliptica Merr., PJS 20 (1922) 426; EPFP 3 (1923) 271; --Type: BS 39069 Ramos & Edaño. Endemic to the Philippines. MINDANAO: Bukidnon. Streamside thickets, 900m.
  4. Embelia loheri Merr., PJS 27 (1925) 45; EPFP 4 (1925) 253; --Type: Loher 13845 (PNH, holo, ?lost; UC, iso), Luzon: Rizal Prov, Mt Oriud, Mar-1914. Endemic to the Philippines. LUZON: Rizal (Mt Paningtingan; Mt Oriud).
  5. Embelia luzoniensis Merr., PJS 20 (1922) 427; EPFP 3 (1923) 271; --Type: BS 20098 McGregor. Endemic to the Philippines. LUZON: Nueva Vizcaya. Medium elevation thickets or forests.
  6. Embelia nigropunctata Merr., PJS 7 c (1912) Bot. 326; EPFP 3 (1923) 271. Endemic to the Philippines. LUZON: Bataan, Rizal, MINDORO, PALAWAN. Low and medium elevation thickets and forests. Photos
  7. Embelia obtusiuscula Mez, Fedde Repert. 16 (1920) 422; --Merr., EPFP 3 (1923) 271; --Type: Elmer 5881. Endemic to the Philippines. LUZON: Benguet. Steep slopes in thickets, c. 1200m.
  8. Embelia ovatifolia Merr., PJS 20 (1922) 428; EPFP 3 (1923) 272; --Type: BS 39001 Ramos & Edaño. Endemic to the Philippines. MINDANAO: Bukidnon. Forested slopes, c. 1200m.
  9. Embelia philippinensis A.DC., Prodr. 8 (1844) 83; --Merr., EPFP 3 (1923) 272. --Rhamnus lando Llanos, Frag. Fl. Philip. (1851) 57; --Fern.-Vill. & Naves in Blanco, Fl. Filip. ed. 3, 41 (1880) 41. --Samara philippinensis S.Vidal, Sinopsis Atlas (1883) 30. Borneo, Philippines. LUZON: Benguet, Pangasinan, Nueva Vizcaya, Nueva Ecija, Bulacan, Rizal. Laguna, Sorsogon. LEYTE, MINDANAO: Lanao, Bukidnon, Davao. Medium elevation forests, often common. Photos
  10. Embelia whitfordii Merr., PJS 1 (1906) Suppl. 113; EPFP 3 (1923) 272; --Type: Whitford 453. --Embelia urdanetensis Elmer, LPB 8 (1915) 2783; --Merr., EPFP 3 (1923) 272; --Type: Elmer 13745. Endemic to the Philippines. LUZON: Mountain Province, Benguet, Bataan, Laguna, MINDANAO: Agusan del Norte.

Labisia Lindl.

Last edited by Pieter B. Pelser, 22 September 2017

Photos
  1. Labisia pumila (Blume) Fern.-Vill., Novis. App. (1880) 123; --Merr., EPFP 3 (1923) 266; --Ardisia pumila Blume, Catalogus (1823) 44. Indochina, Sumatra, Peninsular Malaysia, Java, Borneo, Philippines. MINDANAO, MINDORO, LEYTE, JOLO. Damp primary forests up to 500m. Very rare and local in the Philippines. Photos

Loheria Merr.

Last edited by Pieter B. Pelser, 31 May 2018

Photos
  1. Loheria bracteata Merr., PJS 5 c (1910) Bot. 374; EPFP 3 (1923) 270; --Type: Merrill 4783. Endemic to the Philippines. LUZON: Mountain Province, Benguet. Mossy forests, 1900-2400m. Photos
  2. Loheria crassifolia (Merr.) Stone, Micronesica 24 (1991) 72; --Amblyanthopsis crassifolia Merr., PJS 14 (1919) 445; EPFP 3 (1923) 266. Endemic to the Philippines. LUZON: Ilocos Norte, Aurora. Forested slopes, c. 1000m. Photos
  3. Loheria jubilaria Stone, Micronesica 24 (1991) 74; --Loheria radlkoferi (Mez) Merr., EPFP 3 (1923) 271, nom. illeg.; --Jubilaria radlkoferi Mez, Fedde Repert. 16 (1920) 421; --Type: Merrill 5743 (US, iso), Mindoro: Mindoro Oriental Prov, Alag River, Nov-1906. Endemic to the Philippines. MINDORO. Low elevation damp primary forests.
  4. Loheria porteana (Mez) Merr., EPFP 3 (1923) 271; --Embelia porteana Mez, Pflanzenr. 4, 236 (1902) 302. Endemic to the Philippines. LUZON: Quezon, Rizal, POLILLO. Low elevation damp primary forests.

 Rapanea Aubl. = Myrsine

Lysimachia L.

Last edited by Pieter B. Pelser, 22 September 2017

Photos
  1. Lysimachia capillipes Hemsl., J. Linn. Soc. Bot. 26 (1889) 48; --Merr., EPFP 3 (1923) 274; --Bentvelzen, Fl. Males. ser. 1, 6 (1962) 179. China (Yunnan; Guizhou; Sichuan; Hubei), Taiwan, N Philippines. N LUZON. Damp thickets along streams, 1300-1500m.
  2. Lysimachia decurrens G.Forst., Prodr. (1786) 12; --Merr., EPFP 3 (1923) 274; --Bentvelzen, Fl. Males. ser. 1, 6 (1962) 185. E Himalaya, Myanmar, Thailand, Indochina, S China, Japan, Taiwan, W Java, Lesser Sunda Isls (Timor), N Philippines, Vanuatu, New Caledonia. LUZON: Benguet.
  3. Lysimachia japonica Thunb., Fl. Jap. (1784) 83; --Merr., EPFP 3 (1923) 275; --Bentvelzen, Fl. Males. ser. 1, 6 (1962) 182.
  4. Lysimachia mauritiana Lam., Encycl. 3 (1789) 592; --Merr., EPFP 3 (1923) 275; --Bentvelzen, Fl. Males. ser. 1, 6 (1962) 183. Mauritius, Réunion, E China, Korean islands, Japan, Ryukyu Isls, Taiwan, N Philippines, Bonin Isls, Marianas, Hawaiian Isls, New Caledonia, Loyalty Isls. BATANES. Coastal coral rocks and beaches. Photos
  5. Lysimachia sikokiana Miq., Ann. Mus. Bot. Lugd.-Bat. 3 (1867) 121; --Bentvelzen, Fl. Males. ser. 1, 6 (1962) 178. --Lysimachia ramosa Wall. ex Duby in DC., Prodr. 8 (1844) 65; --Merr., EPFP 3 (1923) 275; --Wall., Cat. (1828) No. 1490, nom. nud. S Japan, Ryukyu Isls, Taiwan, S China (Yunnan; Guangdong), Hainan, N Thailand, Philippines. LUZON: Mountain Province, MINDORO, NEGROS, CAMIGUIN. Mossy forests, damp ravines, 1200-2300m. Photos

Maesa Forssk.

Last edited by Pieter B. Pelser, 31 May 2018

Photos
  1. Maesa cumingii Mez in Engl., Pflanzenreich  9 (1902) 49; --Merr., EPFP 3 (1923) 253; --Utteridge & Saunders, Bot. J. Linn. Soc. 145 (2004) 20; --Type: Cuming 1474 (K, lecto; BM, K, W, iso), Luzon: Camarines Sur Prov, 1836-1839. --Maesa pachyphylla Merr., PJS 7 c (1912) 328; EPFP 3 (1923) 255; --Type: BS 11030 Ramos (PNH, holo, lost; BM, lecto; A (fragment), isolecto), Cebu: Hills behind the City of Cebu (Merrill 1912), Mar-1912. --Maesa undulata Merr., PJS 20 (1922) 422; EPFP 3 (1923) 255; --Type: BS 26872 Edaño (PNH, holo, lost; K, lecto), Luzon: Quezon Prov., near Pagbilao, Mar-1917. Endemic to the Philippines, but widespread throughout the archipelago. Along clearings and streams, secondary growth; 50-900m. Fl. Oct-May; fr. throughout the year. Conservation status: Not Threatened (Utteridge & Saunders 2004). Photos
  2. Maesa davaensis Quisumb., PJS 41 (1930) 341; --Utteridge & Saunders, Bot. J. Linn. Soc. 145 (2004) 22; --Type: BS 49123 Ramos & Edaño (PNH, holo, lost; UC, lecto; A, NY, P, iso), Mindanao: Davao Oriental Prov, Mati, Dawau Streams, 9-Apr-1927. MINDANAO: Davao Oriental. Low elevation forests. Fl. & fr. Apr.  This species is only known from a single locality, from a single collection made in 1927. Conservation status: Critically Endangered B1a+b(iii) (Utteridge & Saunders 2004).
  3. Maesa denticulata Mez in Engl., Pflanzenreich 9 (1902) 48; --Merr., EPFP 3 (1953) 253; --Utteridge & Saunders, Bot. J. Linn. Soc. 145 (2004) 23; --Type: Warburg 15025 (E, lecto; B, isolecto, lost), Sulu Archipelago: 1888. --Maesa vulcanica Elmer in Merr., EPFP 3 (1923) 253 (1923), nom. nud.; --Voucher: Elmer 14966 (GH, L, PNH, UC), Luzon: Sorsogon Prov, Irosin (Mt Bulusan), Nov-1915. Borneo (Sabah), Sulawesi, Philippines. BATAN and N LUZON to PALAWAN and MINDANAO. Forest edges, disturbed forests, secondary forests, lowland, submontane and mossy forests; 100-1550m. Fl. Dec-Aug; fr. throughout the year. Conservation status: Least Concern (Utteridge & Saunders 2004). Photos
  4. Maesa ferruginea Merr., PJS 7 c (1912) 327; EPFP 3 (1923) 254; --Utteridge & Saunders, Bot. J. Linn. Soc. 145 (2004) 24; --Type: BS 7109 Ramos (PNH, holo, lost; NY, lecto), Luzon: Abra Prov, Mt Paraga, 7-Feb-1909. Endemic to the Philippines. LUZON (Abra). In forest, c. 3000m. Only known from two collections. Fl. Feb, May; fr. May. Maesa ferruginea is only known from Abra Province (c. 4000km2). The narrow distribution range of the species, combined with the forest habitat degradation in the Philippines, suggests that Maesa ferruginea faces a very high risk of extinction in the wild. Conservation status: Endangered B1a+b(iii) (Utteridge & Saunders 2004).
  5. Maesa gaudichaudii A.DC., Ann. Sci. Nat. Bot. Sér. II, 16 (1841) 80; --Merr., EPFP 3 (1923) 254; --Utteridge & Saunders, Bot. J. Linn. Soc. 145 (2004) 25; --Type: Gaudichaud s.n. (G- DEL, holo), Luzon: Manila, Dec-1836. --Maesa piscatorum Mez, Repert. Spec. Nov Regni Veg. 16 (1920) 309; --Type: BS 19 Foxworthy (PNH, holo, lost; K, lecto; BO, GH, NSW, NY, PNH lost, isolecto), Luzon: Rizal Prov, Jan-1906. Endemic to the Philippines. LUZON, MINDORO, MINDANAO, NEGROS, PALAWAN. Forest edges, secondary growth; 100-300m. Fl. & fr. throughout the year. Conservation status: Not Threatened (Utteridge & Saunders 2004).
  6. Maesa haenkeana Mez in Engl., Pflanzenreich 9 (1902) 32; --Merr., EPFP 3 (1923) 254; --Utteridge & Saunders, Bot. J. Linn. Soc. 145 (2004) 26; --Type: Haenke 471 (PR, lecto; NY, isolecto), Luzon: Sorsogon Prov, 1792. --Maesa embelioides Elmer, LPB 2 (1908) 444; --Type: Elmer 7459 (B, holo, lost; K, lecto; A, E, L, LE, isolecto), Luzon: Quezon, Lucban, May-1906. --Maesa longipetiolata Merr., PJS 11 c (1916) 28; EPFP 3 (1923) 254; --Type: BS 23358 Ramos (PNH, holo, lost; L, lecto; A, BO, K, NY, P, isolecto), Luzon: Sorsogon Prov, Mt Pacdal, 5-Aug-1915. --Maesa brunnea Merr., PJS 13 c (1918) Bot. 321; EPFP 3 (1923) 253; --Type: BS 30500 Ramos (PNH, holo, lost; K, lecto; A, P, isolecto), Catanduanes: Mt Mariguidon, 14-Nov to 11-Dec-1917. --Maesa cuprifolia Elmer in Merr., EPFP 3 (1923) 254, nom. nud.; --Voucher: Elmer 16521 (A, K, L, PNH, US), Luzon: Sorsogon Prov, Irosin (Mt Bulusan), Jul-1916. --Maesa platyphylla var. minor Elmer in Merr., EPFP 3 (1923) 254, nom. nud.; --Voucher: Elmer 15558 (A, BO, GH, L), Luzon: Sorsogon Prov, Irosin (Mt Bulusan), Nov-1915. --Maesa irosinensis Elmer, LPB 10 (1939) 3765, nom. inval., no Latin descr.; --Utteridge & Saunders, Bot. J. Linn. Soc. 145 (2004) 37; --Type: Elmer 15558 (PNH, holo, lost; US, lecto; A 2 sheets, BO, C, GH, L, MO, NY 2 sheets, isolecto), Luzon: Sorsogon Prov, Mt Bulusan, Nov-1915. Endemic to the Philippines. LUZON: Quezon, Camarines, Albay, Sorsogon, CATANDUANES, SAMAR. Lowland forests; 150-760m. F. Mar-Jul; fr. throughout the year. Conservation status: Not Threatened (Utteridge & Saunders 2004).
  7. Maesa indica (Roxb.) A.DC., Trans. Linn. Soc. 17 (1834) 133; --Utteridge & Saunders, Bot. J. Linn. Soc. 145 (2004) 29; --Baeobotrys indica Roxb., Fl. Ind. 2 (1824) 230. --Maesa laxa Mez in Engl., Pflanzenreich 9 (1902) 32; --Merr., EPFP 3 (1923) 254; --Type: Cuming 983 (P, lecto; BM, E, K 2 sheets: Herb. Benth. & Herb. Hook., LE, MO, W 2 sheets, isolecto), Luzon: Pangasinan Prov, 1836-1839. --Maesa elmeri Mez, PJS 1 (1906) Suppl. 271; --Merr., EPFP 3 (1923) 253; --Type: Elmer 5925 (US, iso), Luzon: Benguet Prov, Baguio, Mar-1904. --Maesa grossedentata Mez, Repert. Spec. Nov. Regni Veg. 16 (1920) 309; --Merr., EPFP 3 (1923) 255; --Type: BS 1769 Ramos (PNH, holo, lost; NY, lecto), Luzon: Rizal Prov, Jan-1907. --Maesa ramosii Quisumb. & Merr., PJS 37 (1928) 186; --Type: BS 47272 Ramos & Edaño (PNH, holo, lost; A, lecto; K, NY, isolecto), Luzon: Isabela Prov, Mt Moises, 4-Mar-1926. Widespread in SE Asia, occurring throughout the Philippines. Secondary forests, mossy forests, and upper pine forests; 100-2000m. Fl. & fr. throughout the year. Conservation status: Least Concern (Utteridge & Saunders 2004). Photos
  8. Maesa manillensis Mez in Engl., Pflanzenreich 9 (1902) 34; --Utteridge & Saunders, Bot. J. Linn. Soc. 145 (2004) 31; --Type: Calléry 2 (P, holo), Luzon: ?Mountain Province, Mt Igorrotes, May-1840. Endemic to the Philippines. N LUZON: Ilocos Norte, Mountain Province. Forests; 150-1500m. Fl. Feb-Mar, Nov; fr. Aug, Oct, Nov. Conservation status: Not Threatened (Utteridge & Saunders 2004).
  9. Maesa megalobotrya Merr., PJS 20 (1922) 422; EPFP 3 (1923) 255; --Utteridge & Saunders, Bot. J. Linn. Soc. 145 (2004) 32; --Type: Merrill 9176 (PNH, holo, lost; L, lecto; BO, K, NSW, NY, P, isolecto), Palawan: Taytay, Apr-1913. Endemic to the Philippines. PALAWAN. Closed broad-leaved forests over ultrabasic rock; 490-590m. Fl. Apr; fr. Apr-May. This species has only been collected twice. Conservation status: Vulnerable B1ab(iii) (Utteridge & Saunders 2004).
  10. Maesa megaphylla Merr., PJS 12 c (1917) Bot. 158; EPFP 3 (1923) 255; --Utteridge & Saunders, Bot. J. Linn. Soc. 145 (2004) 33; --Type: Wenzel 1510 (PNH, holo, lost; A 2 sheets, lecto; BM, GH 2 sheets, NSW, PNH, isolecto), Leyte: Tigbao near Tacloban, 3-Aug-1915. BOHOL, LEYTE and MINDANAO. Dry forests; sea level to 600m. Fl. Jun, Aug; fr. Feb, Apr, Aug, Oct. This species is only known from seven collections, the last of which was made in 1923. Conservation status: Vulnerable B1ab(iii) (Utteridge & Saunders 2004).
  11. Maesa platyphylla Elmer, LPB 2 (1910) 667; --Utteridge & Saunders, Bot. J. Linn. Soc. 145 (2004) 34; --Type: Elmer 10570 (PNH, holo, lost; NY, lecto; E, GH 2 sheets, K, L, LE, MO, isolecto), Mindanao: Davao, Mt Apo, Todaya, May-1909. MINDANAO: Davao (Mt Apo). Primary forests; 910-1200m. Only known from two collections. Fl. May; fr. May, July and August. Conservation status: Not Threatened (Utteridge & Saunders 2004).
  12. Maesa reflexa Utteridge & R.M.K.Saunders, Kew Bull. 55 (2000) 981; Bot. J. Linn. Soc. 145 (2004) 34; --Type: BS 38973 Ramos & Edaño (A, holo; K, iso), Mindanao: Bukidnon Prov, Mt Dumalucpihan, June-July 1920. Endemic to the Philippines. MINDANAO: Bukidnon. Habitat information not available but presumably from montane forests; c. 2350m. Fl. Jun-Jul; fr. Apr, Jun-Jul. Maesa reflexa is known from only two localities (three collections made in 1920 and 1949) from Bukidnon province (c. 9000 km2). The lack of collections from other areas on MINDANAO (especially Mt Apo, which has been relatively well collected) implies that this species has a very restricted distribution in specific mountain ranges. Conservation status: Vulnerable B1ab(iii) (Utteridge & Saunders 2004).
  13. Maesa stonei Utteridge & R.M.K.Saunders, Bot. J. Linn. Soc. 145 (2004) 37; --Type: PNH 109009 Gutierrez, Yen & Reynoso (PNH, holo; BISH, iso), Mindanao: South Cotabato Prov, Tasaday forest, Jul/Aug-1972. MINDANAO: South Cotabato. Dipterocarp forests, c. 1220m. Fl. & fr. Jul-Aug. Maesa stonei is only known from a single locality, in an area which has not been extensively collected; the species is furthermore only known from one collection (dating from 1972). Conservation status: Data Deficient (Utteridge & Saunders 2004).

Doubtful and excluded names:

  1. Maesa lobuligera Mez, Repert. Spec. Nov. Regni Veg. 16 (1920) 310; Merr., PJS 20 (1922) 421; --Utteridge & Saunders, Bot. J. Linn. Soc. 145 (2004) 39; --Type: BS 727 Foxworthy (PNH, holo, lost), Palawan. See Utteridge & Saunders (2004).
  2. Maesa membranacea A.DC., Ann. Sci. Nat. Bot. Sér. II, 16 (1841) 80. --Utteridge & Saunders, Bot. J. Linn. Soc. 145 (2004) 39; --Type: from Vietnam, 1836-1837, Gaudichaud, number not cited (G-DEL 2 sheets, syn). See Utteridge & Saunders (2004).
  3. Maesa philippinensis Gaudich., Bull. Soc. Bot. France 60 (1913) 24. --Utteridge & Saunders, Bot. J. Linn. Soc. 145 (2004) 39; --Type: from Peninsular Malaysia. Malacca, 1839-1840, Cuming 2286 (L, iso). This is Maesa ramentacea (Roxb.) A.DC., see Utteridge & Saunders (2004).
  4. Maesa tenera Mez in Engl., Pflanzenreich 9 (1902) 28; --Hatusima, Mem. Fac. Agr. Kagoshima Univ. 5 (1966) 43. S Japan, Ryukyus, Taiwan, S China to Indochina, Philippines. BATAN (Mt Iraya). In thickets on the ridge of Mt Iraya, 400m. Locally rare. Possibly synonymous with Maesa indica (Roxb.) A.DC. (Utteridge, pers. comm. 2-Jan-18).

Myrsine L.

Last edited by Pieter B. Pelser, 31 May 2018

Photos
  1. Myrsine amorosoana Pipoly, Sida 17 (1996) 126; --Rapanea retusa Merr., PJS 2 c (1907) Bot. 297, nom. illeg., auct. non Aiton; EPFP 3 (1923) 274; --Type: Merrill 5734 (US, lecto; K, NY, isolecto). Endemic to the Philippines. LUZON: Benguet (Mt Pulag), Quirino (Mt Alzapan), Aurora (Mt Dingalan), Quezon (Mt Binuang; Quezon National Forest Park, Atimonan), Rizal (Angilog), Camarines Norte, Camarines Sur (Bicol National Park; Mt Isarog), Sorsogon, MINDORO: Mindoro Oriental (N coast, Subaan River headwaters, inland from San Teodoro; Mt Halcon), SIBUYAN (Mt Giting-giting), NEGROS: Negros Occidental (Mt Kanlaon), MINDANAO: Bukidnon (Mt Candoon; Mt Lipa; Kitanglad Range at Mt Kinasalapi and Mt Dulang-dulang). Mid- to high-montane mossy forests, 1400-2825m. Photos
  2. Myrsine apoensis (Elmer) Pipoly, Sida 17 (1996) 125; --Rapanea apoensis Elmer, LPB 2 (1910) 669; --Merr., EPFP 3 (1923) 273; --Type: Elmer 10629 (A, lecto; BISH, GH 2 sheets, K, L, NY, isolecto), Mindanao: Davao, Mt Apo, Todaya, May-1909. Endemic to the Philippines. MINDANAO: Davao (Mt Apo, Todaya; Mt McKinley), North Cotabato (Mt Apo, Kidapawan, Geothermal Project Site). High-montane mossy forest, 1666-2900m. 
  3. Myrsine aralioides (Philipson) Pipoly, Sida 17 (1996) 152; --Rapanea aralioides Philipson, J. Bot. (London) 78 (1939) 104; --Type: from Sabah. Borneo (Sabah), Philippines. MINDANAO: Davao Oriental (Mt Hamiguitan; Mati, South Basiao; along Biga Creek), Surigao (del Sur Prov?), PALAWAN (Mt Bloomfield). 150-200m in Palawan. Seen at c. 1200m on Mt Hamiguitan. On ultramafics.
  4. Myrsine avenis (Blume) A.DC., Trans. Linn. Soc. 17 (1834) 107; --Pipoly, Sida 17 (1996) 144; --Rapanea avenis (Blume) Mez in Engl., Pflanzenreich 9 (1902) 357; --Merr., EPFP 3 (1923) 273; --Ardisia avenis Blume, Bijdr. (1825) 691. --Type: from Java. Sumatra, Java, Buru, Borneo, Philippines, Sulawesi. LUZON: Ilocos Norte (Mt Palimlim), Abra, Mountain Province, Ifugao (Banawe, Bayninan), Benguet (Mt Pulogloco; Mt Sto Tomas; Baguio and vicinity), Bataan (Mt Mariveles, Lamao River), Rizal (Mt Susong Dalaga; Sumag; Montalban), Laguna, Quezon, Camarines Norte (Paracale), Camarines Sur, MINDORO, NEGROS, MINDANAO: Bukidnon (Mt Kitanglad), Davao (Mt Apo). Hill to mossy forests and wind-swept ridges, (300-)1400-2400m.
  5. Myrsine cruciata (Philipson) Pipoly, Sida 17 (1996) 130; --Rapanea cruciata Philipson, J. Bot. (London) 77 (1939) 105; --Type: from Mt Kinabalu, Sabah. Philippines, Borneo (Sabah: Mt Kinabalu). LUZON: Abra, Zambales, PALAWAN (Mt Beaufort). At least recorded between 490-815m in Palawan, on presumably ultramafic habitat.
  6. Myrsine densiflora Scheffer, Comm. Myrs. Archipelago Ind. (1876) 50; --Pipoly, Sida 17 (1996) 132. Philippines, Sulawesi, Moluccas, Papua New Guinea. BATAN, LUZON: Pangasinan, Zambales, PALAWAN, PANAY: Capiz, NEGROS, SAMAR, DINAGAT, MINDANAO: Surigao. Photos
  7. Myrsine fastigiata (Elmer) Pipoly, Sida 17 (1996) 149; --Rapanea fastigiata Elmer, LPB 2 (1910) 670; --Merr., EPFP 3 (1923) 273; --Type: Elmer 11447 (GH, lecto; A, K, L, NY, isolecto), Mindanao: Davao, Mt Apo, Todaya, 2000m, Aug-1909. Endemic to the Philippines. LUZON: Ifugao (Mt Polis), Mountain Province, Benguet (Atok, Pauai and vicinity; Mt Tabayoc; Mt Pulag; Mt Singakalsa; Mt Nangaoto; Baguio; Mt Sto Tomas), Zambales (Sta Cruz),  Bataan (Mt Mariveles, upper Lamao River), Camarines Norte (Paracale), BILIRAN (Mt Suiro), MINDANAO: Bukidnon (Mt Lipa), Davao (Mt Apo). Low to high montane sites, but most conspicuous at 2000-2500m.
  8. Myrsine glandulosa (Elmer) Pipoly, Sida 17 (1996) 154; --Rapanea glandulosa Elmer, LPB 8 (1915) 2784; --Merr., EPFP 3 (1923) 273; --Type: Elmer 14148 (A, lecto; BISH, GH, L, K, NY, isolecto), Mindanao: Agusan del Norte Prov, Cabadbaran, Mt Urdaneta, 1524m, Oct-1912. Endemic to the Philippines. NEGROS: Negros Occidental (Mt Kanlaon, SW slopes), Negros Oriental (Cuernos Mtns), LEYTE, MINDANAO: Bukidnon (Malaybalay, Kaatoan), Davao del Sur (Mt McKinley), South Cotabato (Mt Matutum), Agusan del Norte. Montane forests, 1350-2242m.
  9. Myrsine medeciloae Pipoly, Sida 17 (1996) 122; --Rapanea angustifolia Merr., PJS 20  (1922) 429 (non Myrsine angustifolia (A.DC.) D.Dietr., nec Myrsine angustifolia E.Meyer ex Schimper (nom. nud.), nec Myrsine angustifolia Heller, nec Myrsine angustifolia (Mez) Hosaka); --Type: BS 33243 Ramos (A, lecto), Luzon: Ilocos Norte Prov, Mt Nagapatan, Aug-1918. Endemic to the Philippines. LUZON: Ilocos Norte (Mt Nagapatan; Burgos; Sicapoo Range, en-route from miner’s camp to Mt Burnay), Zambales (Mt Tapulao), Bataan (Mt Mariveles, Lamao River). Submontane mossy forests, 1000-1800m.
  10. Myrsine mindanaensis (Elmer) Pipoly, Sida 17 (1996) 140; --Rapanea mindanaensis Elmer, LPB 2 (1910) 671; --Merr., EPFP 3 (1923) 273; --Type: Elmer 11332 (K, lecto; A, BISH, GH, L, NY, isolecto), Mindanao: Davao, Mt Apo, Todaya, Aug-1909. Endemic to the Philippines. LUZON: Camarines Sur (Mt Isarog), BILIRAN: Biliran (Naval, Libtong), LEYTE: Leyte (Cabalian), MINDANAO: Davao del Sur; Davao-North Cotabato (Mt Apo). Low to high montane forests, 895-2200m. Photos
  11. Myrsine oblongibacca (Merr.) Pipoly, Sida 17 (1996) 129; --Rapanea oblongibacca Merr., PJS 20 (1922) 429; EPFP 3 (1923) 273; --Type: BS 33256 Ramos (A, lecto; K, US, isolecto), Luzon: Ilocos Norte Prov, Mt Palimlim, Aug-1918. Endemic to the Philippines. LUZON: Ilocos Norte (Mt Palimlim), Cagayan (Mt Cagua; Mt Dos Cuernos; Mt Tabuan, Bauan district; Mt Balatongan), Nueva Ecija (Mt Umingan = Mingan), Aurora (Mt Dingalan), Rizal (Mt Susong Dalaga), DINAGAT. Possibly a submontane species at perhaps 800-1500m elevation. In Ilocos Norte and Dinagat possibly growing on ultramafics.
  12. Myrsine penibukana (Philipson) Pipoly, Sida 17 (1996) 147; --Rapanea penibukana Philipson, London J. Bot. 78 (1939) 105; --Type: from Mt Kinabalu, Sabah. Borneo (Mt Kinabalu), Philippines. LUZON: Abra (Mt Posuey), Aurora (Mt Dingalan), Rizal. 1000-1200m.
  13. Myrsine peregrina (Mez) Pipoly, Sida 17 (1996) 136; --Rapanea peregrina Mez, Repert. Spec. Nov. Regni Veg. 16 (1920) 424; --Type: M.S. Clemens 917 (US, lecto; A, isolecto), Mindanao: Lanao del Sur Prov, Camp Keithley, Lake Lanao, Jun-1907. --Rapanea venosa Elmer, LPB 2 (1910) 672; --Merr., EPFP 3 (1923) 274, auct. non (A.DC.) Mez, nec Myrsine venosa A.DC.; --Type: Elmer 11445 (GH, lecto; A, BISH, K, L, NY, isolecto), Mindanao: Davao, Mt Apo, Todaya, 2000m, Aug-1909. Endemic to the Philippines. BATAN, LUZON: Mountain Province, Nueva Ecija, MINDORO, PALAWAN: Palawan, PANAY, MINDANAO: Misamis Occidental, Lanao del Sur, Bukidnon, Davao-North Cotabato, Davao del Sur, Davao Oriental, South Cotabato. Submontane and montane mossy forests, up to 2800m.
  14. Myrsine philippinensis A.DC., Prodr. 8 (1844) 94; --Pipoly, Sida 17 (1996) 141; --Rapanea philippinensis (A.DC.) Mez, in Engl., Pflanzenreich 9 (1902) 364; --Merr., EPFP 3 (1923) 273; --Type: Cuming 869 (G, holo; fragm. in A; K 2 sheets, L, iso), Luzon: Albay Prov, 1841. Endemic to the Philippines. LUZON: Batangas, Quezon, Camarines Norte, Albay, POLILLO, MINDORO: Mindoro Oriental (Baco), SIBUYAN (Mt Giting-giting), CUYO ISL GROUP (PAMALICAN), GUIMARAS, LEYTE, SAMAR: Northern Samar, JOLO, MINDANAO: Surigao del Norte. Coastal forests and secondary vegetation, on sands and mangrove swamps, sea level to 305m.  Photos

Acknowledgements

Derek Cabactulan, Andrew Pranada, and Racquel Tan Chua-Barcelo kindly provided some photographs for this webpage. Liezel Molina-Magtoto helped with the identification of several of the Ardisia photos.

Literature

Bentvelzen, PAJ. 1962. Primulaceae. Flora Malesiana series 1, 6: 173-192

DENR Administrative Order. 2017. Updated national list of threatened Philippine plants and their categories. DAO 2017-11

Dubéarnès, A, A Julius & TA Utteridge. 2015. A synopsis of the genus Embelia in Peninsular Malaysia and Singapore. Studies in Malaysian Myrsinaceae III. Kew Bulletin 70: 1-33

Hu Chi Ming, 1999. New synonyms and combinations in Asiatic Ardisia. Blumea 44: 391-406

Molina-Magtoto, L & IE Buot Jr. 2020. Species of Philippine Ardisia Sw. (Primulaceae). Journal on New Biological Reports 9: 289-298

Pipoly III, JJ. 1996. Contributions towards a new flora of the Philippines: I. A synopsis of the genus Myrsine (Myrsinaceae). SIDA 17: 115-162.

Stone, BC. 1994. New and noteworthy Malesian Myrsinaceae, VIII. SIDA 16: 263-272.

Utteridge, TMA & RMK Saunders. 2004. The genus Maesa (Maesaceae) in the Philippines. Botanical Journal of the Linnean Society 145: 17-43.


Pelser, P.B., J.F. Barcelona & D.L. Nickrent (eds.). 2011 onwards. Co's Digital Flora of the Philippines. www.philippineplants.org

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Last updated 21 October 2020