Arthropteridaceae

Edited by Pieter B. Pelser, 1 September 2013

Arthropteris J.Sm. ex Hook.f.

1. Arthropteris beckleri (Hook.) Mett. --Arthropteris obliterata (R.Br.) J.Sm.
2. Arthropteris palisotii (Desv.) Alston. --Arthropteris glabra Copel.
3. Arthropteris repens (Brackenr.) C.Chr. Possibly not distinct from Arthropteris palisotii (Liu et al. 2013). Photos
4. Arthropteris wollastonii (Ridley) Holttum. --Arthropteris orientalis (J. Gmel.) C.Chr.

Aspleniaceae

Asplenium L.

Edited by Julie F. Barcelona, 8 February 2013

1. Asplenium affine Sw., Schrad. J. Bot. 1800 (2) (1801) 56. --Asplenium spathulinum J.Sm. ex Hook. --Asplenium gracilifolium Copel. Photos
2. Asplenium amboinense Willd., Sp. Pl. 5 (1810) 303; 
3. Asplenium apoense Copel. Photos
4. Asplenium callipteris (Fée) E.Fourn. --Asplenium epiphyticum Copel. Photos
5. Asplenium caudatum G Forst., Prodr. (1786) 80. --? Asplenium chamissoanum C.Presl Photos
6. Asplenium cheilosorum Kunze ex Mett., Aspl., 133, n. 104 (1859) t. 5, Fig. 12-13. Photos
7. Asplenium colubrinum Christ 
8. Asplenium crinicaule Hance
9. Asplenium cuneatum Lam. Photos
10. Asplenium cymbifolium Christ, Bull. Boiss. 2, 6 (1906) 999. Photos  
11. Asplenium elmeri Christ, PJS 2 c (1907) Bot. 164 Photos
12. Asplenium excisum C.Presl --Asplenium obscurum Blume. Photos
13. Asplenium exiguum Bedd. Photos
14. Asplenium griseum Copel.
15. Asplenium haenkei Copel. Photos
16. Asplenium insiticium Brackenr.
17. Asplenium lepturus J.Sm. ex C.Presl --Asplenium contiguum Kaulf. 
18. Asplenium lobulatum Mett., Linnaea 36 (1869) 100. Photos
19. Asplenium longissimum Blume. --Asplenium acutiusculum Blume. --?Asplenium anisodontum C.Presl Photos  
20. Asplenium mantalingahanum PM Zamora & Co, Nat. Appl. Sc. Bull. Univ. Philipp. 32 (1980) 46, t. 3.
21. Asplenium nidus L., Sp. Pl. (1753) 1079. Photos
    • var. nidus
    • var. curtisorum (Christ) Holtt., Gard. Bull. Sing. 27 (1974) 148; --Asplenium curtisorum Christ, Ann. Jard. Bot. Buitenzorg II. 4 (1904) 39.
22. Asplenium nigrescens Blume, Enum. Pl. Javae 2 (1828) 180. --Asplenium militare Copel. Photos
23. Asplenium normale D.Don, Prodr. Fl. Nepal. (1825) 7. Photos  
24. Asplenium ofeliae Salgado, Amer. Fern J. 93(4) (2003) 193; --Type: Merrill Phil. Plts. 700. (US, holo; iso: PRC). Luzon: Benguet prov., May 1911. Photos  
25. Asplenium pellucidum Lam., Encycl. 2 (1786) 305. Photos  
26. Asplenium persicifolium J.Sm. ex Mett., Aspl. 97 (1859) no. 32. --Asplenium tripinnatifidum Copel. Photos
27. Asplenium phyllitidis D.Don
    
    • ssp. malesicum Holttum, Gard. Bull. Sing. 27 (1974) 153; --Asplenium nidus L. var. phyllitidis (D. Don) Alderw., Handb. Malayan Ferns Suppl. (1917) 282.  --Probably incl. Asplenium colubrinum. Photos
28. Asplenium polyodon G.Forst., Prodr. (1786) 80; --Asplenium falcatum non Lam. Photos  
29. Asplenium robustum Blume. --Asplenium laserpitiifolium Lam.
30. Asplenium scandens J.Sm. ex Mett. Photos
31. Asplenium schizocarpum Copel.
32. Asplenium scolopendrioides J.Sm. ex Hook., Icon. Pl. (1854) t. 930
33. Asplenium semivarians Viane & Reichst., Pteridol. New Millennium (2003) 78 (-81; fig. 3). Tamil Nadu (India, Indian Subcontinent, Asia-Tropical); Yunnan (China South-Central, China, Asia-Temperate); West-Central Tropical Africa (Africa); Manipur (Assam, Indian Subcontinent, Asia-Tropical); Philippines (Malesia, Asia-Tropical); Sri Lanka (Indian Subcontinent, Asia-Tropical); Uganda (East Tropical Africa, Africa); Zimbabwe (South Tropical Africa, Africa); --Type: Collector(s): Fraser-Jenkins 9063. Locality: Shevaroy Hills, NW of Yercaud, NE of Salem, c. 1550m. 1978-12-13. holotype GENT, isotype Herb. R.Viane 
34. Asplenium setisectum Blume, Enum. Pl. Jav. (1828) 187; 
35. Asplenium stantoni Copel.
36. Asplenium steerei Harrington 
37. Asplenium subnormale Copel. in Perkins, Fragm. Fl. Philip. (1905) 183, t. 4, Fig. B. Photos
38. Asplenium subresectum Copel. 
39. Asplenium tenerum G Forst., Prodr. (1786) 80. Photos  
40. Asplenium tenuifolium Don
41. Asplenium thunbergii Kunze, Linnaea 10 (1836) 517; --Asplenium decorum Kunze. Photos
42. Asplenium unilaterale Lam., Encycl. 2 (1786) 305; --Asplenium filipes Copel., PJS 3 c (1908) Bot. 304. Photos  
43. Asplenium vittiforme Cav., Descr. (1802) 255. Conservation status: Vulnerable (DAO 2017-11). Photos
44. Asplenium vulcanicum Blume, Enum. Pl. Jav. (1828) 176; Possibly conspecific with Asplenium salignum Blume
45. Asplenium yoshinagae Makino var. planicaule (Wall. ex Mett.) C.V.Morton, Contr. US Nat’l. Herb. 38 (1973) 227; --Asplenium planicaule Wall. ex Mett., Abhandl. Senckenb. Naturf. Gessell. 3 (1859) 201, non Lowe (1858).

Athyriaceae

Edited by Julie F. Barcelona, 22 January 2013

Anisocampium C.Presl

Edited by Julie F. Barcelona, 1 September 2013

1. Anisocampium cumingianum C.Presl; --Athyrium cumingianum (C.Presl) Ching. Photos

Athyrium Roth

Last edited by Pieter B. Pelser, 29 July 2018

1. Athyrium anisopterum Christ, Bull. Herb. Boissier 6(12) (1898) 962; --Type: A. Henry 10109 (MO, iso). China, Tibet, Taiwan, India, Nepal, Sri Lanka, Myanmar, Vietnam, Thailand, Sumatra, Borneo, Sulawesi, Java, Peninsular Malaysia, Philippines. LUZON. Photos
2. Athyrium aristulatum Copel., PJS 1 (Suppl. 4) (1906) 253; --Type: Copeland 1880 (MICH, holo). Sulawesi, Sumatra, Philippines. Synonym of Athyrium tozanense (Hayata) Hayata? Photos
3. Athyrium brevipinnulum Copel., PJS 3 C (1908) 291; --Type: Copeland 1939 (MICH, holo). Endemic to the Philippines? Synonym of Athyrium nitidulum (Kunze) Milde? Photos
4. Athyrium drepanopteron (Kunze) Milde, Fil. Eur. (1867) 49; --Polypodium drepanopteron Kunze, Linnaea 23 (1850) 318. Peninsular Malaysia, Sumatra, Philippines. Photos
5. Athyrium elmeri Copel., PJS 3 C (1908) 292. Endemic to the Philippines. Photos
6. Athyrium erythropodum Hayata, Icon. Pl. Formosan. 4 (1914) 233–234, f. 163. Taiwan, Java, Lesser Sunda Islands (Flores), Philippines.
7. Athyrium fauriei (Christ) Makino, Bot. Mag. (Tokyo) 17 (1903) 160; --Nephrodium fauriei Christ, Bull. Boiss. 4 (1896) 671; --Aspidium fauriei (Christ) Christ, Bull. Boiss. 6 (1898) 193. China, Japan, North Korea, South Korea, ?Philippines. Synonym of Athyrium fallaciosum Milde?
8. Athyrium mearnsianum (Copel.) Alderw., Mal. Ferns Suppl. (1917) 279; --Athyrium nigripes var. mearnsianum Copel., PJS 3 C (1908) 291; --Type: Elmer 6543. Sulawesi, Java, Philippines. Synonym of Athyrium nitidulum (Kunze) Milde?
9. Athyrium minutum Copel., Univ. Calif. Publ. Bot. 18 (1942) 221. Sulawesi, New Guinea, Philippines.
   
10. Athyrium oreopteris Copel., PJS 40 (1929) 302; --Type: E.B. Copeland 2305 (MICH, holo; UC, iso). Endemic to the Philippines? Synonym of Athyrium oppositipennum Hayata?
11. Athyrium philippinense Christ, PJS 1 (Suppl. 4) (1906) 254; --Athyrium sarasinorum var. philippinense Christ, Bull. Boiss. 6 (1898) 154. Endemic to the Philippines? Synonym of Athyrium nitidulum (Kunze) Milde?
    
12. Athyrium puncticaule (Blume) T.Moore, Index Fil. (1860) 186. Borneo, India, Japan, Java, Lesser Sunda Isls, Moluccas, Sri Lanka, Sulawesi, Sumatra, Taiwan, Vietnam, Philippines. LUZON. Photos
13. Athyrium ramosii Copel., PJS 38 (1929) 140. Synonym of Diplazium geophilum (Copel.) Alderw.?
14. Athyrium sarasinorum Christ, Verh. Nat. Ges. Basel 11 (1896) 434. Sulawesi, Philippines.

Callipteris Bory = Diplazium

Cornopteris Nakai

Edited by Pieter B. Pelser, 3 March 2012

1. Cornopteris banahaoensis (C.Chr.) K Iwats. & M.G.Price. --Athyrium nudum Copel.
2. Cornopteris decurrenti-alata (Hook.) Nakai. Photos
3. Cornopteris opaca (D. Don) Tagawa. --Athyrium gymnocarpum Copel.

Deparia Hook. & Grev.

Edited by Julie F. Barcelona, 7 September 2013

1. Deparia boryana (Willd.) M.Kato, Bot. Mag. Tokyo 90 (1977) 36; --Dryoathyrium boryanum (Willd.) Ching; --Ctenitis boryana (Willd.) Copel.
2. Deparia confluens (Kunze) M.Kato. Photos
3. Deparia formosana (Rosenst.) Sano, Act. Phytotax. Geobot. 51 (2000) 17; --Diplaziopsis formosana (Rosenst.) M.G.Price, Contrib. Univ. Mich. Herb. 17 (1990) 269; -- Diplazium formosanum Rosenst., Hedw. 56 (1915) 337.
4. Deparia japonica (Thunb.) M.Kato, Bot. Mag. Tokyo 90 (1977) 37; --Lunathyrium japonicum (Thunb.) Kurata; --Athyrium japonicum (Thunb.) Ching. --Athyrium grammitoides (C.Presl) Milde. Photos
5. Deparia lancea (Thunb.) Fraser-Jenkins, New Sp. Syndr. Indian Pteridol. (1997) 101; --Asplenium lanceum Thunb., Fl. Jap. (1784) 333; --Athyrium lanceum (Thunb.) Milde (1870, non Moore 1860); --Copel., Philip. J. Sci. 7C (1912) 55; --Diplazium lanceum (Thunb.) C. Presl, Tent. Pterid. (1836) 113 (non Bory 1833); --Micropodium lanceum (Thunb.) J.Smith; --Triblemma lancea (Thunb.) Ching. --Asplenium subsinuatum Wall. ex Hook. & Grev., Icon. Fil. (1827) t. 27; --Diplazium subsinuatum (Hook. & Grev.) Tagawa, Col. Ill Jap. Pterid. (1959) 203, f. 298; --Price, Kalikasan 1 (1972) 42;. --Athyrium dubium (D.Don) Ohwi (1956), not Ching (1949); --Copel., Fern Fl. Philip. 3 (1961) 412 (non Diplazium dubium Link (1833)); --Scolopendrium dubium D.Don. --Diplazium simplicifolium Kodama. Sri Lanka, India to China, Indochina, Taiwan to Japan, south to the Philippines (Batanes, Luzon and Panay) and Borneo. Photos
6. Deparia petersenii (Kunze) M.Kato. Photos
7. Deparia subfluviatilis (Hayata) M.Kato.

Diplazium Sw.

1. Diplazium accedens Blume Polillo, PNBG 184 Sotalbo (PUH). New record for the Philippines? Most specimens in D. proliferum; the rest in D. vestitum & D. dolichosorum. Photos
2. Diplazium allantoideum M.G.Price.
3. Diplazium atratum Christ; --Athyrium atratum (Christ) Copel.
4. Diplazium bantamense Blume; --Athyrium bantamense (Blume) Milde.
5. Diplazium bolsteri Copel.; --Athyrium bolsteri (Copel.) Copel.
6. Diplazium brachysoroides Copel.; --Athyrium brachysoroides (Copel.) Copel.
7. Diplazium calliphyllum (Copel.) M.G.Price, Contrib. Univ. Mich. Herb. 17 (1990) 270; --Athyrium calliphyllum Copel. Conservation status: Other Threatened Species (DAO 2017-11).
8. Diplazium cordifolium Blume; --Athyrium cordifolium (Blume) Copel. --Athyrium parvum Copel. Photos
9. Diplazium costulisorum (Copel.) C.Chr.; --Athyrium costulisorum Copel. Conservation status: Endangered (DAO 2017-11).
10. Diplazium crenato-serratum (Blume) T.Moore, Index Fil. (T. Moore) 121 (1859) 325; --Athyrium crenato-serratum (Blume) Mildez. Photos
11. Diplazium cultratum C.Presl; --Athyrium cultratum (C.Presl) Milde. Photos
12. Diplazium cumingii (C.Presl) C.Chr.; --Athyrium cumingii (C.Presl) Milde. Photos
13. Diplazium davaoense Copel.; --Athyrium davaoense (Copel.) Copel. Photos
14. Diplazium deltoideum (C.Presl) C.Presl; --Athyrium deltoideum (C.Presl) Milde.
15. Diplazium dilatatum Blume; --Athyrium dilatatum (Blume) Milde. Photos
16. Diplazium doederleinii (Luerssen) Makino. --Athyrium irayense Copel.
17. Diplazium dolichosorum Copel.; --Athyrium dolichosorum (Copel.) Copel.
18. Diplazium egenolfioides M.G.Price, Brit. Fern Gaz. 10 (1973) 258, t. 1 LUZON: Mt. Makiling (several collections from this type locality); Nueva Vizcaya prov., Sta. Fe, mtns N of Imugan, c. 1200m, L.L.Co 1640 (MICH, PNH, PUH). Conservation status: Endangered (DAO 2017-11).
19. Diplazium esculentum (Retz.) Sw.; --Athyrium esculentum (Retz.) Copel. --Asplenium manilense Sprengel.
20. Diplazium forbesii (Baker) C.Chr., Index Filic. (1905) 232. --Diplazium pariens (Copel.) C.Chr.; --Athyrium pariens Copel. Photos
21. Diplazium fraxinifolium C.Presl; --Athyrium fraxinifolium (C.Presl) Milde. Photos
22. Diplazium fructuosum Copel.; --Athyrium fructuosum (Copel.) Copel.
23. Diplazium fuliginosum (Hook.) M.G.Price, Brit. Fern Gaz. 10 (1973) 260; --Asplenium fuliginosum Hook. --Diplazium longissimum (Copel.) C.Chr.; --Athyrium longissimum Copel.
24. Diplazium geophilum (Copel.) Alderw.; --Athyrium geophilum Copel.
25. Diplazium griffithii T.Moore. --Athyrium platyphyllum Copel., 1908,  non Diplazium platyphyllum Christ, 1906. --Diplazium mearnsii Hieron. --Athyrium platyphyllum Copel. Photos
26. Diplazium irigense (Copel.) M.G.Price, Contrib. Univ. Mich. Herb. 17 (1990) 273; --Athyrium irigense Copel.
27. Diplazium lomariaceum (Christ) M.G.Price, Gard. Bull. Sing. 36 (1983) 27. --Diplazium altum (Copel.) C.Chr.; --Athyrium altum Copel. --Diplazium merrillii Copel.; --Athyrium merrillii (Copel.) Copel. Photos
28. Diplazium macrosorum (Copel.) M.G.Price, Contrib. Herb. Univ. Mich. 17 (1990) 273; --Athyrium macrosorum Copel. Conservation status: Other Threatened Species (DAO 2017-11 as Diplazium maximum (D.Don) C.Chr.).
29. Diplazium magnificum (Copel.) M.G.Price, 1982; --Athyrium magnificum Copel.
30. Diplazium melanopodium Fée; --Athyrium melanopodium (Fée) Copel. Photos
31. Diplazium mollifrons (C.Chr.) M.G.Price
32. Diplazium muricatum (Mett.) Alderw.; --Athyrium muricatum (Mett.) Copel.
33. Diplazium oligosorum Copel.; --Athyrium oligosorum (Copel.) Copel.
34. Diplazium pallidum (Blume) Moore; --Athyrium pallidum Blume. Photos
35. Diplazium petiolare C.Presl; --Athyrium petiolare (C.Presl) Milde.
36. Diplazium pinatubicum M.G.Price, Contrib. Herb. Univ. Michigan 17 (1990) 273.
37. Diplazium polypodioides Blume. --Athyrium blumei (Bergsmann) Copel. --Diplazium ophiodontum (Copel.) C.Chr.; --Athyrium ophiodontum Copel. --Diplazium asperum Blume; --Athyrium asperum (Blume) Milde.
38. Diplazium proliferum (Lam.) Thouars; --Callipteris prolifera (Lam.) Bory. --Diplazium accedens Blume.
39. Diplazium propinquum (Copel.) Alderw.; --Athyrium propinquum Copel. Conservation status: Endangered (DAO 2017-11).
40. Diplazium pseudocyatheifolium Rosenst.; --Athyrium pseudocyatheifolium (Rosenst.) Copel. --Athyrium ebenirachis Copel. Photos
41. Diplazium sibuyanense (Copel.) Alderw.; --Athyrium sibuyanense Copel. Conservation status: Other Threatened Species (DAO 2017-11). Photos
42. Diplazium sorzogonense (C.Presl) C.Presl; --Athyrium sorzogonense (C.Presl) Milde. Photo
43. Diplazium symmetricum (Copel.) M.G.Price, Gard. Bull. Sing. 36 (1983) 26.; --Athyrium symmetricum Copel. SAMAR: Mt. Cabayanan, Edano PNH 15148, lectotype (MICH).
44. Diplazium tenuifolium (Copel.) Lellinger, Am. Fern J. 74 (1954) 57; --Athyrium tenuifolium Copel. Formally published new combination?
45. Diplazium tomentosum Blume. --Diplazium vestitum C.Presl; --Athyrium vestitum (C.Presl) Milde. Conservation status: Other Threatened Species (DAO 2017-11 as Diplazium vestitum C.Presl).
46. Diplazium whitfordii Copel.; --Athyrium whitfordii (Copel.) Copel.
47. Diplazium williamsii Copel.; --Athyrium williamsii (Copel.) Copel. --Athyrium banahaoense Copel.; --Diplazium banahaoense (Copel.) C.Chr.
48. Diplazium woodwardioides (C.Presl) Holttum. --Diplazium sylvaticum (Bory) Sw.; --Synopsis Filicum (1867). --Athyrium sylvaticum (Blume) Milde.
49. Diplazium xiphophyllum (Baker) Copel.; --Athyrium xiphophyllum (Baker) Copel.

Dryoathyrium Ching = Deparia

Lunathyrium Koidz. = Deparia

Blechnaceae

Blechnum L.

1. Blechnum egregium Copel. in Perkins, Fragm. Fl. Philip. (1905) 187. Conservation status: Vulnerable (DAO 2017-11). Photos
2. Blechnum finlaysonianum Hook. & Grev., Icon. Fil., t. (1831) 225. Photos
3. Blechnum fraseri (A. Cunningham) Luerss., Flora (1876) 292. Conservation status: Vulnerable (DAO 2017-11). Photos
4. Blechnum melanocaulon (Brack.) T.C.Chambers & P.A.Farrant, Blumea 46(2) (2001) 318. MINDANAO: North Cotabato prov. Photos
5. Blechnum orientale L., Sp. Pl. (1753) 1077. Photos
6. Blechnum patersonii (R.Br.) Mett., Fil. Lips. (1856) 64, t. 4, Fig. 4-10.
7. Blechnum vestitum (Blume) Kuhn, Ann. Lugd.-Bat. 4 (1869) 284. --Blechnum capense (L.) Schlechtr. Photos
8. Blechnum vulcanicum (Blume) Kuhn, Ann. Lugd.-Bat. 4 (1869) 284. Photos

Brainea J.Sm.

1. Brainea insignis (Hook.) J.Sm. MINDORO. Conservation status: Endangered (DAO 2017-11). Photos

Stenochlaena J.Sm.

1. Stenochlaena areolaris (Harrington) Copel.
2. Stenochlaena cumingii Holttum. --Stenochlaena laurifolia C.Presl
3. Stenochlaena milnei Underw. --Stenochlaena juglandifolia C.Presl Photos
4. Stenochlaena palustris (NL Burm.) Bedd., Ferns Brit. Ind. Suppl. (1876) 26. Photos

Woodwardia Sm.

1. Woodwardia orientalis Sw. var. prolifera (Hook. & Arn.) Hook.
2. Woodwardia unigemmata (Makino) Nakai. Photos

Cibotiaceae 

Cibotium Kaulf.

1. Cibotium barometz (L.) J.Sm. Assam to S China, W Malesia. LUZON: Mountain Province (near Bontoc). Known in the Philippines by a single collection, CAHUP 16635 Pancho. Conservation status: Endangered (DAO 2017-11). Photos
2. Cibotium cumingii Kunze. Conservation status: Endangered (DAO 2017-11). Photos

Cyatheaceae

Cyathea Sm.

1. Cyathea acuminata Copel. Conservation status: Endangered (DAO 2017-11).
2. Cyathea apoensis Copel. --Cyathea lobata Copel. Conservation status: Endangered (DAO 2017-11).
3. Cyathea atropurpurea Copel.; --Gymnosphaera atropurpurea (Copel.) Copel. Conservation status: Endangered (DAO 2017-11).
4. Cyathea biliranensis Copel. 
5. Cyathea binuangensis Copel. Conservation status: Endangered (DAO 2017-11).
6. Cyathea callosa Christ. --Cyathea foxworthyi Copel. Conservation status: Vulnerable (DAO 2017-11).
7. Cyathea camiguinensis Copel. 
8. Cyathea caudata (J.Sm.) Copel. --Cyathea arborescens Copel. --Cyathea brevipes Copel. --Cyathea dupaxensis Copel. Conservation status: Vulnerable (DAO 2017-11).
9. Cyathea christii Copel. Conservation status: Endangered (DAO 2017-11).
10. Cyathea cinerea Copel. Conservation status: Vulnerable (DAO 2017-11).
11. Cyathea contaminans (Wall.) Copel., PJS 4 c (1909) Bot. 60; --Cyathea clementis Copel. New Guinea to India. Throughout the Philippines. Usually common in open places and light forests from near sea-level to 1300m. Conservation status: Endangered (DAO 2017-11). Photos
12. Cyathea curranii Copel., Philipp. J. Sci., C 3 (1909) 356. Conservation status: Critically Endangered (DAO 2017-11). Photos
13. Cyathea edanoi Copel. Conservation status: Endangered (DAO 2017-11). Photos  
14. Cyathea elmeri (Copel.) Copel. --Cyathea argyrolepis Copel. --Cyathea dimorphotricha Copel. Conservation status: Vulnerable (DAO 2017-11).
15. Cyathea fenicis Copel. Conservation status: Vulnerable (DAO 2017-11).
16. Cyathea ferruginea Christ. --Cyathea ferrugineoides Copel. Conservation status: Endangered (DAO 2017-11).
17. Cyathea fuliginosa (Christ) Copel. --Cyathea bicolor Copel. --Cyathea dura Copel. --Cyathea lanaensis Christ. --Cyathea lepidigera Copel. --Cyathea mindanaensis (Christ) Copel. --Cyathea squamicostata Copel. --Cyathea sulitii Copel. --Cyathea warihon Copel. Conservation status: Vulnerable (DAO 2017-11).
18. Cyathea halconensis Christ. --Cyathea mearnsii Copel. --Cyathea melanophlebia Copel. Conservation status: Vulnerable (DAO 2017-11).
19. Cyathea heterochlamydea Copel. --Cyathea merrillii Copel. Conservation status: Vulnerable (DAO 2017-11). Photos  
20. Cyathea integra J.Sm. ex Hook. --Cyathea arguta Copel. --Cyathea breviloba Copel. --Cyathea bulusanensis Copel. Photos  
21. Cyathea latebrosa (Wall.) Copel., PJS 4 c (1909) Bot. 52;
22. Cyathea latipinnula Copel. Conservation status: Critically Endangered (DAO 2017-11).
23. Cyathea lepifera (J.Sm. ex Hook.) Copel. --Cyathea calocoma (Christ) Copel. --Cyathea pteridioides Copel. Conservation status: Endangered (DAO 2017-11).
24. Cyathea loheri Christ, Bull. Herb. Boiss. 2, 6 (1906) 1007. --Cyathea campbellii Copel. --Cyathea fructuosa Copel. --Cyathea indusiosa Copel. --Cyathea matthewii (Christ) Domin. --Cyathea mitrata Copel. Photos
25. Cyathea lurida (Blume) Copel. --Gymnosphaera melanorhachis (Copel.) Copel.
26. Cyathea masapilidensis Copel. Conservation status: Endangered (DAO 2017-11).
27. Cyathea microchlamys Holttum. Conservation status: Critically Endangered (DAO 2017-11).
28. Cyathea negrosiana Christ. Conservation status: Vulnerable (DAO 2017-11).
29. Cyathea obliqua Copel.; --Gymnosphaera obliqua (Copel.) Copel. Conservation status: Critically Endangered (DAO 2017-11). Photos
30. Cyathea philippinensis Baker, Ann. Bot. 5 (1891) 186; --Copel., FFP 2 (1960) 205; --Holttum,  Fl. Males. ser. 2, 1 (1963) 149. --Type: From a cultivated plant originating from the Philippines, ex Hort. Veitch., Feb. 1878), cult. Kew (K). --Cyathea bicolana Copel. in Elmer, LPB 9 (1920) 3108; --Copel., FFP 2 (1960) 204; --Type: Elmer 16528  (L, U, US, isotypes). Luzon: Sorsogon prov., Mt Bulusan, July 1916. --Cyathea bontocensis Copel., PJS 46 (1931) 209; FFP 2 (1960) 205; --Type: Vanoverbergh 813. Luzon:  Mountain Province. --Cyathea heteroloba Copel., PJS 38 (1929) 134; FFP 2 (1960) 204; --Type: BS 45633 Ramos & Edaño (NY, isotype). Luzon: Quirino prov., Mt Alzapan, altitude 1500m, May-June 1925. --Cyathea ramosiana Alderw., Bull. Jard. Bot. Buitenz. iii, 2 (1920) 137; --Copel., FFP 2 (1960) 206. --Type: BS 23411 Ramos (L, US, isotypes). Luzon: Sorsogon prov., July-Aug. 1915. LUZON: Mountain Province, Quirino (Mt Alzapan), Aurora (Mt Camatis), Quezon (Mt Banahaw), Camarines Sur (Mt Isarog, apparently common), Sorsogon (Mt Bulusan), MINDORO: Mindoro Oriental (Mt Halcon). Montane forests, alt. 800-1500m. Conservation status: Vulnerable (DAO 2017-11). Photos  
31. Cyathea polypoda Baker in Stapf, Trans. Linn. Soc. Lond. Bot. 4 (1894) 250. --Gymnosphaera dinagatensis (Copel.) Copel. --Gymnosphaera glabra sensu Copel. Fern Fl. Philip. 2 (1960) 235, non Blume.
32. Cyathea recommutata Copel., PJS 4 (1909) 36; --Alsophila commutata Mett., Ann. Mus. Bot. Lugduno-Batavi 1 (1863) 53; --Type: from Malaysia. Borneo, Malay Peninsula, Sumatra, Philippines. MINDANAO: Bukidnon (Mt Kiamo, Mt Limbawon). Montane forests c. 1600-1700m. Photos
33. Cyathea robinsonii Copel. --Cyathea pseudoalbizzia Copel. Conservation status: Vulnerable (DAO 2017-11).
34. Cyathea rufopannosa Christ. Conservation status: Endangered (DAO 2017-11).
35. Cyathea setulosa Copel. Conservation status: Vulnerable (DAO 2017-11).
36. Cyathea sibuyanensis Copel. Conservation status: Critically Endangered (DAO 2017-11). SIBUYAN. Photos
37. Cyathea squamulata (Blume) Copel., PJS 4 c (1909) Bot. 37; --Gymnosphaera squamulata Blume
38. Cyathea suluensis Baker. --Cyathea sessilipinnula Copel.
39. Cyathea trichophora Copel., PJS 6 c (1911) Bot. 363; --Gymnosphaera trichophora (Copel.) Copel.
40. Cyathea tripinnata Copel., PJS 1 (1906) Suppl. 251. --Cyathea densisora Alderw. --Cyathea leucostegia Copel. --Cyathea leytensis Copel.
41. Cyathea zamboangana Copel., PJS 30 (1926) 325. --Cyathea urdanetensis Copel., PJS 38 (1929) 132. Endemic to the Philippines. MINDANAO, NEGROS, ?SAMAR. Conservation status: Endangered (DAO 2017-11). Photos

Gymnosphaera Blume = Cyathea

Cystopteridaceae

Currania Copeland = Gymnocarpium

Cystopteris Bernhardi

1. Cystopteris tenuisecta (Blume) Mett., Ann. Lugd. Bat. 1 (1864) 241; --Acystopteris tenuisecta (Blume) Tagawa. --Ctenitis tenerifrons (Christ) Copel.

Gymnocarpium Newman

1. Gymnocarpium oyamense (Baker) Ching. --Currania gracilipes Copel. Conservation status: Vulnerable (DAO 2017-11).

Davalliaceae

Araiostegia Copel. = Davallia

Davallia J.Sm.

1. Davallia angustata Hook. & Grev.; --Humata angustata (Wall.) J.Sm. --Humata microsora Alderw.
2. Davallia brevipes Copel.; --Hovenkamp & de Joncheere, Blumea 33 (1988) 408. Philippines, Sulawesi, New Guinea.
3. Davallia denticulata (NL Burm.) Mett., ex Kuhn, Fil. Deck., (1867) 27. Photos
   
   • var. denticulata
   • var. elata (G Forst.) Kuhn. Conservation status: Other Threatened Species (DAO 2017-11).
4. Davallia divaricata Blume var. divaricata.
5. Davallia embolostegia Copel., PJS 1 (1906) Suppl. 147, t. 3. Conservation status: Other Threatened Species (DAO 2017-11). Photos
6. Davallia falcinella (J.Sm.) C.Presl; --Trogostolon falcinellus (C.Presl) Copel.
7. Davallia heterophylla Sm.; --Humata heterophylla (J.Sm.) Desv., Prodr. (1825) 323. --Humata ophioglossa Cav. Photos
8. Davallia hymenophylloides (Blume) Kuhn; --Araiostegia hymenophylloides (Blume) Copel., PJS 34 (1927) 241. Photos
9. Davallia pectinata Sm.; --Humata pectinata (J.Sm.) Desv. Photos
10. Davallia repens (L.f.) Kuhn.; Humata repens (L.f.) Diels in Engl. & Prant, Nat. Pflanzenfam. 14 (1899) 209. --Humata intermedia C.Chr. --Humata obtusata Alderw. --Humata pusilloides Copel. --Humata trifoliata Cav. --Humata vestita (Blume) T.Moore, Index (1857) 92. Photos
11. Davallia sessilifolia Blume
12. Davallia solida (Forst.) Sw., Schrad. J. Bot. 1800 (1801) 87. --Davallia elmeri Copel. --Davallia robinsonii Copel. Malesia to Polynesia. Throughout the Philippines. Epiphytic or epilithic, sea-level to c. 600m. Photos
    
    • var. solida. Conservation status: Other Threatened Species (DAO 2017-11).
13. Davallia trichomanoides Blume. Photos
    • var. trichomanoides
    • var. lorrainii (Hance) Holttum; --Davallia lorrainii Hance, Ann. Sci. Nat. 5, 5 (1866) 254. Conservation status: Other Threatened Species (DAO 2017-11 as Davallia lorrainii Hance).
14. Davallia wagneriana Copel. Photos

Davallodes (Copel.) Copel.

1. Davallodes hirsutum (C.Presl) Copel. --Davallodes grammatosorum Copel. --Davallodes gymnocarpum Copel. --Davallodes laxum Copel. --Davallodes congestum Copel. Photos

Humata Cav. = Davallia

Trogostolon Copel. = Davallia falcinella

Dennstaedtiaceae 

including Hypolepidaceae, Monachosoraceae, and Pteridiaceae  

Dennstaedtia Bernh.

1. Dennstaedtia articulata Copel. Conservation status: Endangered (DAO 2017-11).
2. Dennstaedtia dennstaedtioides (Copel.) Copel.
3. Dennstaedtia elmeri Copel.
4. Dennstaedtia fusca Copel. Conservation status: Endangered (DAO 2017-11).
5. Dennstaedtia glabrata (Cesati) C.Chr.
6. Dennstaedtia hooveri Christ. Photos  
7. Dennstaedtia macgregori Copel.
8. Dennstaedtia merrillii Copel.
9. Dennstaedtia philippinensis Copel.
10. Dennstaedtia resinifera (Blume) Mett. ex Kuhn. --Dennstaedtia cuneata (J.Sm. ex Hook.) T.Moore.
11. Dennstaedtia rufidula C.Chr. in C.Chr. & Holltum, Gard. Bull. Sing. 7 (1934) 226, t. 51.
12. Dennstaedtia scabra (Wall.) T.Moore. Photos
13. Dennstaedtia scandens (Blume) T.Moore.
14. Dennstaedtia smithii (Hook.) T.Moore.
15. Dennstaedtia williamsii Copel.

Histiopteris (Agardh) J.Sm.

1. Histiopteris incisa (Thunb.) J.Sm., Hist. Fil. (1875) 295. --Histiopteris montana Copel. Photos  

Hypolepis Bernh.

1. Hypolepis alpina (Blume) Hook., Sp. Fil. 2 (1852) 63.
2. Hypolepis bamleriana Rosenst., Feddes Repert. 10 (1912) 325.
3. Hypolepis glandulifera Brownsey & Chinnock, J. Adelaide Bot. Gard. 10 (1987) 16.
4. Hypolepis malesiana Brownsey.
5. Hypolepis pallida (Blume) Hook.?, Sp. Fil. 2 (1852) 64; Doubtfully occurring in the Philippines (Negros: Amlan, M.G.Price 2479).
6. Hypolepis polypodioides (Blume) Hook.
7. Hypolepis punctata (Thunb.) Mett. ex Kuhn Doubtfully occurring in the Philippines (Luzon: Ilocos Norte, Mt. Bubungkayo, Iwatsuki et al. P375 (L).
8. Hypolepis tenuifolia (Forst. f.) Bernh. ex C.Presl 

Microlepia C.Presl

1. Microlepia herbacea Ching & C.Chr. apud Tard. ex C.Chr. --Microlepia trapeziformis (Roxb.) Kuhn. Photos
2. Microlepia platyphylla (D.Don) J.Sm. Photos
3. Microlepia protracta Copel. Conservation status: Endangered (DAO 2017-11).
4. Microlepia speluncae (L.) T.Moore. --Microlepia sablanensis Christ. --? Microlepia manilensis C.Presl Photos  
5. Microlepia strigosa (Thunb.) C.Presl Photos
6. Microlepia todayensis Christ
7. Microlepia trichotricha J.Sm.

Monachosorum Kunze

1. Monachosorum henryi Christ Luzon: Cordillera Central Mtn. Range, Mt. Data, alt. 2220 m, Co 4086 (L, PNH, PUH). New to the Philippines. Previously known from Taiwan and Mainland China (Guangdong, Guangxi, Guizhou and NE & W Yunnan). Photos  
2. Monachosorum subdigitatum (Blume) Kuhn, Chaetopt. (1882) 345; 

Paesia A.St.-Hil.

1. Paesia elmeri Copel. Photos
2. Paesia luzonica Christ, Philipp. J. Sci., C 3 (1908) 275. Photos
3. Paesia radula (Baker) C.Chr.

Pteridium Gled. ex Scop.

1. Pteridium aquilinum (L.) Kuhn Photos
   • var. wightianum (J Agardh) Tryon. 
2. Pteridium caudatum (L.) Maxon. --Pteridium aquilinum (L.) Kuhn var. yarrabense Domin

Dicksoniaceae

Calochlaena (Maxon) M.D.Turner & R.A.White

1. Calochlaena javanica Culcita javanica (Blume) Maxon. --Culcita copelandii (Christ) Maxon. --Culcita straminea (Labill.) Maxon. Photos  

Dicksonia L’Her.

1. Dicksonia amorosoana Lehnert & Coritico, Blumea 63 (2018) 269; --Type: F.P. Coritico FPC 016 (PNH, holo; BONN, CMUH iso), Mindanao: North Cotabato, Mt Apo, Kidapawan, Mandarangan, trail to Lake Venado, 2106m, 5-May-2012. Endemic to the Philippines. MINDANAO: North Cotabato.
   
2. Dicksonia mollis Holttum, Kew Bull. 16 (1962) 64; --Type: Elmer 9874 (K, holo; BM, L, MICH, MO, NY, P, U, US, iso), Negros: Negros Oriental, Dumaguete, Cuernos Mts,
   Apr-1908. Borneo, Philippines. NEGROS. Conservation status: Endangered (DAO 2017-11). Photos  

Diplaziopsidaceae

Edited by Julie F. Barcelona, 22 January 2013

Diplaziopsis C.Chr.

1. Diplaziopsis javanica (Blume) C.Chr., Index Filic. (1905) 227; --Diplazium javanicum (Blume) Makino, Phan. Pter. Jap. Ic. I11. 1(1901) pl. 46. Photos

Dipteridaceae

Cheiropleuria C.Presl

1. Cheiropleuria bicuspis (Blume) C.Presl Photos  

Dipteris Reinw.

1. Dipteris conjugata Reinw. Photos  
2. Dipteris lobbiana (Blume) Moore. Conservation status: Critically Endangered (DAO 2017-11). Photos

Dryopteridaceae 

Acrophorus C.Presl = Dryopteris

Acystopteris Nakai = Cystopteris

Arachniodes Blume

1. Arachniodes amabilis (Blume) Tindale; --Rumohra amabilis (Blume) Ching. Photos
2. Arachniodes aristata (Forster) Tindale; --Rumohra aristata (Forst.) Ching.
3. Arachniodes coniifolia (Wall. ex T.Moore) Ching. --Rumohra wallichii Ching.
4. Arachniodes hasseltii (Blume) Ching; --Rumohra hasseltii (Blume) Ching.
5. Arachniodes tripinnata (Goldm.) Sledge. --Rumohra carvifolia (Kunze) Ching.

 Arthrobotrya J.Sm. = Teratophyllum

Bolbitis Schott

1. Bolbitis angustipinna (Hayata) H.Ito.
2. Bolbitis appendiculata (Willd.) K Iwats. ssp. appendiculata; --Egenolfia appendiculata (Willd.) J.Sm.
3. Bolbitis heteroclita (C.Presl) Ching. --Bolbitis cuspidata (C.Presl) Ching. --Bolbitis pseudoscalpturata (Copel.) Ching. --Bolbitis foxworthyi (Copel.) Ching. --Bolbitis tenuissima (Copel.) Copel. --Bolbitis membranacea (Copel.) C.Chr. --Bolbitis edanyoi Copel. --Edanyoa difformis Copel. Photos
4. Bolbitis quoyana (Gaudich.) Ching. --Bolbitis enormis (Copel.) C.Chr. --Stenosemia dimorpha Copel.
5. Bolbitis repanda (Blume) Schott. Photos
6. Bolbitis rhizophylla (Kaulf.) Hennipman. --Egenolfia intermedia (J.Sm.) Fée. --Egenolfia rhizophylla (Kaulf.) Fée. --Egenolfia fluviatilis Copel. Photos
7. Bolbitis scalpturata (Fée) Ching. Photos
8. Bolbitis sinuata (C.Presl) Hennipman. --Bolbitis diversifolia (Blume) Schott. --Bolbitis subsimplex (Fée) Ching. --Bolbitis hydrophylla (Copel.) Ching. Photos

Hybrids

1. Bolbitis X arguta (Fée) Ching.  Hennipman doubts whether this fern warrants a separate treatment as a hybrid; it may be close to Edanyoa difformis (= B. X heteroclita).
2. Bolbitis heteroclita X rhizophylla Hennipman, Leid. Bot. Ser. 2 (1977) 291, Fig. 831-o.
3. Bolbitis X sinuosa (Fée) Copel., two nothomorphs distinguished: nm. sinuosa nm. foxii (Copel.) Hennipman and Bolbitis X foxii Copel. Photos

Dubious species

1. Leptochilus stolonifer Christ, Bull. Herb. Boiss. Ii, 6 (1906) 1004. Luzon: Rizal prov., Angilog (type locality). Possibly synonymous with B. heteroclita but the venation pattern as given by Christ is deviating. Type not traced (Hennipman, Fl. Males. Ser ii 1 (1978) 330).

Excluded taxa

1. Bolbitis presliana (Fée) Ching  Included in Copeland’s treatment in Fern Fl. Philipp. 2 (1960) 260. This taxon does not occur in the Philippines nor anywhere in Malesia.

Cyrtomium C.Presl

1. Cyrtomium fortunei J.Sm. forma latipinna Ching. Photos

 Diacalpe Blume = Dryopteris

Dryopsis Holttum & P.J.Edwards = Dryopteris

Dryopteris Adans.

1. Dryopteris cacaina Tagawa.
2. Dryopteris cochleata (Don) C.Chr.
3. Dryopteris chrysocoma C.Chr. Conservation status: Endangered (DAO 2017-11).
4. Dryopteris formosana (Christ) C.Chr. Photos
5. Dryopteris hendersonii (Bedd.) C.Chr., Index Filic. (1905) 270; --Nothoperanema hendersonii (Bedd.) Ching. --Ctenitis mearnsii Copel. As Ct. mearnsii: LUZON: Benguet (Pauai; Mt Pulag). Terrestrial in mossy forest, altitude 2100-2750m. Photos
6. Dryopteris hirtipes (Blume) Kuntze. Photos
7. Dryopteris manipurensis (Bedd.) C.Chr., Index Filic. (1905) 276; --Dryopsis manipurensis (Bedd.) Holttum & P.J.Edwards, Kew Bull. 41 (1986) 200. --Ctenitis copelandii (Christ) Copel. --Ctenitis manipurensis (Bedd.) Ching.
8. Dryopteris nodosa (C. Presl) Li Bing Zhang; --Acrophorus nodosus C.Presl --Acrophorus stipellatus (Wall.) T.Moore. Photos
9. Dryopteris permagna M.G.Price. Conservation status: Endangered (DAO 2017-11).
10. Dryopteris polita Rosenst. Conservation status: Vulnerable (DAO 2017-11).
11. Dryopteris pseudocaenopteris (Kunze) Li Bing Zhang, Taxon 61(6) (2012) 1209; --Peranema aspidioides (Blume) Mett.; --Diacalpe aspidioides Blume. Photos
12. Dryopteris pulvinulifera (Bedd.) Kuntze.
13. Dryopteris purpurascens (Blume) Christ. Photos
14. Dryopteris sparsa (Don) Kuntze. Photos
15. Dryopteris subarborea (Baker) C.Chr.
16. Dryopteris uropinna M.G.Price. --Dryopteris erythrosora sensu Copel. (1960), non (Eaton) Kuntze. Photos
17. Dryopteris varia (L.) Kuntze.
18. Dryopteris wallichiana (Sprengel) Alston & Bonner. --Dryopteris paleacea (Sw.) C.Chr., nom. illeg. Photos
19. Dryopteris zhuweimingii Li Bing Zhang, Taxon 61(6) (2012) 1211; --Peranema cyatheoides D.Don var. luzonicum (Copel.) Ching & SH Wu, Acta Phytotax. Sin. 21 (1983) 373; --Peranema luzonica Copel. Conservation status: Critically Endangered (DAO 2017-11). Photos

 Edanyoa Copel. = Bolbitis heteroclita

 Egenolfia Schott = Bolbitis

Elaphoglossum Schott ex J.Sm.

1. Elaphoglossum angulatum (Blume) T.Moore.
2. Elaphoglossum apoense Holttum.
3. Elaphoglossum blumeanum (Fée) J.Sm. Photos
   • var. blumeanum. --Elaphoglossum copelandii Christ
   • var. philippinense Christ ex Holttum., Flora Malesiana Series II, 1 (1978) 311.
4. Elaphoglossum calanasanicum Holttum.
5. Elaphoglossum callifolium (Blume) T.Moore. Photos
6. Elaphoglossum luzonicum Copel. --Elaphoglossum elmeri Copel. Photos
7. Elaphoglossum melanostictum (Blume) T.Moore. --Elaphoglossum basilanicum Copel.
8. Elaphoglossum negrosensis Holttum. --Elaphoglossum angulatum sensu Copel. (1960), pro parte, non (Blume) T.Moore.
9. Elaphoglossum ophioglossoides (Goldm.) Holttum. --Elaphoglossum merrillii Christ. Photos

Hemigramma Christ = Tectaria

Lastreopsis Ching

1. Lastreopsis tenera (R.Br.) Tindale; --Ctenitis recedens (J.Sm.) Copel.

Lomagramma J.Sm.

1. Lomagramma brooksii Copel.
2. Lomagramma copelandii Holttum.
3. Lomagramma merrillii Holttum. Photos
4. Lomagramma pteroides J.Sm. --Lomagramma subcoriacea Copel. --Lomagramma pedicellata Copel. --Lomagramma cordata Copel. Photos

Nothoperanema (Tagawa) Ching = Dryopteris

Peranema D.Don = Dryopteris

Phanerophlebia C.Presl = Cyrtomium

Polystichum Roth

1. Polystichum biaristatum (Blume) T.Moore. --Polystichum aquilinum Copel.
2. Polystichum copelandii Christ.
3. Polystichum deltodon (Baker) Diels.
4. Polystichum doianum Tagawa. --Polystichum pulogense Copel.
5. Polystichum elmeri Copel. Photos
6. Polystichum falcatum (L.f.) Diels, Nat. Pflanzenfam. 1 (1899) 194; --Phanerophlebia falcata (L. f.) Copel. Philippine record based on a Loher collection from Mt. Data [Copeland, Fern Fl. Philipp. 2 (1960) 248] at the turn of the 1900s. Photos
   • var. caryotideum (Wall. ex Hook. & Grev.) Matsum., Index Pl. Jap. 1 (1904) 342; --Phanerophlebia caryotidea (Wall. ex Hook. & Grev.) Copel. Philippine record based on a sterile specimen by Mearns from Mt. Sto. Tomas [Copeland, Fern Fl. Philipp. 2 (1960) 248].
7. Polystichum fuscum Copel.
8. Polystichum horizontale C.Presl Photos
9. Polystichum nepalense (Sprengel) C.Chr.
10. Polystichum nudum Copel. Conservation status: Endangered (DAO 2017-11). Photos
11. Polystichum obliquum (D.Don) T.Moore.
12. Polystichum obtusum J.Sm. ex C.Presl --Polystichum edanoi Copel.

Rumohra sensu Copel. (1960) non Raddi = Arachniodes

Stenolepia Alderw.

1. Stenolepia tristis (Blume) Alderw., Bull. Dept. Agric. Indes Neerl. 27 (1909) 46 t. 7; --Aspidium triste Blume, Enum. Pl. Javae 2 (1828) 169; --Type from Java. Widespread throughout the Malesian region. MINDANAO: Bukidnon, North Cotabato. 2300-2900m. Conservation status: Other Threatened Species (DAO 2017-11).

Teratophyllum Mett.

1. Teratophyllum aculeatum (Blume) Mett. ex Kuhn. Photos
   • var. aculeatum. 
2. Teratophyllum arthropteroides (Christ) Holttum.
3. Teratophyllum articulatum (J.Sm. ex Fée) Mett. ex Kuhn; --Arthrobotrya articulata J.Sm.
4. Teratophyllum koordersii Holttum. Photos
5. Teratophyllum leptocarpum (Fée) Holttum. Photos
6. Teratophyllum luzonicum Holttum. Photos

Equisetaceae

Equisetum L.

1. Equisetum ramosissimum Desf. ssp. debile (Roxb. ex Vauch.) Hauke, Amer. Fern J. 52 (1962) 33. Photos.  

Gleicheniaceae

Edited by Pieter B. Pelser, 10 June 2012, following Christenhusz et al. 2011

Dicranopteris Bernh.

1. Dicranopteris curranii Copel., PJS 81 (1952) 4; --Copel., FFP 1 (1958) 46; Sumatra, Peninsular Malaysia, W Java, Lesser Sunda Isls (Flores), Borneo, Sulawesi, Philippines. LUZON (Laguna), NEGROS, MINDANAO. Photos  
2. Dicranopteris linearis (Burm.) Underw., Bull. Torrey Bot. Club 34 (1907) 250; --Copel., FFP 1 (1958) 45. Photos  
   • var. altissima Holttum. 
   • var. latiloba Holttum.
   • var. rigida (Blume) Holttum. Dicranopteris crassifolia (C.Presl) Ching, Sunyatsenia 5 (1940) 275; --Copel., FFP 1 (1958) 46.
   • var. subspeciosa Holttum.

Diplopterygium (Diels) Nakai

Photos

1. Diplopterygium elmeri (Copel.) Nakai, Bull. Natl. Sci. Mus. 29 (1950) 49; --Gleichenia elmeri Copel. in Elmer, LPB 3 (1910) 799; --Hicriopteris elmeri (Copel.) Copel., FFP 1 (1958) 43. Photos
2. Diplopterygium laevissimum (H.Christ) Nakai, Bull. Natl. Sci. Mus. 29 (1950) 52; --Gleichenia laevissima Christ, Bull. Ac. Inst. Geogr. Bot. 3, 11 (1902) 268; --Hicriopteris laevissima (Christ) Ching, Sunyatesenia 5 (1940) 280; --Copel., FFP 1 (1958) 44.
3. Diplopterygium longissimum (Blume) Nakai, Bull. Natl. Sci. Mus. 29 (1950) 53; --Gleichenia longissima Blume, Enum. Pl. Jav. (1828) 250. --Hicriopteris glauca (Thunb.) Ching, Sunyatsenia 5 (1940) 279; --Copel., FFP 1 (1958) 43 S China to Indochina through Malesia to Melanesia (Fiji, Tahiti). In Malesia not collected in the Lesser Sunda Isls E of Flores, Sulawesi, the Moluccas and New Guinea. Throughout the Philippines, usually at 800-1500m alt. in exposed places. Photos 

Gleichenia Sm.

1. Gleichenia dicarpa R.Br., Prodr. Fl. Nov. Holl. (1810) 161; --Copel., FFP 1 (1958) 38. Australia, New Caledonia, Malesia: New Guinea and Philippines. MINDANAO (Agusan del Norte, Mt Urdaneta; Davao, Mt Apo). Photos  
2. Gleichenia hirta Blume, Enum. Pl. Jav. (1828) 250; --Copel., FFP 1 (1958) 40. Sticherus hirtus (Blume) Ching, Sunyatsenia 5 (1940) 203 (err. ‘hirsutus’) Moluccas and the Philippines. LUZON, MINDORO, MINDANAO. Photos  
3. Gleichenia microphylla R.Br., Prodromus Florae Novae Hollandiae (1810). Photos
4. Gleichenia paleacea (Copel.) Holttum, Reinwardtia 4 (1957) 265; --Hicriopteris paleacea Copel., PJS 81 (1952) 3; FFP 1 (1958) 44. 
5. Gleichenia peltophora Copel., PJS 40 (1929) 292, t. 1; --Copel., FFP 1 (1958) 39. New Guinea, S C Sulawesi, Borneo and Philippines. MINDANAO: South Cotabato (Mt Matutum), alt. 1600m.
6. Gleichenia sordida Copel. in Elmer, LPB 3 (1910) 798; --Hicriopteris sordida (Copel.) Copel., FFP 1 (1958) 44.
7. Gleichenia vulcanica Blume, Enum. Pl. Jav. (1828) 251; Sumatra, Peninsular Malaysia, W java, Borneo, Philippines, Sulawesi and New Guinea. MINDORO: Mindoro Oriental (Mt Halcon), PALAWAN (Mt Pulgar and Mt Mantalingahan, Co 1773), MINDANAO. Photos

Sticherus C.Presl

Edited by Julie F. Barcelona, 23 July 2013

Photos

1. Sticherus hispidus (Mett. ex Kuhn) Copel., Gen. Fil. (1947) 27; FFP 1 (1958) 41; --Gleichenia hispida Mett. ex Kuhn, Verh. K.K. Zool.-Bot. Ges. Wien 25 (1875) 600. --Sticherus koordersii (Christ) Copel., FFP 1 (1958) 40. --Sticherus pinnatus Copel. Sumatra, Java, Bali, N Sulawesi, Ternate and Philippines. LUZON: Benguet, Albay (Mt Mayon), NEGROS.
2. Sticherus loheri (Christ) Copel., FFP 1 (1958) 42; --Gleichenia loheri Christ, Bull. Herb. Boiss. 2, 6 (1906) 1009. --Sticherus perpaleaceus Copel., PJS 81 (1952) 3; FFP 1 (1958) 42. Photos
   • var. loheri LUZON: Benguet (Mt Pulog), Rizal, Quezon (Mt Banahaw).
   • var. major Holttum, Reinwardtia 4 (1959) 272; NEGROS: Negros Occidental (Mt Kanlaon). 
3. Sticherus milnei (Baker) Ching, Sunyatsenia 5 (1940) 284; --Gleichenia milnei Baker, Syn. Fil. (1874) 449; Sulawesi, Moluccas, New Guinea, Admiralty Isls, Solomons, New Hebrides. LUZON. Photos
4. Sticherus truncatus (Willd.) Nakai, Bull. Natl. Sci. Mus. 29 (1950) 20; --Gleichenia truncata (Willd.) Sprengel, Syst. Veg. ed. 16, 4 (1827) 25. Photos
• var. truncata. --Sticherus laevigatus (Willd.) C.Presl, Tent. (1836) 52; --Copel., FFP 1 (1958) 41; Throughout Malesia, apparently most common in the west. 

Hymenophyllaceae

Edited by Pieter B. Pelser, 10 June 2012, following Ebihara et al. 2006.

Abrodictyum C.Presl

Callistopteris C.Presl

Cephalomanes C.Presl

1. Cephalomanes atrovirens C.Presl, Hymen. (1843) 17, pl. 5; --Copel., FFP 1 (1958) 77; --Trichomanes atrovirens (C.Presl) Kunze, Bot. Zeit. 6 (1847) 371; --Copel., PJS 51 (1933) 251, pl. 52, Fig. 3; pl. 55, Fig. 2. Photos.  
2. Cephalomanes crassum (Copel.) M.G.Price; --Trichomanes crassum Copel.
3. Cephalomanes javanicum (Blume) Bosch var. asplenioides (C.Presl) K Iwats. --Cephalomanes oblongifolium C.Presl, Epim. Bot. (1849) 19, pl. 10; --Copel., FFP 1 (1923) 78. --Trichomanes asplenioides C.Presl --Trichomanes laciniatum Roxb. Photos

Crepidomanes (C.Presl) C.Presl

1. Crepidomanes bipunctatum (Poir.) Copel., PJS 67 (1938) 59; FFP 1 (1958) 72; --Trichomanes bipunctatum Poir. in Lam., Encycl. 8 (1808) 69; --Copel., PJS 51 (1933) 177, pl. 18, Fig. 3, 1-4. Photos
2. Crepidomanes brevipes (C.Presl) Copel., PJS 67 (1938) 60; FFP 1 (1958) 71; --Trichomanes brevipes (C.Presl) Baker, Syn. Fil. (1867) 84; --Copel., PJS 51 (1933) 182, pl. 20; --Didymoglossum brevipes C.Presl Photos
3. Crepidomanes christii (Copel.) Copel., PJS 67 (1938) 60; FFP 1 (1958) 73; --Trichomanes christii Copel., PJS 1 (1906) Suppl. 251; PJS 51 (1933) 185, pl. 21.
4. Crepidomanes grande (Copel.) Ebihara & K. Iwats., Blumea 51 (2006) 239; --Cephalomanes grande (Copel.) K Iwats.; --Trichomanes grande Copel., PJS 6 c (1911) 70; PJS 51 (1933) 224, pl. 40, Fig. 1-4; --Nesopteris grandis (Copel.) Copel., PJS 67 (1938) 66; FFP 1 (1958) 76. Photos
5. Crepidomanes humile (Forst.) Bosch; --Trichomanes humile Forst., Prodr. (1786) 12; --Copel., PJS 51 (1933) 164, pl. 12; --Crepidopteris humilis (Forst.) Copel., PJS 67 (1938) 58; FFP 1 (1958) 70. --Trichomanes gracillimum Copel., PJS 51 (1933) 168, pl. 13; --Crepidopteris gracillima (Copel.) Copel., PJS 67 (1938) 58; FFP 1 (1958) 70. Photos
6. Crepidomanes minutum (Blume) K Iwats.; --Trichomanes minutum Blume, Enum. Pl. Jav. (1828) 223; --Trichomanes minutum Blume forma alagensis (C.Chr.) Copel; --Gonocormus minutus (Blume) Bosch, Hymen. Javan. (1861) 7, pl. 3; --Copel., FFP 1 (1958) 68. --Trichomanes alagensis Christ, PJS 3 c (1908) Bot. 270; --Copel., PJS 51 (1933) 152, pl. 6, Fig. 4-7; --Gonocormus alagensis (Christ) Copel., PJS 67 (1938) 57. Photos
7. Crepidomanes rupicolum (Raciborski) Copel., PJS 67 (1938) 59; FFP 1 (1958) 73; --Trichomanes rupicolum Raciborski, Pterid. Buitenz. (1898) 24. 
8. Crepidomanes schmidtianum (Zenker ex Taschner) K. Iwats., J. Fac. Sci. Univ. Tokyo, Sect. 3, Bot. 13(5) (1985) 526
   
   • var. latifrons (Bosch) K. Iwats.; --Trichomanes latifrons Bosch; --Vandenboschia latifrons (Bosch) Copel.
9. Crepidomanes thysanostomum (Makino) Ebihara & K.Iwats., Blumea 51 (2006) 240; --Cephalomanes thysanostomum (Makino) K Iwats.; --Trichomanes thysanostomum Makino, Bot. mag. Tokyo 12 (1898) 193; --Nesopteris thysanostoma (Makino) Copel., PJS 67 (1938) 66. 

Crepidopteris Copel. = Crepidomanes

Didymoglossum Desv.

1. Didymoglossum mindorense (H.Christ) K.Iwats., Blumea 51 (2006) 236; -- Trichomanes mindorense Christ, PJS 3 c (1908) Bot. 270; --Copel., PJS 51 (1933) 209, pl. 34,  Fig. 1, 2; --Microgonium mindorense (Christ) Copel., PJS 67 (1938) 61; PJS 51 (1933) 164, pl. 12. --Trichomanes craspedoneurum Copel., PJS 7 c (1912) Bot. 58; PJS 51 (1933) 208, pl. 33, Fig. 5-7; --Microgonium craspedoneurum (Copel.) Copel., PJS 67 (1938) 61; FFP 1 (1958) 74.
2. Didymoglossum motleyi (Bosch) Ebihara & K. Iwats., Blumea 51 (2006) 236; --Trichomanes motleyi Bosch, Ned. Kruid. Arch. 5 (1861) 145; --Copel., PJS 51 (1933) 201, pl. xx, Fig. 1-4; --Microgonium motleyi Bosch, Hymen. Javan. (1861) 5, pl. 1; --Copel., FFP 1 (1958) 75. --Trichomanes beccarianum Cesati, Atti Accad. Napoli, Part 8 7 (1876) 8, pl. 1, Fig. 2; --Copel., PJS 51 (1933) 200, pl. 29; --Microgonium beccarianum (Cesati) Copel., PJS 67 (1938) 63; FFP 1 (1958) 74. Photos

Gonocormus Bosch = Crepidomanes

Hymenophyllum Sm.

1. Hymenophyllum acanthoides (Bosch) Rosenst., Bull. Jard. Bot. Buitenz. 2, 2 (1911) 25; --Copel., PJS 64 (1937) 45, pl. 17; --Meringium acanthoides (Bosch) Copel., PJS 67 (1938) 42; FFP 1 (1958) 63. Photos
2. Hymenophyllum acutum (C. Presl) Ebihara & K. Iwats., Blumea 51 (2006) 232; --Crepidomanes acutum (C. Presl) K Iwats.; --Trichomanes acutum C.Presl, Epim. Bot. (1849) 258. 
3. Hymenophyllum angulosum Christ, PJS 3 c (1908) Bot. 269; --Copel., PJS 64 (1937) 33, pl.  13; --Mecodium angulosum (Christ) Copel., PJS 64 (1937) 109, pl. 50; FFP 1 (1958) 49. Photos
4. Hymenophyllum badium Hook. & Grev., Icon. Fil. (1828) pl. 76; --Copel., PJS 64 (1937) 144, pl. 76; --Mecodium badium (Hook. & Grev.) Copel., PJS 67 (1938) 23; FFP 1 (1958) 53. Photos
5. Hymenophyllum bartlettii (Copel.) C.V.Morton; --Meringium bartlettii Copel., PJS 73 (1940) 464; FFP 1 (1958) 62. 
6. Hymenophyllum bicolanum Copel., PJS 64 (1937) 31, pl. 10; --Meringium bicolanum (Copel.) Copel., PJS 67 (1938) 40; FFP 1 (1958) 60.
7. Hymenophyllum blandum Raciborski, Pterid. Buitenz. (1898) 20; --Copel., PJS 64 (1937) 50,  pl. 18; --Meringium blandum (Raciborski) Copel., PJS 67 (1938) 43; FFP 1 (1958) 63.
8. Hymenophyllum bontocense Copel., PJS 64 (1937) 33, pl. 12; --Meringium bontocense (Copel.) Copel., PJS 67 (1938) 41; FFP 1 (1958) 60.
9. Hymenophyllum campanulatum H.Christ, PJS 2 c (1907) Bot.155; --Copel., PJS 64 (1937) 32,  pl. 11; --Meringium campanulatum (Christ) Copel., PJS 67 (1938) 40; FFP 1 (1958) 60.
10. Hymenophyllum crispatum Wall. in Hook. & Grev., Icon. Fil. (1828) pl. 72; --Copel., PJS 64 (1937) 148, pl. 77; --Mecodium crispatum (Wall.) Copel., PJS 67 (1938) 23.
11. Hymenophyllum denticulatum Sw., Schrad. J. (1801) 100; --Copel., PJS 64 (1937) 41, pl. 15; --Meringium denticulatum (Sw.) Copel., PJS 67 (1938) 42; FFP 1 (1958) 62. Photos
12. Hymenophyllum digitatum (Sw.) Fosberg; --Crepidomanes digitatum (Sw.) K Iwats.; --Trichomanes digitatum Sw., Syn. Fil. (1806) 370; --Copel., PJS 51 (1933) 159, pl. 7, Fig. 3, 4; --Microtrichomanes digitatum (Sw.) Copel., PJS 67 (1938) 36; FFP 1 (1958) 55.
13. Hymenophyllum edanoi (Copel.) C.V.Morton; --Mecodium edanoi Copel., PJS 81 (1952) 2; FFP 1 (1958) 50.
14. Hymenophyllum emarginatum Sw., Schrad. J. 2 of 1800 (1801) 101; --Copel., PJS 64 (1937)  118, pl. 58; --Mecodium emarginatum (Sw.) Copel., PJS 67 (1938) 21; FFP 1 (1958) 52. Photos  
15. Hymenophyllum fimbriatum J.Sm., J. Bot. 3 (1841) 418, nomen; --Hook., Sp. Fil. (1844) 102,  pl. 36C; --Copel., PJS 64 (1937) 122, pl. 60; --Mecodium fimbriatum (J.Sm.) Copel., PJS 67 (1938) 21; FFP 1 (1958) 52. Photos
16. Hymenophyllum holochilum (Bosch) C.Chr., Index Fil. (1905) 226, 362; --Copel., PJS 64  (1937) 34, pl. 14; --Meringium holochilum (Bosch) Copel., PJS 67 (1938) 41; FFP 1 (1958) 62. --Hymenophyllum merrillii Christ, PJS 2 c (1907) Bot. 154; --Copel., PJS 64 (1937) 33, pl. 13; --Meringium merrillii (Christ) Copel., PJS 67 (1938) 41; FFP 1 (1958) 61. Photos
17. Hymenophyllum imbricatum Blume, Enum. Pl. Jav (1828) 220; --Copel., PJS 64 (1937) 137,  pl. 70, 71; --Mecodium imbricatum (Blume) Copel., PJS 67 (1938) 22; FFP 1 (1958) 53. Photos
18. Hymenophyllum javanicum Spreng., Syst. Veg. (ed. 16) [Sprengel] 4 (1827) 132.
19. Hymenophyllum johorense Holttum, Gard. Bull. Sing. 4 (1929) 408, with fig.; --Copel., PJS 64  (1937) 51, pl. 19; --Meringium johorense (Holttum) Copel., FFP 1 (1958) 64. --Hymenophyllum reductum Copel., PJS 64 (1937) 53, pl. 20; --Meringium reductum (Copel.) Copel., PJS 67 (1938) 43; FFP 1 (1958) 64. Photos
20. Hymenophyllum klabatense H.Christ, Verh. Nat. Ges. Basel 11 (1894) 4; --Copel., PJS 64 (1937) 30; --Meringium klabatense (Christ) Copel., PJS 67 (1938) 40; FFP 1 (1958) 59.
21. Hymenophyllum nitiduloides Copel., PJS 64 (1937) 112, pl. 52; --Mecodium nitiduloides (Copel.) Copel., PJS 64 (1937) 112, pl. 52; FFP 1 (1958) 50. 
22. Hymenophyllum nitidulum (Bosch) Ebihara & K.Iwats., Taxon 53(4) (2004) 941; --Trichomanes nitidulum Bosch, Pl. Jungh. 1 (1856) 547; --Gonocormus nitidulus (Bosch) Prantl; --Microtrichomanes nitidulum (Bosch) Copeland; --Sphaerocionium nitidulum (Bosch) K.Iwats. Photos
23. Hymenophyllum pachydermicum Cesati, Atti Accad. Napoli 8 (1876) 8; --Meringium pachydermicum (Cesati) Copel., PJS 67 (1938) 41; FFP 1 (1958) 57. Photos
24. Hymenophyllum pallidum (Blume) Ebihara & K. Iwats.; --Crepidomanes pallidum (Blume) K Iwats.; --Pleuromanes pallidum (Blume) C. Presl, Epim. Bot. (1849) 258; --Copel., FFP 1 (1958) 67; --Trichomanes pallidum Blume, Enum. Pl. Jav. (1828) 226; --Copel., PJS 51 (1933) 141. Photos
25. Hymenophyllum paniculiflorum C.Presl, Hymen. (1843) 147; --Copel., PJS 64 (1937) 110,  pl. 51; --Mecodium paniculiflorum (C.Presl) Copel., PJS 67 (1938) 19; FFP 1 (1958) 50. Photos
26. Hymenophyllum pilosissimum C.Chr., Gard. Bull. 7 (1934) 213; --Sphaerocionium pilosissimum (C.Chr.) Copel., PJS 67 (1938) 33; FFP 1 (1958) 54. Photos
27. Hymenophyllum polyanthos (Sw.) Sw., Schrad. J. ii of 1800 (1801) 102; --Copel., PJS 64  (1937) 97, pl. 46, 47; --Mecodium polyanthos (Sw.) Copel., PJS 67 (1938) 19; FFP 1 (1958) 49.
28. Hymenophyllum productum Kunze, Bot. Zeit. 6 (1848) 305; --Copel., PJS 64 (1937) 118, pl.  54; --Mecodium productum (Kunze) Copel., PJS 67 (1938) 20; FFP 1 (1958) 51.
29. Hymenophyllum ramosii Copel., PJS 64 (1937) 34, pl. 9, Fig. 4-6; --Meringium ramosii (Copel.) Copel., PJS 67 (1938) 31; FFP 1 (1958) 61. Photos
30. Hymenophyllum reinwardtii Bosch., Pl. Jungh. 1 (1856) 567; --Copel., PJS 64 (1937) 115, pl.  55; --Mecodium reinwardtii (Bosch.) Copel., PJS 67 (1938) 20; FFP 1 (1958) 51. Photos
31. Hymenophyllum serrulatum (C.Presl) C.Chr. --Meringium meyenianum C.Presl, Hymen. (1843) 116, pl. 8 B; --Copel., FFP 1 (1958) 58; --Hymenophyllum meyenianum (C.Presl) Copel., PJS 64 (1937) 25, pl. 8; FFP 1 (1958) 58. --Hymenophyllum edentulum (Bosch) C.Chr., Index Fil. (1905) 360; --Meringium edentulum (Bosch) Copel., PJS 67 (1938) 41; FFP 1 (1958) 58. --Hymenophyllum bakeri Copel., Sarawak Mus. J. 2 (1917) 309; --Copel., PJS 64 (1937) 29; --Meringium bakeri (Copel.) Copel., PJS 67 (1938) 40; FFP 1 (1958) 59. --Hymenophyllum pulchrum Copel., PJS 64 (1937) 22; --Meringium pulchrum (Copel.) Copel., PJS 67 (1938) 42; FFP 1 (1958) 57. Photos
32. Hymenophyllum thuidium Harrington, J. Linn. Soc. Bot. 16 (1877) 25; --Copel., PJS 64 (1937) 116, pl. 56; --Mecodium thuidium (Harrington) Copel., PJS 67 (1938) 20; FFP 1 (1958) 52.
33. Hymenophyllum vittatum Copel., PJS 64 (1937) 31, pl. 9, Fig. 1-3; FFP 1 (1958) 59; --Meringium vittatum (Copel.) Copel., PJS 67 (1938) 40; FFP 1 (1958) 59.
34. Hymenophyllum zamboanganum ined; --Microtrichomanes zamboanganum Copel., PJS 81 (1952) 2; FFP 1 (1958) 56; --Trichomanes zamboanganum (Copel.) C.V.Morton.

Sphaerocionium C.Presl = Hymenophyllum

Vandenboschia Copel.

Photos

1. Vandenboschia auriculata (Blume) Copel.; --Crepidomanes auriculatum (Blume) K Iwats.; --Trichomanes auriculatum Blume. Photos
2. Vandenboschia johnstonensis (Bailey) Copel.; --Crepidomanes johnstonense (Bailey) K Iwats.; --Trichomanes johnstonense Bailey.
3. Vandenboschia maxima (Blume) Copel.; --Crepidomanes maximum (Blume) K Iwats.; --Trichomanes maximum Blume. Photos

Hypodematiaceae 

Edited by Julie F. Barcelona, 20 January 2013

Didymochlaena Desv.

1. Didymochlaena truncatula (Sw.) J.Sm. Photos

Hypodematium Kunze

1. Hypodematium crenatum (Forssk.) Kuhn. Photos

Leucostegia C.Presl

1. Leucostegia immersa (Wall.) C.Presl, Tent. (1836) 95, t. 4, Fig. 11. Photos
2. Leucostegia pallida Mett. Photos

Lindsaeaceae

Isoloma J.Sm. = Lindsaea

Lindsaea Dryand. ex Sm.

1. Lindsaea adiantoides J.Sm. --Lindsaea tropidorhachis Alderw. Photos
2. Lindsaea apoensis Copel. --Lindsaea havicei Copel. Photos
3. Lindsaea bouillodii Christ, Not. Syst. 1 (1909) 59;  Thailand and N Vietnam to Malesia: Peninsular Malaysia, Sumatra, W Java, Borneo, Banka and Natuna Is. New to the Philippines: PALAWAN: Mt. Mantalingahan, alt. 1200m, Co 1793 (A, PNH, PUH). 
4. Lindsaea capillacea Christ
5. Lindsaea cultrata (Willd.) Sw., Syn. Fil. (1806) 119; Lindsaea decomposita sensu Copel. (1960) non Willd., pro parte.
6. Lindsaea divergens Hook. & Grev. Isoloma divergens (Hook. & Grev.) J.Sm.
7. Lindsaea doryphora Kramer Lindsaea parasitica sensu Copel. (1960) non Wall. Photos  
8. Lindsaea ensifolia Sw. ssp. ensifolia Photos
9. Lindsaea fissa Copel. Lindsaea hymenophylloides sensu Copel. (1960), non Blume. Photos  
10. Lindsaea gueriniana (Gaudich.) Desv., Mem. Soc. Linn. Paris 6 (1827) 312; --Isoloma guerinianum (Gaudich.) Fée. --Isoloma fuligineum (Copel.) Copel. Photos
11. Lindsaea hamiguitanensis D.N.Karger & V.B.Amoroso, Phytotaxa 56: 18. --Type: Karger 444 (holo PNH, UC, Z, iso). Endemic to the Philippines. MINDANAO: Mt. Hamiguitan. Conservation status: Vulnerable (DAO 2017-11). Photos
12. Lindsaea heterophylla Dryander
13. Lindsaea javanensis Blume
14. Lindsaea lobata Poir. in Lam., Encycl. Suppl. 3 (1813) 448; 
15. Lindsaea longifolia Copel. 
16. Lindsaea lucida Blume, Enum. Pl. Jav. (1828) 216; Lindsaea securifolia C.Presl ex Goldm. Lindsaea gracilis Blume in Copel. (1960), excl. syn. ssp. lucida. Photos  
17. Lindsaea merrillii Copel. ssp. merrillii
18. Lindsaea oblanceolata Alderw., Bull. Jard. Bot. Buitenz. 2, 23 (1916) 15. Photos
19. Lindsaea obtusa J.Sm. ex Hook., Sp. Fil. 1 (1846) 224; Lindsaea decomposita sensu Copel. (1960) non Willd., excl. syn. Lindsaea davallioides sensu Copel. (1960) non Blume. Photos  
20. Lindsaea orbiculata (Lam.) Mett. ex Kuhn Photos
    • var. orbiculata
    • var. conmixta (Tagawa) Kramer
21. Lindsaea ovata J.Sm. Isoloma ovatum (J.Sm.) C.Presl
22. Lindsaea pellaeformis Christ Luzon: Isabela prov., Divilacan, Aubarede peninsula, 50 m alt., L. Co 3580, distributed as L. gueriniana (A, PNH, PUH, CAHUP, L, US, K, MICH). In here sympatrically occurring with L. gueriniana, which according to Kramer (1971), might just be an extreme form of that variable species. New to the Philippines. Previously known only from Sulawesi. 
23. Lindsaea philippinensis Kramer. --Isoloma angustum (Copel.) Tard., comb. superfl.; non L. angusta Copel. (1952). According to Kramer (1971), doubtfully distinct from L. jamesonioides Baker of Sabah. Photos  
24. Lindsaea polyctena Kramer. Photos
25. Lindsaea pulchella (J.Sm.) Mett. ex Kuhn. Photos
    
    • var. pulchella. --Lindsaea adiantifolia (Hook.) Copel. --Lindsaea adiantoides sensu Copel. (1960), pro parte.
    • var. blanda (Mett. ex Kuhn) Kramer. --Lindsaea cyathicola Copel.
26. Lindsaea ramosii Copel. Very doubtfully distinct from L. longifolia.
27. Lindsaea repens (Bory) Thwaites
    
    • var. sessilis (Copel.) Kramer, Blumea 15 (1968) 568. --Lindsaea longa Copel. --Lindsaea cultripinna Copel.
    • var. pectinata (Blume) Mett. ex Kuhn in Miq., Ann. Mus. Bot. Lugd.-Bat. 4 (1869) 277; --Lindsaea pectinata Blume. --Lindsaea macraeana (Hook. & Arn.) Copel., at least in part. --? Lindsaea roxasii Copel. --Lindsaea angusta Copel. Aberrant form from Mindanao called L. angusta Copel. may be distinguished as forma angusta (Copel.) Kramer, Blumea 15 (1968) 568. Photos
28. Lindsaea rigida J.Sm. ex Hook., Sp. Fil. 1 (1846) 217, t. 63A. --Lindsaea monosora Copel.
29. Lindsaea tetragona Kramer var. tetragona. --Lindsaea tenuifolia sensu Copel. (1960) non Blume, excl. syn.

Doubtful species

1. Lindsaea decrescens Copel. “As stated by Copeland, this is intermediate between L. repens (‘L. macraena, L. longa’) and L. fissa. The isotype in MICH from Copeland’s herbarium is rather scrappy. I doubt whether it represents a distinct species” –Kramer, Fl. Males. Ser. ii 1 (1971) 253.
2. Lindsaea longa Copel. Probably only an usually deeply incised form of L. repens (Bory) Thwaites var. sessilis (Copel.) Kramer.

Sphenomeris Maxon: see Odontosoria for the Philippine species

Odontosoria (Pr.) Fée

Photos

1. Odontosoria biflora (Kaulf.) C.Chr., Index Filic. (1905) 207; --Sphenomeris biflora (Kaulf.) Tagawa. --Sphenomeris chusana sensu Copel. (1960), pro parte 
2. Odontosoria chinensis (L.) J.Sm., Bot. Voy. Herald [Seemann] 10 (1857) 430. 1857; --Sphenomeris chinensis (L.) Maxon. --Sphenomeris chusana (L.) Copel. ssp. chinensis Tropical to subtropical Asia and Oceania. Throughout the Philippines. Terrestrial or epilithic in exposed sites, from low altitudes to 2100m. Photos
   • ssp. divaricata (Christ) Kramer
3. Odontosoria retusa (Cav.) J.Sm., Bot. Voy. Herald [Seemann] 10 (1857) 430. 1857; --Sphenomeris retusa (Cav.) Maxon. Photos  

Osmolindsaea (K.U.Kramer) Lehtonen & Christenh.

Edited by Julie F. Barcelona, 2 May 2013

Photos

1. Osmolindsaea odorata (Roxb.) Lehtonen & Christenh., Bot. J. Linn. Soc. 163 (2010): 335 (-336); --Lindsaea odorata Roxb., Calc. J. Nat. Hist. 4 (1844) 511; --Lindsaea cultrata sensu Copel. (1960), pro parte Lindsaea cultrata (Willd.) Sw. var. varia Copel. --?Lindsaea cultrata var. minor Hook. --Lindsaea loheriana Christ. Photos

Tapeinidium (C.Presl) C.Chr.

1. Tapeinidium acuminatum Kramer. Photos  
2. Tapeinidium calomelanos Kramer
3. Tapeinidium gracile (Blume) Alderw.
4. Tapeinidium luzonicum (Hook.) Kramer Photos
   • var. luzonicum
   • var. leptophyllum Kramer
5. Tapeinidium pinnatum (Cav.) C.Chr. Photos  
6. Tapeinidium biserratum (Blume) Alderw. Probably a natural hybrid. According to Kramer (Fl. Males. Ser. ii 1 (1971): “The type of this ‘species’ is intermediate between T. luzonicum and T. pinnatum; there are 25-30 intermediates… Their status is not clear. They occur throughout the common range of the two species”.

Lomariopsidaceae 

Cyclopeltis J.Sm.

1. Cyclopeltis crenata (Fee) C.Chr. --Cyclopeltis zamboangana Copel. Photos
2. Cyclopeltis presliana (J.Sm.) Berkeley. --Cyclopeltis cumingiana (Fee) C.V.Morton.

Lomariopsis Fée

1. Lomariopsis kingii (Copel.) Holttum.
2. Lomariopsis lineata (C.Presl) Holttum. --Lomariopsis smithii Fée. --Lomariopsis papyracea Copel. Photos
3. Lomariopsis subtrifoliata (Copel.) Holttum.

Doubtfully occurring in the Philippines

1. Lomariopsis spectabilis (Kunze) Mett. Philippine specimens formerly so-named to this species all appear to have narrower fertile pinnae and hence believed by Holttum to be L. kingii. (Fl. Males. ser. 2, 1 (1978) 262. Photos

Lygodiaceae

Lygodium Sw.

1. Lygodium auriculatum (Willd.) Alston, Reinwardtia 5 (1959) 16. --Lygodium semihastatum (Cav.) Desv., Prodr. (1827) 203; --Copel., FFP 1 (1958) 34. Indochina, Micronesia, Malesia: E Borneo, Philippines. LUZON, POLILLO, MINDORO, SAMAR, MINDANAO. Terrestrial, usually in thickets and disturbed forests. Photos  
2. Lygodium circinnatum (Burm.) Sw., Syn. Fil. (1806) 153; --Copel., FFP 1 (1958) 33. --Lygodium basilanicum Christ, PJS 2 c (1907) Bot. 179; --Copel., FFP 1 (1958) 34. Sri Lanka and India to S China, across Malesia to the Solomons. Throughout the Philippines. Terrestrial, common in brush and open sites and disturbed lowland forests. Photos  
3. Lygodium flexuosum (L.) Sw., Schrader’s J. (1801) 106; --Copel., FFP 1 (1958) 35. Sri Lanka, India, Himalayas to S China, Hongkong, Taiwan, Ryukyus, across Malesia, N Queensland, Melanesia. Terrestrial, in brush and open forests at low altitudes. Photos  
4. Lygodium japonicum (Thunb.) Sw., Schrader’s J. (1801) 106; --Copel., FFP 1 (1958) 36. --Lygodium mearnsii Copel., PJS 3 c (1908) Bot. 37; --Copel., FFP 1 (1958) 35. Sri Lanka, Himalayas (Kashmir eastwards) to E China (Zejiang), Korea, Japan and southwards to Thailand and S China; in Malesia: Banka, C & E Java, Sulawesi, Philippines, Moluccas, Lesser Sunda Isls and New Guinea. Naturalized in Florida and Texas. Terrestrial in open forests and throroughly cleared sites at low to medium altitudes. Photos  
5. Lygodium longifolium (Willd.) Sw. in Schrader’s J. Bot. 1801, pt. 2 (1803) 305; S India, Hainan. In Malesia: Sumatra, Peninsular Malaysia, Riouw and Lingga Isls., Borneo, Philippines. LUZON (doubtful), PALAWAN.
6. Lygodium merrillii Copel., PJS 2 c (1907) Bot. 146, t. 4; FFP 1 (1958) 37. --Lygodium matthewii Copel., PJS 3 c (1908) Bot. 36; --Copel., FFP 1 (1958) 37. SW China (Guizhou), N Vietnam, Sumatra, Borneo (Sarawak), Philippines. LUZON: Isabela; S Luzon, MINDORO, LEYTE. Photos
7. Lygodium microphyllum (Cav.) R.Br., Prodr. Fl. Nov. Holl. (1810) 162. --Lygodium scandens (L.) Sw., Schrader’s J. (1801) 106; --Copel., FFP 1 (1958) 36. Tropical Africa, SE Asia, N to Bangladesh and Hongkong and the Ryukyus, across Malesia to the Solomons, New Caledonia, N and E Australia S to New South Wales. LUZON to MINDANAO (along the Pacific side), MINDORO, CULION, BASILAN. Terrestrial in moe or less open sites. Photos
8. Lygodium salicifolium Sw.
9. Lygodium versteegii Christ, Nova Guinea 8 (1909) 161; --Copel., FFP 1 (1958) 37. New Guinea and Philippines. LUZON: Laguna (Mt Makiling), Quezon (Guinayangan), Camarines (Bicol National Park), MINDANAO: Agusan del Norte (Mt Hilong-hilong). Photos  

Marattiaceae

Angiopteris Hoffm.

1. Angiopteris evecta Sw., Syn. Fil. (1806) 395; --Copel., FFP 1 (1958) 22, in obs., pro syn. A. palmiformis. --Angiopteris palmiformis (Cav.) C.Chr., Dansk Bot. Arki 3, 9 (1937) 30, t. 3, Fig. 8, 9; --Copel., FFP 1 (1958) 22. --Angiopteris elmeriana Copel. in Elmer, LPB 5 (1913) 1679; --Copel., FFP 1 (1958) 23. --Angiopteris uncinata Christ, PJS 2 c (1907) Bot. 186, non Vriese (1853); --Copel., FFP 1 (1958) 24. --Angiopteris caudata Vriese, Monog. (1853) 20; --Copel., FFP 1 (1958) 24. --Angiopteris cartilagidens Christ, Bull. Boiss. 6 (1898) 207, t. 4, Fig. 8; --Copel., FFP 1 (1958) 24. --Angiopteris pruinosa Kunze, Bot. Zeit. 4 (1846) 417; --Copel., FFP 1 (1958) 25. --Angiopteris angustifolia C Presl var. prunifera Hieron., Hedwigia 61 (1919) 257; --Copel., FFP 1 (1958) 25. Conservation status: Other Threatened Species (DAO 2017-11 as Angiopteris palmiformis (Cav.) C.Chr.). Photos  

Christensenia Maxon

1. Christensenia aesculifolia (Blume) Maxon, Proc. Biol. Soc. Wash. 18 (1905) 240. Conservation status: Endangered (DAO 2017-11). Photos
   • ssp. aesculifolia; --Copel., FFP 1 (1958) 27; --Aspidium aesculifolium Blume, Enum. Pl. Javae 2 (1898) 143; --Kaulfussia aesculifolia (Blume) Blume, Enum. Pl. Javae 2 (1898) 260; --Type: Blume? (L51148), Java. India (Assam) to Malesia.  
2. Christensenia lobbiana (de Vriese) Rolleri, Amer. Fern Jour. 83(1) (1993) 17; --Kaulfussia lobbiana de Vriese & Harting, Monogr. Marattiac. (1853) 14; --Type: "Habitat in insulam Javam ". --Christensenia cumingiana Christ, Philipp. J. Sci. 2 (1907) 186; --Type: Copeland, s.n. (holo, MICH (HC1110)?, iso, PNH62411?).  Philippines, Borneo, Thailand, Sumatra and Java. Photos. 
   There may very well be one morphologically variable species. I have seen both 3-5-foliate fronds in one plant in Panay. Larger fronds tend to be more dissected than smaller ones.

Macroglossum Copel. [sometimes included in Angiopteris]

1. Macroglossum alidae Copel.

Marattia Sw. p.p. = Ptisana

Ptisana Murdock

Edited by Julie F. Barcelona, 13 January 2013

Photos 

1. Ptisana fraxinea (Sm.) Murdock, Taxon 57(3) (2008) 746; --Marattia fraxinea Sm., Pl. Ic. ined. 2. (1790) t. 48. Photos
2. Ptisana pellucida (C.Presl) Murdock, Taxon 57(3) (2008) 747; --Marattia pellucida C Presl, Suppl. Tent. (1845) 10; --Copel., FFP 1 (1958) 26. --Marattia ternatea Christ, PJS 2 c (1907) Bot. 185; --Copel., FFP 1 (1958) 26. C LUZON southwards to MINDANAO: Mt. Matutum. Photos  
3. Ptisana sylvatica (Blume) Murdock, Taxon 57(3) (2008) 747; --Marattia sylvatica Blume, Enum. (1828) 256; --Copel., FFP 1 (1958) 27. LUZON: Cordillera Central. Common in mossy montane forests to c. 2400m. Photos

Marsileaceae

Marsilea L.

1. Marsilea crenata C.Presl. Conservation status: Vulnerable (DAO 2017-11 as Marsilea minuta L.). Photos  

Matoniaceae

Matonia R.Br.

1. Matonia foxworthyi Copel. Conservation status: Endangered (DAO 2017-11). Photos

Phanerosorus Copel.

1. Phanerosorus major Diels, Notizbl. Bot. Gart. Berlin-Dahlem 11 (1932) 311. New Guinea, Seram (= Ceram), Southern Moluccas, East Indonesia, Philippines. SAMAR. Conservation status: Critically Endangered (DAO 2017-11). Photos

Nephrolepidaceae

Edited by Julie F. Barcelona, 16 January 2013

Nephrolepis Schott

1. Nephrolepis acutangula C.Presl --Nephrolepis pubescens Copel.
2. Nephrolepis auriculata (L.) Trimen.
3. Nephrolepis biserrata (Sw.) Schott.
4. Nephrolepis clementis Christ. Probably synonymous with Nephrolepis radicans.
5. Nephrolepis cordifolia (L.) Presl Nephrolepis glabra Copel. Photos
6. Nephrolepis exaltata (L.) Schott. Photos
7. Nephrolepis falcata (Cav.) C.Chr. Photos
8. Nephrolepis hirsutula (G Forst.) C.Presl
9. Nephrolepis multiflora (Roxb.) Jarrett ex C.V.Morton.
10. Nephrolepis radicans (Burm.) Kuhn.

Oleandraceae

Oleandra Cav.

1. Oleandra cumingii J.Sm., J.Sm., J. Bot. (Hooker) 4 (1842) 413; --Type: Cuming 60 (holo, K; iso, B, BM, SING, US), Luzon. --Oleandra macrocarpa C.Presl, Epimel. Bot. (1851) 41; --Fée, Mém. Foug., 5. Gen. Filic. (1852) 304; --Type: Cuming 60 p.p. (holo, PRC, teste Holttum; iso, B, BM, K, SING, US), Luzon. --Oleandra scandens Copel., Philipp. J. Sci. 46 (1931) 218; --Type: Elmer 6513 (holo, US), Baguio. Photos
2. Oleandra herrei Copel., Philipp. J. Sci. 81 (1952) 12; --Type: Herre s.n. (holo, UC). This is considered a synonym of O. neriiformis by Hovenkamp & Ho (2012).
3. Oleandra maquilingensis Copel., Philipp. J. Sci. 46 (1931) 217; --Type: Matthew s.n. (lecto, MICH 1210440). --Oleandra colubrina sensu Copel., Fern Fl. Philipp. 1 (1958) 181, pro parte. This is considered a synonym of O. neriiformis by Hovenkamp & Ho (2012).
4. Oleandra musifolia Blume, Enum. Pl. Javae (1828) 141 (as Oleandra musaefolium); --Type: Blume s.n. (holo, L), Indonesia, Java, Gedeh. --Oleandra benguetensis Copel., Philipp. J. Sci. 46 (1931) 217; --Fern Flora of the Philippines (1958) 183. 19; --Type: Elmer 6286 (holo, US), Luzon, Baguio.
5. Oleandra neriiformis Cav., Anales Hist. Nat. 2 (1799) 115 (as “neriformis”); --Type: Née s.n. (holo, MA 476029, teste Gariletti), Luzon, Mauban. Photos
6. Oleandra nitida (Copel.) Copel., Fern Flora of the Philippines (1958) 181. --Oleandra colubrina var. nitida Copel., Philipp. J. Sci., C 3 (1908) 33; --Type: Copeland 1474 (lecto, US), Mindanao, Mt. Apo. This is considered a synonym of O. neriiformis by Hovenkamp & Ho (2012).
7. Oleandra sibbaldii Grev., Ann. and Mag. Nat. Hist. Ser. 2 1 (1848) 327; --Oleandra cumingii var. sibbaldii (Grev.) Baker, Syn. Fil. (Hooker & Baker) (1868) 303.
   1868; --Type: Sibbald s.n. (holo, E, barcode E00417634), French Polynesia, Tahiti. Photos

Ophioglossaceae

Botrychium Sw.

1. Botrychium daucifolium Wall. ex Hook. & Grev., Icon. Fil. (1829) t. 161; --Copel., FFP 1 (1958) 20. Conservation status: Vulnerable (DAO 2017-11). Photos  
2. Botrychium lanuginosum Wall. ex Hook. & Grev., Icon. Fil. (1828) t. 79; --Copel., FFP 1 (1958) 19. Conservation status: Endangered (DAO 2017-11).

Helminthostachys Kaulf.

1. Helminthostachys zeylanica (L.) Hook., Genera (1840) t. 47; --Copel., FFP 1 (1958) 20; -- Holttum, Rev. Fl. Mal. 2 (1954) 42, Fig. 2. Sri Lanka and India to Hainan, Taiwan, across Malesia to New Caledonia. Throughout the Philippines. Terrestrial in open brushy places near sea level on damp ground. Conservation status: Critically Endangered (DAO 2017-11). Photos  

Ophioglossum L.

1. Ophioglossum costatum R.Br.
2. Ophioglossum gramineum Willd. var. gramineum .
3. Ophioglossum intermedium Hook., Icon. Pl. 10 (1854) t. 995; --Copel., FFP 1 (1958) 18. Photos
4. Ophioglossum nudicaule L.f., Suppl. (1781) 443; --Copel., FFP 1 (1958) 18.
   • var. nudicaule
   • var. macrorrhizum (Kunze) Clausen
5. Ophioglossum pendulum L., Sp. Pl. ed. 2 (1763) 1518; --Copel., FFP 1 (1958) 19; --Holttum, Rev. Fl. Mal. 2 (1954) 40. Conservation status: Endangered (DAO 2017-11). Photos.  
   • forma pendulum Old World tropics from Madagascar through tropical Asia to Polynesia. Throughout the Philippines. Epiphytic in everwet rainforests.
   • forma pendulum X forma nutans Alderw.
   • forma pendulum X forma falcatum (C Presl) Wief. 
6. Ophioglossum ramosii Copel., PJS 56 (1935) 97, pl. 1, Fig. 1, 2; FFP 1 (1958) 18. Conservation status: Endangered (DAO 2017-11). Photos
7. Ophioglossum reticulatum L., Sp. Pl. (1753) 1063; --Copel., FFP 1 (1958) 17; forma reticulatum. --Ophioglossum petiolatum Hook., Exot. Fl. (1823) pl. 56; --Copel., FFP 1 (1958) 17. --Ophioglossum nudicaule L. f. var. grandifolium Clausen, Mem. Torr. Bot. Club 19 (1938) 151; --Copel., FFP 1 (1958) 18. Pantropic. Probably throughout the Philippines. Terrestrial in humid mostly shaded sites. Only present during the wet season, 0-1600m. Photos  
   • forma dilatatum (Miq.) Wief.
   • forma complicatum (Miq.) Wief. 

Osmundaceae

Osmunda L.

1. Osmunda banksiifolia (C Presl) Kuhn, Ann. Lugd.-Bat. 4 (1869) 299; --Copel., FFP 1 (1958) 30. Conservation status: Other Threatened Species (DAO 2017-11). Photos
2. Osmunda bromeliifolia (C Presl) Copel., PJS 2 c (1907) Bot. 147; --Copel., FFP 1 (1958) 30.
3. Osmunda herbacea Copel., PJS 40 (1929) 291; FFP 1 (1958) 31.

Plagiogyriaceae

Plagiogyria (Kunze) Mett.

 
1. Plagiogyria adnata (Blume) Bedd.
2. Plagiogyria egenolfioides (Baker) Copel. var. latipinna (Copel.) XC Zhang & Noot. --Plagiogyria tuberculata Copel. Photos
3. Plagiogyria euphlebia (Kunze) Mett. --Plagiogyria christii Copel. Photos  
4. Plagiogyria falcata Copel.
5. Plagiogyria glauca (Blume) Mett. --Plagiogyria nana Copel.; --Plagiogyria glauca var. nana (Copel.) C.Chr. Photos
6. Plagiogyria pycnophylla (Kunze) Mett. Photos
7. Plagiogyria stenoptera (Hance) Diels.

Polypodiaceae

including Drynariaceae, Grammitidaceae, Loxogrammaceae, Platyceriaceae

Edited by Barbara Parris, 30 September 2012

Acrosorus Copel.

1. Acrosorus friderici-et-pauli (Christ) Copel. --Acrosorus exaltatus Copel. --Acrosorus merrillii Copel. --Acrosorus symmetricus Copel. --Acrosorus triangularis (Scort. ex Bedd.) Copel.
2. Acrosorus nudicarpus PM Zamora & Co, Nat. Appl. Sc. Bull. Univ. Philipp. 32 (1980) 47,  t. 2; --Xiphopteris nudicarpa (PM Zamora & Co) Parris in Parris et al., Pl. Mt Kinabalu 1 (1992) 77. --Grammitis palasaba M.G.Price, Contr. Univ. Mich. Herb. 16 (1987) 196. Conservation status: Vulnerable (DAO 2017-11 as Xiphopterella nudicarpa (P.M.Zamora & Co) Parris).
3. Acrosorus streptophyllus (Baker) Copel. Recorded by Price (1972); some material so identified belongs to A. friderici-et-pauli. Occurrence in the Philippines needs confirmation.

Aglaomorpha Schott = Drynaria

 Anapausia C.Presl  = Leptochilus

Arthromeris (T.Moore) J.Sm.

1. Arthromeris proteus (Copel.) Tagawa. Conservation status: Other Threatened Species (DAO 2017-11 as Arthromeris mairei (Brause) Ching).

Belvisia Mirbel = Lepisorus (J.Sm.) Ching

Calymmodon C.Presl

Edited by Barbara Parris, 30 September 2012

1. Calymmodon clavifer (Hook.) T.Moore.
2. Calymmodon cucullatus (Nees & Blume) C.Presl Photos
3. Calymmodon gracilis (Fée) Copel. Photos
4. Calymmodon muscoides (Copel.) Copel. Photos
5. Calymmodon ordinatus Copel.

Christopteris Copel. = Drynaria

Chrysogrammitis Parris

1. Chrysogrammitis glandulosa (J.Sm.) Parris. --Ctenopteris merrittii (Copel.) Tagawa. Photos
2. Chrysogrammitis musgraviana (Baker) Parris. --Xiphopteris apoensis (Copel.) Copel.

Ctenopteris Blume = Chrysogrammitis, Ctenopterella, Dasygrammitis,  Prosaptia, Themelium, Tomophyllum

  Christiopteris Copel. = Drynaria

 Colysis C.Presl = Leptochilus

  Crypsinus C.Presl = Selliguea

Ctenopterella Parris

1. Ctenopterella blechnoides (Grev.) Parris, Gard. Bull. Sing. 58 (2007) 235; --Ctenopteris blechnoides (Grev.) Wagner & Grether. Photos
2. Ctenopterella denticulata (Blume) Parris. --Ctenopteris solida Kunze in Zoll., Syst. Verz. (1854) 37.
3. Ctenopterella pediculata (Baker) Parris, Gard. Bull. Sing. 58 (2007) 237; --Ctenopteris pediculata (Baker) Copel.

Dasygrammitis Parris

1. Dasygrammitis fuscata (Blume) Parris, Gard. Bull. Sing. 58 (2007) 239; --Ctenopteris fuscata (Blume) Kunze.
2. Dasygrammitis malaccana (Baker) Parris. --Ctenopteris matutumensis Copel., Fern Fl. Philip. 3 (1960) 528, anglice, non rite publice. --Ctenopteris pachycaulis (Copel.) Copel.
3. Dasygrammitis mollicoma (Nees & Blume) Parris, Gard. Bull. Sing. 58 (2007) 239; --Ctenopteris mollicoma (Nees & Blume) Kunze. Photos

 Dendroglossa C.Presl = Leptochilus

 Diblemma J.Sm. = Microsorum samarense

 Drymoglossum C.Presl = Pyrrosia

Drynaria (Bory) J.Sm.

1. Drynaria aglaomorpha Christenh., Phytotaxa 230 (2015) 299; --Aglaomorpha splendens (J.Sm.) Copel., PJS 6 c (1911) 141; --Dryostachyum splendens J.Sm., J. Bot. (Hooker) 3 (1841) 399. Conservation status: Vulnerable (DAO 2017-11). Photos
   
2. Drynaria cornucopia (Copel.) Alderw., Bull. Dépt. Agric. Indes Néerl. 21 (1908) 8; --Aglaomorpha cornucopia (Copel.) M.C.Roos, Blumea 31 (1985) 153; --Thayeria cornucopia Copel., PJS 1 (1906) Suppl. 165, t. 28. Conservation status: Vulnerable (DAO 2017-11). Photos
   
3. Drynaria descensa Copel., PJS 3 c (1908) 36. Photos
   
4. Drynaria heraclea (Kunze) T.Moore, Index Fil. (T. Moore) (1862) 346; --Aglaomorpha heraclea (Kunze) Copel., Univ. Calif. Publ. Bot. 16 (1929) 117; --Drynariopsis heraclea (Kunze) Ching, Sunyatsenia 5 (1940) 262; --Polypodium heracleum Kunze, Bot. Zeitung (Berlin) 6 (1848) 117. Conservation status: Vulnerable (DAO 2017-11). Photos
   
5. Drynaria meyeniana (Schott) Christenh., Phytotaxa 230 (2015) 299; --Aglaomorpha meyeniana Schott, Gen. Fil. (1834) pl. 20. Conservation status: Vulnerable (DAO 2017-11). Photos
   
6. Drynaria pilosa (J.Sm. ex Kunze) Christenh., Phytotaxa 230 (2015) 299; --Aglaomorpha pilosa (J.Sm. ex Kunze) Copel., PJS 6 c (1911) 141; --Dryostachyum pilosum J.Sm. ex Kunze, Die Farrnkräuter in kolorirten Abbildungen naturgetreu erläutert und beschrieben 1 (1844) 139. Conservation status: Vulnerable (DAO 2017-11). Photos
   
7. Drynaria quercifolia (L.) J.Sm., J. Bot. (Hooker) 3 (1841) 398; --Polypodium quercifolium L., Sp. Pl. 2 (1753) 1087. India and S China, across Malesia to tropical Australia and Fiji. Throughout the Philippines. A very common epiphyte or lithophyte in seasonal areas, in open forests, about settlements, etc., usually at low elevation. Photos
   
8. Drynaria rigidula (Sw.) Bedd., Ferns Brit. India (1869) pl. 314; --Polypodium rigidulum Sw., J. Bot. (Schrader) 1800 (1801) 26. Photos
   
9. Drynaria sagitta (Christ) Christenh., Phytotaxa 230 (2015) 299; --Christopteris sagitta (Christ) Copel. in Perkins, Fragm. Fl. Philipp. (1905) 188; --Selliguea sagitta (Christ) Fraser-Jenk., Taxon. Revis. Indian Subcontinental Pteridophytes (2008) 45; --Polypodium sagittum Christ, Bull. Herb. Boissier 6 (1898) 199. --Christopteris copelandii Christ, Bull. Herb. Boissier II. 6. (1906) 990. Conservation status: Vulnerable (DAO 2017-11 as Selliguea sagitta (Christ) Fraser-Jenk.). Photos
   
10. Drynaria sparsisora (Desv.) T.Moore, Index Fil. (1862) 348; --Polypodium sparsisorum Desv., Ges. Naturf. Freunde Berlin Mag. Neuesten Entdeck. Gesammten Naturk. 5 (1811) 315. Photos
    
11. Drynaria speciosa (Blume) Christenh., Phytotaxa 230 (2015) 299; --Photinopteris speciosa (Blume) C.Presl, Abh. Königl. Böhm. Ges. Wiss., ser. 5 6 (1851) 624; --Lomaria speciosa Blume, Enum. Pl. Javae fasc. 2 (1828) 202. --Aglaomorpha acuminata (Willd.) Hovenkamp, Fl. Males., Ser. 2, Pterid. 3 (1998) 13; --Photinopteris acuminata (Willd.) C.V.Morton, Contr. U.S. Natl. Herb. 38 (1963) 31; --Acrostichum acuminatum Willd., Sp. Pl. 5 (1810) 116. Conservation status: Vulnerable (DAO 2017-11). Photos

Drynariopsis (Copel.) Ching = Aglaomorpha

Goniophlebium (Blume) C.Presl

1. Goniophlebium benguetense (Copel.) Copel., Fern Fl. Philip. 3 (1961) 461; --Schellolepis benguetensis (Copel.) Pic.-Serm.; --Polypodium benguetense Copel., PJS 1, Suppl. 2 (1906) 256. Photos
2. Goniophlenium coadunatum Barcelona & M.G.Price.
3. Goniophlebium mengtzeense (Christ) Rödl-Linder; --Goniophlebium argutum auctt. philipp. non J.Sm.
4. Goniophlebium percussum (Cav.) Wagner & Grether; --Schellolepis percussa Pic.-Serm.
5. Goniophlebium persicifolium (Desv.) Bedd.; --Schellolepis persicifolia (Desv.) Pic.-Serm. Photos
6. Goniophlebium pseudoconnatum (Copel.) Copel. Photos
7. Goniophlebium subauriculatum (Blume) C.Presl --Goniophlebium tomentellum (C.Chr.) Copel.; --Schellolepis tomentella (C.Chr.) Pic.-Serm. Goniophlebium integrum Copel. Photos
8. Goniophlebium terrestre Copel.; --Schellolepis terrestris (Copel.) M.G.Price. Conservation status: Critically Endangered (DAO 2017-11).

 Grammitis auctt. Males., non Sw. = Oreogrammitis

Lecanopteris Reinw.

Edited by Julie F. Barcelona, 15 May 2012

1. Lecanopteris deparioides (Cesati) Baker, J. Bot. 19 (1881) 366; --NPfl. 328. --Lecanopteris curtisii Baker, J. Bot. (1881) Icon. Plant, pl. 1607; --H Gay, Gard. Bull. Sing. 45 (1993) 301. --Lecanopteris pumila Copel., Polyp. Philipp. (1905) 134; Fern Fl. Philip. 3 (1960) 499, non Blume (1829.)-- Lecanopteris philippinensis Alderw., Bull. Jard. Bot. Buitenz. Ser. 2, no. 1 (1911) 8. Sumatra, Peninsular Malaysia, Philippines to the Moluccas. MINDANAO: Lanao del Sur, Davao (Mt Apo), Davao Oriental (Mt Hamiguitan). 800-1200m. Conservation status: Endangered (DAO 2017-11). Photos
2. Lecanopteris lomarioides (Mett.) Copel. LUZON: Nueva Ecija. Conservation status: Endangered (DAO 2017-11 as Lecanopteris sarcopus (Teijsm. & Binn.) Copel.).
3. Lecanopteris luzonensis Hennipman, Blumea 32. --Lecanopteris carnosa auctt. philipp. non (Reinw.) Blume  LUZON: Pangasinan, Pampanga, Bataan (Mt Mariveles), Laguna, Quezon, Camarines Norte. Conservation status: Endangered (DAO 2017-11). Photos
4. Lecanopteris sinuosa (Wall. ex Hook.) Copel. Conservation status: Endangered (DAO 2017-11). Photos

Lemmaphyllum C.Presl

1. Lemmaphyllum carnosum (J.Sm. ex Hook.) C.Presl, Epimel. Bot. (1851) 158; -- Drymoglossum carnosum (Wall.) J.Sm. ex Hook., Gen. Fil. (1841) t.78A. Photos
   • var. carnosum. --?Lemmaphyllum spathulatum (C.Presl) C.Presl, Epimel. Bot. (1851) 158. ?Cambodia, China, India, ?Laos, Myanmar, Thailand, Vietnam, Philippines. LUZON

Lepisorus (J.Sm.) Ching

1. Lepisorus accedens (Blume) Hosok., Trans. Nat. Hist. Soc. Formosa 31 (1941) 477; --Lemmaphyllum accedens (Blume) Donk, Reinwardtia 2 (1954) 409; --Weatherbya accedens (Blume) Copel., Gen. Fil. (1947) 191; --Polypodium accedens Blume, Enum. Pl. Javae 2 (1828) 121. Australia, Bismarck Arch, Borneo, Fiji, Java, Lesser Sunda Isls, Micronesia, Moluccas, New Guinea, Peninsular Malaysia, Samoa, Solomon Isls, Sulawesi, Sumatra, Thailand, Philippines. Photos
   
2. Lepisorus longifolius (Blume) Holttum; --Pleopeltis longifolia (Blume) Bedd.; --Paragramma longifolia (Bl) T.Moore. Photos
3. Lepisorus mucronatus (Fée) Li Wang, Bot. J. Linn. Soc. 162 (2010): 35; --Belvisia mucronata (Fée) Copel.
4. Lepisorus platyrhynchos (Kunze) Li Wang, Bot. J. Linn. Soc. 162 (2010): 35; --Belvisia platyrhynchos (Kunze) Copel. --Belvisia glauca (Copel.) Copel.; --Hymenolepis platyrhynchos Kunze var. glauca Copel.; --Hymenolepis glauca (Copel.) C.Chr. Conservation status: Endangered (DAO 2017-11). Photos
5. Lepisorus spicatus (L.f.) Li Wang, Bot. J. Linn. Soc. 162 (2010): 35; --Belvisia spicata (L.f.) Copel. --Belvisia revoluta (Blume) Copel.
6. Lepisorus thunbergianus (Kaulf.) Ching; --Pleopeltis thunbergiana Kaulf.
7. Lepisorus validinervis (Kunze) Li Wang, Bot. J. Linn. Soc. 162 (2010): 36; --Belvisia validinervis (Kunze) Copel. Photos

Leptochilus Kaulf.

1. Leptochilus axillaris (Cav.) Kaulf., Enum. Filic. (1824) 147; --Acrostichum axillare Cav., Ann. Hist. Nat. 1 (1799) 101. Bangladesh, Borneo, China, India, Java, Lesser Sunda Isls, Moluccas, Myanmar, New Guinea, Nicobar Isl, Peninsular Malaysia, Sulawesi, Sumatra, Thailand, Vietnam, Philippines. PALAWAN. 
2. Leptochilus decurrens Blume, Enum. Pl. Javae 2 (1828) 206; --Anapausia decurrens (Blume) C.Presl, Epimel. Bot. (1851) 186. Bhutan, Bonin Isl, Borneo, ?Cambodia, China, Christmas Isl, India, Japan, Java, Laos, Lesser Sunda Isls, Moluccas, Myanmar, Nepal, New Guinea, Peninsular Malaysia, South Korea, Sri Lanka, Sulawesi, Sumatra, Taiwan, Thailand,  Vietnam, Philippines.
3. Leptochilus ellipticus (Thunb.) Noot., Blumea 42 (1997) 283; --Colysis elliptica (Thunb.) Ching, Bull. Fan Mem. Inst. Biol. 4 (1933) 333; --Polypodium ellipticum Thunb., Fl. Jap. (Thunberg) (1784) 335. Australia, Bhutan, China, Hong Kong, India, Japan, Laos, Myanmar, Nepal, Ryukyu Isls, South Korea, Taiwan, Thailand, Tibet, Vietnam, Philippines. LEYTE, LUZON, MINDANAO, SAMAR. Photos
4. Leptochilus fluviatilis (Lauterb.) Liang Zhang & Li Bing Zhang, Phytotaxa 374 (2018) 173; --Leptochilus macrophyllus var. fluviatilis (Lauterb.) Noot., Blumea 42 (1997) 289; --Polypodium fluviatile Lauterb., Bot. Jahrb. Syst. 44 (1910) 507; --Type: from Borneo. Borneo, Sulawesi, Philippines.
   
5. Leptochilus hemionitideus (C.Presl) Noot., Blumea 42 (1997) 285; --Selliguea hemionitidea C.Presl, Tent. Pterid. (1836) 216. Bangladesh, Bhutan, China, India, Japan, Myanmar, Nepal, Ryukyu Isls, Taiwan, Thailand, Tibet, Vietnam, Philippines. Photos
6. Leptochilus macrophyllus (Blume) Noot., Blumea 42 (1997) 286; --Colysis macrophylla (Blume) C.Presl, Epimel. Bot. (1851) 147; --Grammitis macrophylla, Enum. Pl. Javae 2 (1828) 119. --Colysis bolsteri (Copel.) Copel., Fern Fl. Philip. 3 (1960) 489; --Polypodium bolsteri Copel., PJS 1(Suppl. 4) (1906) 257. Bismarck Arch, Borneo, Java, Lesser Sunda Isls, Moluccas, New Guinea, Nicobar Isls, Peninsular Malaysia, Solomon Isls, Sulawesi, Sumatra, Thailand, Philippines. PALAWAN. Photos
7. Leptochilus minor Fée, Mém. Foug., 2. Hist. Acrostich. (1845) 87; --Dendroglossa minor (Fée) Copel., Gen. Fil. (1947) 199; --Colysis minor (Fée) M.G.Price, Kalikasan 3 (1974) 176. Borneo, India, Peninsular Malaysia, Sulawesi, Sumatra, Thailand, Vietnam, Philippines. LUZON. Photos
8. Leptochilus pedunculatus (Hook. & Grev.) Fraser-Jenk., Taxon. Revis. Indian Subcontinental Pteridophytes (2008) 62; --Leptochilus macrophyllus (Blume) Noot. var. pedunculatus (Hook. & Grev.) Noot., Blumea 42 (1997) 290; --Ceterach pedunculatum Hook. & Grev., Icon. Filic. (1827) t. 5. --Colysis membranacea (Blume) C.Presl, Epimel. Bot. (1851) 147; --Grammitis membranacea Blume, Enum. Pl. Javae 2 (1828) 118. Bangladesh, China, India, Java, Langkawi Isl, Laos, Myanmar, Nepal, Peninsular Malaysia, Sri Lanka, Sumatra, Thailand, Vietnam, Philippines.
   
9. Leptochilus pothifolius (D.Don) Fraser-Jenk., Taiwania 53 (2008) 181; --Colysis pothifolia (D.Don) C.Presl, Epimel. Bot. (1851) 148; --Hemionitis pothifolia D.Don, Prodr. Fl. Nepal. (1825) 13. Bhutan, Bonin Isl, Cambodia, China, Hong Kong, India, Japan, Laos, Myanmar, Nepal, Ryukyu Isls, ?Taiwan, Thailand, Vietnam, Philippines.
   

Loxogramme (Blume) C.Presl

1. Loxogramme antrophyoides (Baker) C.Chr.
2. Loxogramme avenia (Blume) C.Presl --Loxogramme malayana Copel. Photos
3. Loxogramme chinensis Ching. Possible new record for the Philippines.
4. Loxogramme conferta Copel.
5. Loxogramme dimorpha Copel.
6. Loxogramme forbesii Copel.
7. Loxogramme parallela Copel. Photos
8. Loxogramme scolopendrioides (Gaudich.) C.V.Morton. --Loxogramme scolopendrina (Bory) C.Presl Photos
9. Loxogramme subselliguea (Baker) Alston. --Loxogramme major Copel.

Micropolypodium Hayata

1. Micropolypodium pulogense (Copel.) AR Sm.; --Xiphopteris pulogensis (Copel.) Copel.

Microsorum Link

1. Microsorum commutatum (Blume) Copel.; --Phymatosorus commutatus (Blume) Pic.-Serm. --Microsorum phanerophlebium (Copel.) Copel. Photos
2. Microsorum congregatifolium (Alderw.) Holttum. --Microsorum mindanaense (Christ) Copel. Photos
3. Microsorum ensatum (Thunb.) H.Ito. --Neocheiropteris ensata (Thunb.) Ching.
4. Microsorum heterocarpum (Blume) Ching. Photos
5. Microsorum heterolobum (C.Chr.) Copel. Probably a hybrid.
6. Microsorum insigne (Blume) Copel. --Microsorum decurrens (Brackenr.) Copel. --Microsorum palmatum auct. non (Blume) Fee. Photos
7. Microsorum longissimum Fée. --Microsorum sablanianum (Christ) Copel. Photos
8. Microsorum membranifolium (R.Br.) Ching. --Microsorum plukenetii (C.Presl) M.G.Price. --Microsorum alternifolium auct.: Copel., Gen. Fil. (1947), pro parte. --Microsorum longipes Copel., nom. nud. --Microsorum nigrescens (Blume) Copel.; --Phymatosorus nigrescens (Blume) Pic.-Serm. --? Microsorum schneideri (Christ) Copel. Photos
9. Microsorum monstrosum (Copel.) Copel. Photos
10. Microsorum pentaphyllum (Baker) Copel. --Microsorum curranii (Copel.) Copel. Photos
11. Microsorum pteropus (Blume) Copel.
12. Microsorum punctatum (L.) Copel. --Microsorum musifolium (Blume) Copel. --Microsorum validum (Copel.) Ching. Photos
13. Microsorum rubidum (J.Sm.) Copel. Photos
14. Microsorum samarense (J.Sm.) Bosman; --Diblemma samarense J.Sm. --Microsorum tenuilore (J.Sm. ex Mett.) Copel. Photos
15. Microsorum sarawakense (Baker) Ching. --Microsorum rizalense Copel. Conservation status: Vulnerable (DAO 2017-11). Photos
16. Microsorum scolopendria (Burm. f.) Copel.; --Phymatosorus scolopendria (Burm. f.) Pic.-Serm. Tropical Africa to Polynesia and Australia. Throughout the Philippines. Epiphytic or lithophytic in exposed sites and disturbed forests at low elevations, even on coastal bluffs subjected to salt spray. Common. Photos
17. Microsorum zippelii (Blume) Ching. Photos

 Neocheiropteris Christ = Microsorum

Oreogrammitis Copel.

1. Oreogrammitis adspersa (Blume) Parris, Gard. Bull. Sing. 58 (2007) 255; --Grammitis adspersa (Blume) Blume.
2. Oreogrammitis caespitosa (Blume) Parris, Gard. Bull. Sing. 58 (2007) 257; --Grammitis caespitosa Blume.
3. Oreogrammitis congener (Blume) Parris, Gard. Bull. Sing. 58 (2007) 258; --Grammitis congener Blume.
4. Oreogrammitis dolichosora (Copel.) Parris, Gard. Bull. Sing. 58 (2007) 258; --Grammitis dolichosora (Copel.) Copel.
5. Oreogrammitis fasciata (Blume) Parris, Gard. Bull. Sing. 58 (2007) 259; --Grammitis fasciata Blume.
6. Oreogrammitis fenicis (Copel.) Parris, Gard. Bull. Sing. 58 (2007) 259; --Grammitis fenicis Copel.
7. Oreogrammitis knutsfordiana (Baker) Parris, Gard. Bull. Sing. 58 (2007) 262; --Grammitis knutsfordiana (Baker) Copel.
8. Oreogrammitis locellata (Baker) Parris, Gard. Bull. Sing. 58 (2007) 262; --Grammitis locellata (Baker) Copel. --Grammitis stenocrypta Copel.
9. Oreogrammitis marivelesensis (Copel.) Parris, Gard. Bull. Sing. 58 (2007) 263; --Grammitis marivelesensis Copel.
10. Oreogrammitis merrillii (Baker) Parris, Gard. Bull. Sing. 58 (2007) 263; --Grammitis merrillii (Copel.) Copel.
11. Oreogrammitis oncobasis (M.G.Price) Parris, Gard. Bull. Sing. 58 (2007) 264; --Grammitis oncobasis M.G.Price.
12. Oreogrammitis padangensis (Baker) Parris, Gard. Bull. Sing. 58 (2007) 264; --Grammitis padangensis (Baker) Copel.
13. Oreogrammitis pilosiuscula (Blume) Parris.
14. Oreogrammitis reinwardtii (Blume) Parris, Gard. Bull. Sing. 58 (2007) 266; --Grammitis reinwardtii Blume.
15. Oreogrammitis scabristipes (Baker) Parris, Gard. Bull. Sing. 58 (2007) 267; --Grammitis scabristipes (Baker) Copel. --Grammitis bulbotricha (Copel.) Copel. --Grammitis limapes Copel.
16. Oreogrammitis sumatrana (Baker) Parris, Gard. Bull. Sing. 58 (2007) 269; --Grammitis sumatrana (Baker) Copel.
17. Oreogrammitis torricelliana (Brause) Parris, Gard. Bull. Sing. 58 (2007) 270; --Grammitis torricelliana (Brause) Parris. --Grammitis microtricha Copel.
18. Oreogrammitis aff. insularis.
19. Oreogrammitis aff. knutsfordiana.

Photinopteris J.Sm. = Aglaomorpha

 Phymatodes C.Presl = Microsorum

 Phymatopsis J.Sm. = Selliguea

 Phymatopteris Pic.Serm. = Selliguea

 Phymatosorus Pic.Serm. = Microsorum

Platycerium Desv.

1. Platycerium coronarium (König ex Müller) Desv. Tenasserim, Thailand and Malesia. In the Philippines reported from LUZON: Cagayan to MINDANAO: Davao. High canopy epiphyte in lowland primary dipterocarp forests. Also cultivated as an ornamental. Conservation status: Critically Endangered (DAO 2017-11). Photos
2. Platycerium grande (Fée) Kunze. Endemic to the Philippines. MINDANAO: Zamboanga, Lanao, Davao (Mt Apo). Also reported by Copeland (1960) from S LUZON (Mt Cristobal), possibly in error. Majestic crown epiphyte in forest, lowland to 1000m. Also cultivated as an ornamental. Conservation status: Critically Endangered (DAO 2017-11). Photos

 Pleopeltis Humb. & Bonpl. = Lepisorus

Podosorus Holttum

1. Podosorus angustatus Holttum, Kew Bull. 20 (1966) 455, f. 1. A monotypic Philippine endemic genus known only from the type (Gutierrez, H. 61-142) collected  along creek in Dipterocarp forest, Brgy. Disulap, Municipality of San Mariano, Isabela Prov., Luzon. Conservation status: Critically Endangered (DAO 2017-11).

Prosaptia C.Presl

1. Prosaptia alata (Blume) Christ. --Prosaptia pinnatifida C.Presl --Prosaptia toppingii Copel.
2. Prosaptia ancestralis Copel.
3. Prosaptia barathrophylla (Baker) M.G.Price. Philippine material with the following two names are referable to this species: Prosaptia khasyana (Hook.) C.Chr. & Tardieu, Notul. Syst. (Paris) 8 (1939) 180; --Ctenopteris khasyana (Hook.) Holttum. 
4. Prosaptia celebica (Blume) Tagawa & K Iwats., Acta Phytotax. Geobot. 24 (1969) 61; --Ctenopteris celebica (Blume) Copel. --Ctenopteris pseudoarticulata (Copel.) Tagawa. Photos
5. Prosaptia contigua (Forst.) C.Presl Photos
6. Prosaptia cryptocarpa Copel. Photos
7. Prosaptia davalliacea (F Muell. & Baker) Copel. --Prosaptia linearis Copel. Photos
8. Prosaptia micropora (Copel.) Parris. --Ctenopteris microcarpa Copel. Fern Fl. Philip. 3 (1960) 531, anglice, non rite publice.
9.  Prosaptia multicaudata (Copel.) Parris, Kew Bull. 50 (1995) 166; --Ctenopteris multicaudata (Copel.) Copel.
10. Prosaptia negrosensis (Copel.) M.G.Price, Contr. Univ. Michigan Herb. 17 (1990) 276; --Ctenopteris negrosensis (Copel.) Copel. --Ctenopteris spongiosa (Copel.) Copel.
11. Prosaptia nutans (Blume) Mett.; --Ctenopteris nutans (Blume) J.Sm.
12. Prosaptia obliquata (Blume) Mett. not C.Chr. & Tardieu, Notul. Syst. (Paris) 8 (1939) 180; --Ctenopteris obliquata (Blume) Copel. Photos
13. Prosaptia pectinata T.Moore. --Prosaptia leysii (Baker) Ching.
14. Prosaptia urceolaris (Hayata) Copel.
15. Prosaptia venulosa (Blume) M.G.Price, Contr. Univ. Michigan Herb. 16 (1987) 197; --Ctenopteris venulosa (Blume) Kunze. Photos
16. Prosaptia aff. cavisora

Pteropsis Desv. = Pyrrosia

Pyrrosia Mirbel

1. Pyrrosia lanceolata (L.) Farw. --Pyrrosia adnascens (Sw.) Ching. --Pyrrosia varia (Kaulf.) Farwell. India and S China, across Malesia to Polynesia. Throughout the Philippines. In exposed sites, open forests, etc., Epiphytic or lithophytic, usually at low altitude. Photos
2. Pyrrosia longifolia (Burm.f.) C.V.Morton. --Pyrrosia acrostichoides (G Forst.) Ching. Photos
3. Pyrrosia nummularifolia (Sw.) Ching. Photos
4. Pyrrosia piloselloides (L.) M.G.Price; --Drymoglossum piloselloides (L.) Presl; --Pteropsis piloselloides (L.) Desv. --Pteropsis heterophyllum auct. non (L.) Trimen India to New Guinea. Throughout the Philippines. Epiphytic on tree trunks and branches or epilithic on outcrops and boulders along rivers and streams at low altitudes. Common. Photos
5. Pyrrosia porosa (C.Presl) Hovenkamp var. porosa. --Pyrrosia mollis auct. non (Kunze) Ching. Photos
6. Pyrrosia rasamalae (Raciborski) Shing. --Pyrrosia lanuginosa (Giesenh.) Copel. This species has often been named P. floccigera, properly a synonym of P.  albicans, a species not known to occur in the Philippines. Photos
7. Pyrrosia samarensis (C.Presl) Ching. --Pyrrosia philippinensis Copel., nom. superfl. Photos
8. Pyrrosia sphaerotricha (Mett.) Ching. Photos
9. Pyrrosia splendens (C.Presl) Ching. Conservation status: Vulnerable (DAO 2017-11). Photos

Radiogrammitis Parris

1. Radiogrammitis alepidota (M.G.Price) Parris, Gard. Bull. Sing. 58 (2007) 241; --Grammitis alepidota M.G.Price, Philip. Agriculturist 57 (1973) 34, Fig. 1;  Taiwan (BJ Wang 17447-B, HAST) and Philippines. LUZON (Ilocos Norte, Mt Burnay; Cagayan, Mt Cetaceo; Quezon, Mt Banahaw, type locality), PALAWAN (Mt Mantalingahan). Photos
2. Radiogrammitis jagoriana (Mett. ex Kuhn) Parris, Gard. Bull. Sing. 58 (2007) 242; --Grammitis jagoriana (Mett.) Copel.
3. Radiogrammitis setigera (Blume) Parris. --Grammitis intromissa (Christ) Parris. --Grammitis fasciculata Blume, nom. illegit. pro Polypodium setigerum Blume. Photos
4. Radiogrammitis subpinnatifida (Blume) Parris; --Xiphopteris subpinnatifida (Blume) Copel.

 Schellolepis Pic.-Serm. = Goniophlebium

Scleroglossum Alderw.

1. Scleroglossum pusillum (Blume) Alderw. --Scleroglossum minus (Fée) C.Chr. Photos
2. Scleroglossum sulcatum (Kuhn) Alderw. --Scleroglossum intermedium (Copel.) C.Chr. --Scleroglossum pusillum Copel., non (Blume) Alderw.

Selliguea Bory

1. Selliguea albidosquamata (Blume) Parris; --Crypsinus albidosquamatus (Blume) Copel. Photos
2. Selliguea caudiformis (Blume) Carruth. --Selliguea calophlebia (Copel.) Copel. --Selliguea feei auct. non Bory p.p.
3. Selliguea elmeri (Copel.) Ching; --Selliguea feei Bory var. elmeri (Copel.) Tagawa. --Selliguea feei auct. non Bory p.p. Photos
4. Selliguea enervis (Cav.) Ching; --Crypsinus enervis (Cav.) Copel. Photos
5. Selliguea glauca (J.Sm. ex T.Moore) Hovenkamp; --Crypsinus glaucus (J.Sm. ex T.Moore) Copel. Photos
6. Selliguea heterocarpa (Blume) Blume.
7. Selliguea lagunensis (Christ) Hovenkamp; --Crypsinus lagunensis (Christ) Copel.; --Phymatopteris lagunensis (Christ) Pic.-Serm. Photos
8. Selliguea lateritia (Baker) Hovenkamp.
9. Selliguea oodes (Kunze) Hovenkamp; --Crypsinus oodes (Kunze) Copel. Photos
10. Selliguea pyrolifolia (Goldm.) Hovenkamp; --Crypsinus pyrolifolius (Goldm.) Copel. Photos
11. Selliguea stenophylla (Blume) Parris; --Crypsinus stenophyllus (Blume) Copel. Photos
12. Selliguea taeniata (Sw.) Parris; --Crypsinus taeniatus (Sw.) Copel., including var. palmatus C.Chr.; --Phymatopteris taeniata (Sw.) Pic.-Serm. --Crypsinus ramosii Copel. Photos
13. Selliguea triloba (Houtt.) M.G.Price; --Crypsinus trilobus (Houtt.) Copel.; --Phymatopteris triloba (Houtt.) Pic.-Serm. --Selliguea matutumensis Copel.
14. Selliguea whitfordii (Copel.) Hovenkamp; --Crypsinus whitfordii (Copel.) Copel. --Crypsinus subdimorphus Copel. Photos

Doubtful occurrence in the Philippines:

1. Selliguea hastata (Thunb.) Copel.; --Crypsinus hastatus (Thunb.) Copel.;  --Phymatopteris hastata (Thunb.) Pic.-Serm. India, China, Japan. In the Philippines known only from LUZON: Benguet. Epiphytic or lithophytic, alt. c. 2400m. According to Hovenkamp et.al. [Fl. Males. ser. 2, 3 (1998) 227], simple fronded plants from the Philippines have been identified as Phymatopteris hastata by Zamora & Co [Guide to Philipp. Flora & Fauna 2 (1986) 29], and that other than through this reference, P. hastata is not known from the Philippines.  This is incorrect as Copeland (Fern Fl. Philipp. 2 (1960) 505) reported this species to be common in Benguet at 2400 m.

Themelium (T.Moore) Parris

1. Themelium cf. blechnifrons (Hayata) Parris. --Ctenopteris curtisii (Baker) Copel. Photos
2. Themelium halconense (Baker) Parris, Kew Bull. 59 (2004) 225; --Ctenopteris halconensis (Copel.) Copel.
3. Themelium loherianum (Christ) Parris, Kew Bull. 59 (2004) 225; --Grammitis loheriana (Christ) Copel.
4. Themelium taxodioides (Baker) Parris, Kew Bull. 52 (1997) 739; --Ctenopteris taxodioides (Baker) Copel.
5. Themelium tenuisectum (Blume) Parris, Kew Bull. 52 (1997) 740; --Ctenopteris tenuisecta (Blume) J.Sm. Photos
6. Themelium yoderi (Copel.) Parris, Kew Bull. 52 (1997) 739; --Ctenopteris yoderi (Copel.) Copel.
7. Themelium aff. tenuisectum sp. 1.
8. Themelium aff. tenuisectum sp. 2.

Thylacopteris Kunze ex J.Sm.

1. Thylacopteris papillosa (Blume) J.Sm. Photos

Tomophyllum (E.Fourn.) Parris

1. Tomophyllum macrum (Copel.) Parris, Gard. Bull. Sing. 58 (2007) 247; --Ctenopteris macra (Copel.) Copel.
2. Tomophyllum millefolium (Blume) Parris, Gard. Bull. Sing. 58 (2007) 247; --Ctenopteris millefolia (Blume) Copel.
3. Tomophyllum minutum (Blume) Parris, Gard. Bull. Sing. 58 (2007) 247; --Ctenopteris minuta (Blume) Holttum Possibly conspecific with C. macra. It is more likely to be a misidentification of Tomophyllum macrum.  Blume’s name has priority.
4. Tomophyllum repandulum (Mett.) Parris.
5. Tomophyllum subsecundodissectum (Zoll.) Parris, Gard. Bull. Sing. 58 (2007) 249; --Ctenopteris subsecundodissecta (Zoll.) Copel. Photos

Psilotaceae

Psilotum L.

1. Psilotum complanatum Sw., Syn. Fil. (1806) 188, 414, t. 4, Fig. 5. Conservation status: Endangered (DAO 2017-11). Photos
2. Psilotum nudum (L.) Griseb., Prodr. Aethéogam. (1805) 112. Conservation status: Vulnerable (DAO 2017-11). Photos.  
3. Psilotum triquetrum Sw., Schrad. J. Bot. 1800 (2) 1801.

Tmesipteris Bernh.

1. Tmesipteris zamorarum Gruezo & Amoroso, Asia Life Sci. 21 (2012) 656 (as "zamorae"); --Type: WS Gruezo WM 14493 (CAPH, holo; WGRZ, iso), Luzon: Quezon, Mt. Banahaw, 19-Mar-1982, 1200m. --Tmesipteris lanceolata non Dang, sensu Bierhorst, J. Linn. Soc. Bot. 67, Suppl. 1 (1973) 46. --Tmesipteris tannensis auctt. non (Spreng.) Bernh. Endemic to the Philippines. LUZON: Benguet (Mt Pulog), Mountain Province (Mt Data), Quezon (Mt Banahaw), MINDANAO: Bukidnon (Mt Kitanglad), Cotabato-Davao (Mt Apo), Davao (Mt McKinley), Misamis Occidental (Mt Malindang), South Cotabato (Mt Matutum), MINDORO: Mt Halcon. 1000-2900m. Conservation status: Endangered (DAO 2017-11). Photos

Pteridaceae 

Including Acrostichaceae, Adiantaceae, Antrophyaceae, Ceratopteridaceae, Cheilanthaceae, Hemionitidaceae, Parkeriaceae, Sinopteridaceae, Taenitidaceae, Vittariaceae

Acrostichum L.

1. Acrostichum aureum L. Photos
2. Acrostichum speciosum Willd. Photos

Adiantum L.

1. Adiantum capillus-veneris L. Photos
2. Adiantum caudatum L. Photos
3. Adiantum cupreum Copel. Conservation status: Vulnerable (DAO 2017-11).
4. Adiantum edgeworthii Hook. 
5. Adiantum flabellulatum L.
6. Adiantum hispidulum Sw. Photos  
7. Adiantum hosei Baker, J. Bot. 26 (1888) 324. --Adiantum scabripes Copel., Philipp. J. Sci., C 7 (1912) 55, t. 5. Conservation status: Vulnerable (DAO 2017-11 as Adiantum monosorum Baker). Photos  
8. Adiantum malesianum Ghatak. Photos
9. Adiantum mindanaoense Copel. Conservation status: Endangered (DAO 2017-11). Photos
10. Adiantum philippense L. Photos  
11. Adiantum setulosum J.Sm. --Adiantum diaphanum sensu Copel. non Blume, Enum. Pl. Jav. (1828) 215. Photos
12. Adiantum soboliferum Wall. ex Hook., Sp. Fil. 2 (1851) 13. Africa, China, Taiwan, Myanmar, Thailand, Laos, ?Cambodia, India, Vietnam, Peninsular Malaysia, Lesser Sunda Isls, Madagascar, Philippines. LUZON, PALAWAN. Photos
13. Adiantum stenochlamys Baker. --Adiantum opacum Copel., PJS 1 (1906) Suppl. 255, t. 3. PALAWAN. Conservation status: Vulnerable (DAO 2017-11). Photos
14. Adiantum tenerum Sw., Prodr. [O. P. Swartz] (1788) 135. Introduced from South America. Photos

Aleuritopteris Fée

1. Aleuritopteris albofusca (Baker) Pic.-Serm. --Cheilanthes anceps Blanf. --Cheilanthes albofusca Baker. Photos
2. Aleuritopteris farinosa (Forssk.) Fée; --Cheilanthes farinosa (Forssk.) Kaulf. --Cheilanthes agetae (Saiki) Kuo
3. Aleuritopteris formosana (Hayata) Tagawa, Acta Phytotax. Geobot. 14 (1952) 191
   
4. Aleuritopteris rufa (D.Don) Copel.; --Cheilanthes rufa D.Don. --Cheilanthes dubia Hope. Photos

Antrophyum Kaulf.

1. Antrophyum immersum (Bory ex Willd.) Mett. --Antrophyum nanum Fée. Photos
2. Antrophyum latifolium Blume. Photos  
3. Antrophyum ledermannii Hieron.
4. Antrophyum plantagineum (Cav.) Kaulf. Photos
5. Antrophyum reticulatum (Forst.) Kaulf. --Antrophyum alatum Brackenr. --Antrophyum callifolium Blume. --Antrophyum clementis Christ. Photos
6. Antrophyum semicostatum Blume. Photos
7. Antrophyum sessilifolium (Cav.) Sprengel. Photos
8. Antrophyum williamsii Benedict. Photos

Calciphilopteris Yesilyurt & H.Schneid.

1. Calciphilopteris wallichii (J.Sm.) Yesilyurt & H.Scnheid., Phytotaxa 7 (2010) 57; --Doryopteris wallichii J.Sm., Journal of Botany 4 (1841) 404; --Type: Cuming 238 (K, holo; BM, C, G, L, iso). Endemic to Philippines. LUZON. Note by J.F. Barcelona (15-Aug-14): Philippine specimens under Doryopteris ludens auct. non. (Wall. ex Hook) J.Sm. refer to this species. Photos 

Ceratopteris Brongn.

1. Ceratopteris oblongiloba Masuyama & Watano, Acta Phytotax. Geobot. 61 (2010) 84; --Type from Indonesia. Cambodia, Thailand, Java, Sumatra, Philippines. LUZON.
2. Ceratopteris thalictroides (L.) Brongn.; --Acrostichum thalictroides L., Sp. Pl. 2(1753) 1070. --Ceratopteris siliquosa (L.) Copel. Asia, Oceania, Central America. LUZON, MINDANAO. Conservation status: Endangered (DAO 2017-11). Photos  

Cheilanthes Sw.

1. Cheilanthes belangeri (Bory) C.Chr.
2. Cheilanthes javensis (Willd.) T.Moore. --Cheilanthes densa (J.Sm.) Fée. --Cheilanthes dilimanensis PM Zamora
3. Cheilanthes mysurensis Wall. ex Hook.
4. Cheilanthes tenuifolia (NL Burm.) Sw., Syn. Fil. (1806) 129, 332. Photos

Coniogramme Fée

1. Coniogramme fraxinea (D.Don) Diels in Engl. & Prantl, Nat. Pflanzenfam. 1 (1899) 262; 
2. Coniogramme macrophylla (Blume) Hieron., Hedwigia 57 (1916) 291. Photos
3. Coniogramme serrulata Fée -- Mém. Foug., 5. Gen. Filic. 167 t. 14 B. f. 2. (1850). Photos
4. Coniogramme subcordata Copel.

Copelandiopteris Stone = Pteris

Doryopteris J.Sm.

1. Doryopteris concolor (Langsdorff & Fischer) Kuhn.; --Cheilanthes concolor (Langsdorff & Fischer) R & A Tryon. --Doryopteris cuspidata Copel. Conservation status: Endangered (DAO 2017-11). Photos

Gymnopteris Bernh. [= Hemionitis]

1. Gymnopteris vestita (Wall.) Underw.

Haplopteris C.Presl

Edited by Julie F. Barcelona, 16 January 2013

Photos

1. Haplopteris alternans (Copel.) Barcelona, ined.; --Vittaria alternans Copel. --Vittaria hecistophylla Copel. Photos
2. Haplopteris anguste-elongata (Hayata) E.H. Crane; --Vittaria anguste-elongata Hayata. --Vittaria stenophylla Copel.
3. Haplopteris crispomarginata (Christ) Barcelona, ined.; --Vittaria crispomarginata Christ. --Vittaria merrillii Christ. Photos
4. Haplopteris elongata (Sw.) E.H. Crane; --Vittaria elongata Sw. --Vittaria intermedia Blume. --Vittaria philippinensis Christ
5. Haplopteris ensiformis (Sw.) E.H. Crane; --Vittaria ensiformis Sw. --Vittaria incurvata Cav. --Vittaria lineata auct. non Smith. --Vittaria pachystemma Christ. Photos
6. Haplopteris scolopendrina (Bory) E.H. Crane; --Vittaria scolopendrina (Bory) Thwaites. --Vittaria subcoriacea Christ. Photos
7. Haplopteris taeniophylla (Copel.) E.H. Crane; --Vittaria taeniophylla Copel.
8. Haplopteris zosterifolia (Willd.) E.H. Crane; --Vittaria zosterifolia Willd. --Vittaria copelandii Alderw.

Monogramma Comm. ex Schkuhr.

1. Monogramma dareicarpa Hook. --Monogramma capillaris Copel., PJS 6c (1911) Bot. 147. Photos

Onychium Kaulf.

1. Onychium japonicum (Thunb.) Kunze. Photos  
2. Onychium siliculosum (Desv.) C.Chr. Photos

Parahemionitis Panigrahi

Edited by Julie F. Barcelona & Pieter B. Pelser, 25 May 2012

1. Parahemionitis cordata (Roxb. ex Hook. & Grev.) Fraser-Jenk., New Sp. Syndr. Indian Pteridol. (1997) 187. --Parahemionitis arifolia (Burm. f.) Panigr.; --Hemionitis arifolia (Burm. f.) T.Moore. Photos  

Pityrogramma Link

1. Pityrogramma calomelanos (L.) Link, Handb. Gewachs. 3 (1833) 20. Photos

Pteris L.

1. Pteris armata C.Presl Photos
2. Pteris biaurita L. Photos  
3. Pteris blumeana J Agardh. Photos  
4. Pteris brevis Copel. Conservation status: Vulnerable (DAO 2017-11).
5. Pteris calocarpa (Copel.) M.G.Price; --Schizostegeopsis calocarpa Copel.; --Copelandiopteris calocarpa (Copel.) Stone. Conservation status: Critically Endangered (DAO 2017-11).
6. Pteris cretica L. --Pteris stenophylla sensu Copel., FFP 1 (1958) 129, non Hook. & Grev. Photos  
7. Pteris dataensis Copel. Conservation status: Vulnerable (DAO 2017-11).
8. Pteris distans J.Sm. ex Hook. Possibly a hybrid or a bizarre freak.
9. Pteris edanoi Copel., PJS 81 (1952) 8, 'edanyoi'.
10. Pteris elmeri Christ.
11. Pteris endoneura M.G.Price; --Copelandiopteris endoneura (M.G.Price) Stone. Conservation status: Endangered (DAO 2017-11). Photos
12. Pteris ensiformis Burm. Photos
    • var. ensiformis. --Pteris mutilata L.
    • var. victoriae Hort.
13. Pteris fauriei Hieron., Hedwigia 55 (1914) 345; --Hatusima, Mem. Fac. Kagoshima Univ. 5 (1966) 20.
14. Pteris glaucovirens Goldm. 
15. Pteris heteromorpha Fée. Photos  
16. Pteris loheri Copel.
17. Pteris longipinnula Wall. ex J Agardh. Photos
18. Pteris luzonensis Hieron.
19. Pteris macgregorii Copel. Conservation status: Vulnerable (DAO 2017-11).
20. Pteris melanocaulon Fée. Photos
21. Pteris melanorhachis Copel. 
22. Pteris mertensioides Willd. --Pteris gardneri (Fée) Hook. --Pteris excelsa Gaudich. Photos  
23. Pteris micracantha Copel. Conservation status: Vulnerable (DAO 2017-11).
24. Pteris mucronulata Copel. Conservation status: Vulnerable (DAO 2017-11).
25. Pteris opaca (C.Presl) J.Sm. ex Fée. --Pteris taenitis Copel. Conservation status: Vulnerable (DAO 2017-11 as Pteris taenitis Copel.). Photos
26. Pteris oppositipinnata Fée. --Pteris asperula J.Sm. --Pteris caesia Copel. Photos  
27. Pteris pachysora (Copel.) M.G.Price; --Schizostegeopsis pachysora Copel.; --Copelandiopteris pachysora (Copel.) Stone. Conservation status: Critically Endangered (DAO 2017-11).
28. Pteris pacifica Hieron. --Pteris cumingii Hieron. Photos
29. Pteris pedicellata Copel. 
30. Pteris pellucida C.Presl Photos
31. Pteris philippinensis Fée.; --Pteris longipes D.Don var. philippinensis (Fée) Christ. --Pteris longipes D.Don, Prod. Fl. Nepal. (1825) 15; --Copel., FFP 1 (1958) 133. Photos  
32. Pteris purpureorhachis Copel. 
33. Pteris ramosii Copel. Possibly a variant of P. philippinensis. Conservation status: Vulnerable (DAO 2017-11).
34. Pteris rangiferina C.Presl
35. Pteris rigidula Copel., FFP 1 (1958) 137. Photos
36. Pteris ryukyuensis Tagawa, Acta Phytotax. Geobot. 4 (1935) 204; --Hatusima, Mem. Fac. Kagoshima Univ. 5 (1966) 20. --?Pteris cretica sensu Copel., FFP 1 (1958) 129, quoad pl. ex Batanes Islands, non L. Photos
37. Pteris schlechteri Brause. Photos  
38. Pteris semipinnata L. Photos  
39. Pteris spinescens C.Presl --Pteris setuloso-costulata Hayata.
40. Pteris squamipes Copel. Conservation status: Vulnerable (DAO 2017-11).
41. Pteris tripartita Sw. --Pteris microdictyon (Fée) Hook. Photos  
42. Pteris vittata L. Photos  
43. Pteris wallichiana J Agardh Confirmed to be in Luzon: Ilocos Norte prov., Mt. Burnay (Iwatsuki & Price, 1975).
44. Pteris whitfordii Copel. Photos

Schizoloma Gaudich. = Taenitis

Syngramma J.Sm.

1. Syngramma alismifolia (C.Presl) J.Sm., Lond. J. Bot. 4 (1845) 168, t. 7-8, Fig. B. Photos  
2. Syngramma vittiformis J.Sm. Photos
3. Syngramma wallichii (Hook.) Bedd. Photos

Taenitis Willd.

1. Taenitis blechnoides (Willd.) Sw., Syn. Fil. 24 (1806) 220. Photos  
2. Taenitis luzonica (Alderw.) Holttum; --Syngramma luzonica Alderw. Photos
3. Taenitis cordata (Gaudich.) Holttum; --Schizoloma cordatum Gaudich. Conservation status: Vulnerable (DAO 2017-11). Photos

Vaginularia Fée

Last edited by Julie F. Barcelona & Cheng-Wei Chen, 5 January 2019

Photos 

1. Vaginularia junghuhnii Mett., Fil. Hort. Bot. Lips. (1856) 25. --Vaginularia paradoxa auct. non (Fée) Mett. --Monogramma paradoxa auct. non (Fée) Mett. Borneo, Lesser Sunda Isls, Moluccas, New Guinea, Sri Lanka, Sulawesi, Sumatra, Thailand, Taiwan, Philippines. LUZON: Camarines Sur (photos), Quezon (photos), MINDANAO: North Cotabato (photos), NEGROS (photos). Photos
2. Vaginularia trichoidea (J.Sm.) Fée; --Monogramma trichoidea J.Sm. ex Hook. Photos

Vittaria Sm. = Haplopteris p.p.

Rhachidosoraceae

Edited by Julie F. Barcelona, 1 September 2013

Rhachidosorus Ching

No photos

1. Rhachidosorus stramineus (Copel.) Ching, Acta Phytotax. Sin. 9 (1964) 74; --Athyrium stramineum Copel., Philipp. J. Sci. 3(5) (1908) 292.

Saccolomataceae

Orthiopteris Copel.

Edited by Pieter B. Pelser, 22 July 2015

Photos

1. Orthiopteris campylura (Kunze) Copel., Fern Fl. Philip. 1 (1958) 87; --Davallia campylura Kunze, Bot. Zeitung (Berlin) 8 (1850) 132; --Saccoloma campylurum (Kunze) Mett. Ann. Sci. Nat. Bot. sér. 4, 15 (1861) 80; --Davallia inaequalis var. minor Hook., Sp. Fil. 1 (1846) 180; --Type: Cuming 119 (K, lecto; K, MICH, isolecto), Luzon. --Saccoloma moluccanum (Blume) Mett. in Kuhn var. stenolobum Christ, Bull. Herb. Boissier, ser. 2 6 (1906) 1005; --Type: Loher s.n. (P, holo), Luzon: Mabacal/Rizal, Mar-1906. Borneo, Fiji, Java, Moluccas, New Guinea, Peninsular Malaysia, Sumatra, Vanuatu, Philippines. LEYTE, LUZON, MINDANAO, NEGROS. Photos
   
   • var. campylura. Moluccas, Papua New Guinea, Philippines. LEYTE, LUZON, MINDANAO, NEGROS.

Saccoloma Kaulf. = Orthiopteris

Salviniaceae (including Azollaceae)

Azolla Lam.

1. Azolla pinnata R.Br. var. imbricata (Roxb. ex Griff.) Bonap. --Azolla imbricata (Roxb. ex Griff.) Nakai Photos
2. Azolla rubra R.Br., Prodr. Fl. Nov. Holland. 167 (1810) Bak. Hdb. 137. Photos

Salvinia Seguier [Not native]

1. Salvinia molesta D.Mitch., Brit. Fern Gaz. 10 (1972) 251. Invasive species (CABI 2017). Photos

Schizaeaceae

Schizaea Sm.

1. Schizaea dichotoma (L.) Sm., Mém. Acad. Turin 5 (1793) 422; --Copel., FFP 1 (1958) 32. Photos  
2. Schizaea digitata (L.) Sw., Syn. Fil. (1806) 150, 380, t. 4, Fig. 1; --Copel., FFP 1 (1958) 32; --Actinostachys digitata (L.) Wall. ex Brackenr. Photos  
3. Schizaea inopinata Selling, Svensk. Bot. Tidskr. 40 (1946) 274, Fig. 1-7; --Copel., FFP 1 (1958) 32; --Actinostachys inopinata (Selling) Reed. Conservation status: Endangered (DAO 2017-11). Photos
4. Schizaea malaccana Baker. Conservation status: Endangered (DAO 2017-11).

Tectariaceae

Aenigmopteris Holttum

1. Aenigmopteris dubia (Copel.) Holttum; --Ctenitis dubia (Copel.) Copel. LUZON: Quezon (Mt Banahaw de Lucban), PANAY: Iloilo prov. (Callado, photos), NEGROS: Negros Oriental (Cuernos Mtns), LEYTE, CAMIGUIN. Terrestrial, altitude 850-1200m. Photos
2. Aenigmopteris mindanaensis Holttum. Endemic to the Philippines. Conservation status: Vulnerable (DAO 2017-11).

Ctenitis (C.Chr.) C.Chr.

1. Ctenitis aciculata (Baker) Ching.
2. Ctenitis atrorubens Holttum. --Ctenitis vilis sensu Copel. (1960), pro parte, non (Kunze) Ching.
3. Ctenitis boholensis Holttum.
4. Ctenitis bulusanica Holttum.
5. Ctenitis decurrentipinnata (Ching) Ching.
6. Ctenitis erythradenia Holttum. --Ctenitis vilis sensu Copel. (1960), pro parte, non (Kunze) Ching.
7. Ctenitis humilis Holttum. Photos
8. Ctenitis paleolata Copel. LUZON: Benguet (Pauai), altitude c. 2400m. Known only from the type collection. Conservation status: Critically Endangered (DAO 2017-11).
9. Ctenitis pallens (Brackenr.) M.G.Price. --Ctenitis rizalensis (Christ) Copel. As C. rizalensis: LUZON: Rizal, Laguna (Mt Makiling), SAMAR, MINDANAO: Zamboanga (San Ramon), Davao (Mt Apo). Terrestrial under moist forest shade,  altitude 400-1200m.
10. Ctenitis propinqua (J.Sm. ex C.Presl) Copel. LUZON: Pampanga (Mt Arayat), Laguna (Mt Makiling), Quezon (Mt Banahaw), Batangas (Mt Makulod).
11. Ctenitis setosa (C.Presl) Holttum. --Ctenitis vilis sensu Copel. (1960), pro parte, non (Kunze) Ching.
12. Ctenitis silvatica Holttum. --Ctenitis vilis sensu Copel. (1960), pro parte, non (Kunze) Ching. Photos
13. Ctenitis subglandulosa (Hance) Ching. --Ctenitis rhodolepis (Clarke) Ching. Photos
14. Ctenitis subobscura (Christ) Holttum.

Excluded

1. Ctenitis subconnexa (Christ) Holttum var. subconnexa and var. alstonii Holttum. A mislocalization to the Philippines by Holttum, Fl. Males. ser. 2, 2 (1991) 28. The locality Batjan is in Halmahera (of the Moluccas).

Heterogonium C.Presl = Tectaria

Pleocnemia C.Presl

1. Pleocnemia brongniartii (Bory) Holttum; --Arcypteris brongniartii (Bory) Holttum. --Tectaria irregularis sensu Copel. (1960), pro parte. Photos
2. Pleocnemia conjugata (Blume) C.Presl
3. Pleocnemia cumingiana C.Presl --Tectaria leuzeana sensu Copel. (1960), pro parte. Photos
4. Pleocnemia irregularis (C.Presl) Holttum; --Arcypteris irregularis (C.Presl) Ching; --Tectaria irregularis sensu Copel. (1960), pro parte. Photos
5. Pleocnemia leuzeana (Gaudich.) C.Presl Photos
6. Pleocnemia macrodonta (Fee) Holttum; --Arcypteris macrodonta (C.Presl) Holttum. --Tectaria irregularis sensu Copel. (1960), pro parte. Photos
7. Pleocnemia presliana Holttum. LUZON: Cagayan, Aurora, Laguna, PALAWAN (Mt Gantung; Imolnod), LEYTE (Palo), MINDANAO: Davao (Mt Apo, Todaya), Surigao del Norte. Terrestrial, usually in partly shaded area, near sea-level to at least 400m. Photos

Psomiocarpa C.Presl

1. Psomiocarpa apiifolia C.Presl LUZON: Cagayan, Isabela, Bataan, Rizal, Laguna (Mt Makiling), Quezon, POLILLO, MASBATE, LEYTE, SAMAR, MINDANAO: Zamboanga (San Ramon). Terrestrial in forest, from near sea-level to c. 450m. Conservation status: Endangered (DAO 2017-11). Photos

Pteridrys C.Chr. & Ching

1. Pteridrys microthecia (Fee) C.Chr. & Ching. Photos
2. Pteridrys syrmatica (Willd.) C.Chr. & Ching. Photos

Stenosemia C.Presl  = Tectaria, Heterogonium

Tectaria Cav.

1. Tectaria adenophora Copel. SIBUYAN. Conservation status: Other Threatened Species (DAO 2017-11).
2. Tectaria angulata (Willd.) Copel. --Tectaria trifolia (Alderw.) C.Chr. Photos
3. Tectaria aspidioides (C.Presl) Copel.; --Heterogonium aspidioides C.Presl Photos
4. Tectaria athyriosora M.G.Price. BOHOL, SAMAR, MINDANAO: Surigao del Norte. Photos
5. Tectaria aurita (Sw.) S. Chandra; --Stenosemia aurita (Sw.) Presl. Photos 
6. Tectaria barberi (Hook.) Copel.
7. Tectaria beccariana (Cesati) C.Chr.
8. Tectaria calcarea (C.Presl) Copel. POLILLO, in wet forest, altitude 50m, BOHOL, LEYTE. Photos
9. Tectaria christii Copel. LUZON: Mountain Province, Benguet, MINDORO: Mindoro Oriental (Mt Halcon). Usually on limestone, c. 1500m. Photos
10. Tectaria crenata Cav. --Tectaria grandifolia sensu Copel. (1960), pro parte. Photos
11. Tectaria decurrens (C.Presl) Copel. --Tectaria peralata Copel. Photos
12. Tectaria devexa (Kunze ex Mett.) Copel. var. devexa. Photos
13. Tectaria dissecta (G Forst.) Lellinger; --Ctenitis dissecta (G Forst.) Copel. Photos
14. Tectaria dolichosora Copel. LUZON: Cagayan, Quezon (Infanta, Tignoan). Steep slopes near summits.
15. Tectaria griffithii (Baker) C.Chr. var. griffithii. --Tectaria malayensis (Christ) Copel. Photos
16. Tectaria hilocarpa (Fée) M.G.Price. --Hemigramma latifolia (Meyen ex Goldm.) Copel., excl. Hemionitis zollingeri and other latter names based on this. Photos
17. Tectaria laxa (Copel.) M.G.Price; --Ctenitis laxa Copel. LUZON: Ilocos Norte (Mt Quebrada; Mt Siminublan). Altitude 80-1100m. Photos
18. Tectaria lobbii (Hook.) Copel., PJS 10 (1915) 146; --Aspidium lobbii Hook., Sp. Fil. 4 (1862) 59. Borneo, Philippines. BOHOL, MINDANAO: Surigao del Sur. Conservation status: Endangered (DAO 2017-11). Photos
    
    • var. lobbii.
19. Tectaria manilensis (C.Presl) Holttum var. manilensis; --Lastreopsis manilensis (C.Presl) M.G.Price; --Ctenitis manilensis (C.Presl) Holttum. --Ctenitis loheri (Christ) Copel.
20. Tectaria melanocaula (Blume) Copel. Tectaria ‘melanocaulis’ in Copel. (1960).
21. Tectaria menyanthidis (C.Presl) Copel.
22. Tectaria mesodon (Copel.) M.G.Price; --Ctenitis mesodon (Copel.) Copel.; --Dryopteris mesodon Copel. LUZON: Ilocos Norte, MINDANAO: Agusan del Norte (Mt Hilong-hilong), 120m.
23. Tectaria nom. nov. ined.; --Heterogonium pinnatum (Copel.) Holttum; --Stenosemia pinnata Copel. A new name might be needed for this species in Tectaria, because Tectaria pinnata is occupied (T. pinnata (C.Chr.) R.M.Tryon & A.F.Tryon, based on Camptodium pinnatum C.CHr.). Photos
24. Tectaria polymorpha (Wall. ex Hook.) Copel.
25. Tectaria profereoides (C.Chr.) S.Y.Dong; --Heterogonium profereoides (C.Chr.) Copel.; --Aspidium profereoides C.Chr. --Heterogonium aspidioides sensu Holttum, Kalikasan 4 (1975) 224, pro parte
26. Tectaria ramosii (Copel.) Holttum; --Ctenitis ramosii Copel. As C. ramosii: BOHOL (Dimiao), SAMAR (Catubig River). Possibly near sea-level to c. 300m. Photos
27. Tectaria repanda (Willd.) Holttum. --Tectaria crenata sensu Copel. (1960), pro parte.
28. Tectaria sagenioides (Mett.) Christenh.; --Heterogonium sagenioides (Mett.) Holttum; --Ctenitis sagenioides (Mett.) Copel. --Ctenitis obscura (Hook.) Copel.; --Heterogonium obscurum (Hook.) Holttum.
29. Tectaria samariana S.Y.Dong; --Heterogonium lobulatum Holttum.
30. Tectaria siifolia (Willd.) Copel. Photos
31. Tectaria stalactica M.G.Price. Conservation status: Endangered (DAO 2017-11).
32. Tectaria sulitii Copel. --Tectaria loheri Copel. --Tectaria grandifolia sensu Copel. (1960), pro parte According to Holttum, Fl. Males. ser. 2, 2 (1991) 91, there may be intermediates between this species and T. crenata. As T. loheri: LUZON: Rizal (Montalban; Sumag). Photos
33. Tectaria suluensis Holttum.
34. Tectaria tabonensis M.G.Price. Conservation status: Other Threatened Species (DAO 2017-11).
35. Tectaria taccifolia (Fée) M.G.Price; --Hemigramma taccifolia (Fée) Copel. Photos
36. Tectaria trichotoma (Fée) Tagawa. --Tectaria balabacensis (Christ) M.G.Price; --Ctenitis balabacensis (Christ) Copel. As Ct. balabacensis: LUZON: Quezon (Guinayangan), PALAWAN, BALABAC, SIBUYAN. Petrophytic along rivers and streams in forests, altitude 100-200m in Palawan. Photos
37. Tectaria trifida (Fée) M.G.Price. --Tectaria irrigua (J.Sm. ex Presl) Copel.
38. Tectaria wenzelii (Copel.) S.Y.Dong; --Heterogonium wenzelii (Copel.) Holttum; --Ctenitis wenzelii Copel. LEYTE. Conservation status: Critically Endangered (DAO 2017-11). Photos

1. Tectaria ramosii X Psomiocarpa apiifolia C.Presl
2. Tectaria ramosii X Tectaria aurita (Sw.) S. Chandra.
3. Tectaria repanda X Tectaria hilocarpa (Fée) M.G.Price.

Tectaridium Copel.

1. Tectaridium macleanii Copel. --Tectaridium primitivum Copel. Endemic to Philippines. Conservation status: Endangered (DAO 2017-11). Photos

Thelypteridaceae

Ampelopteris Kunze

1. Ampelopteris prolifera (Retz.) Copel. Tropical and subtropical regions of the Old World. In the Philippines recorded from LUZON: Abra, Rizal, Laguna, MINDANAO: Davao del Norte (Maco, Lake Leonard), Agusan del Sur (Agusan Marsh). Terrestrial, often forming mats or tangles in swampy sites, lake margins or river embankments. Low altitudes. Photos  

Amphineuron Holttum

1. Amphineuron attenuatum (Kuntze) Holttum. --Cyclosorus alatellus sensu Copel. (1960), excl. syn. Lastrea erubescens sensu Copel. (1960).
2. Amphineuron immersum (Blume) Holttum.; --Lastrea immersa (Blume) T.Moore. --Lastrea verrucosa C.Presl 
3. Amphineuron opulentum (Kaulf.) Holttum. --Cyclosorus extensus (Blume) Ching.
4. Amphineuron terminans (Hook.) Holttum. --Cyclosorus interruptus sensu Copel. (1960), non (Willd.) H.Ito Sri Lanka to Polynesia. Throughout the Philippines in the more seasonal regions. Terrestrial, common in thickets on hillsides and in valleys at low and medium altitudes. Photos

Chingia Holttum

1. Chingia christii (Copel.) Holttum.; --Cyclosorus christii (Copel.) Copel.
2. Chingia clavipilosa Holttum var. clavipilosa.
3. Chingia ferox (Blume) Holttum.; --Cyclosorus ferox (Blume) Ching. Photos  
4. Chingia paucipaleata Holttum. Conservation status: Vulnerable (DAO 2017-11 as Cyclosorus paucipaleatus (Holttum) Mazumdar & Mukhop.).
5. Chingia pricei Holttum. Conservation status: Vulnerable (DAO 2017-11).
6. Chingia urens Holttum. Conservation status: Critically Endangered (DAO 2017-11).

Christella H.Lév. emend. Holttum

1. Christella acuminata (Houtt.) Lév.; --Cyclosorus acuminatus (Houtt.) Ching.
2. Christella arida (D.Don) Holttum; --Cyclosorus aridus (D.Don) Ching. Photos
3. Christella dentata (Forssk.) Brownsey & Jermy.; --Cyclosorus dentatus sensu Copel., 1960, pro parte. --Cyclosorus mindanaensis (Christ) Copel. Pantropic. Throughout the Philippines. Terrestrial in exposed sites, light forests, along stream embankments, etc. Common at low altitudes.
4. Christella hispidula (Decne.) Holttum. --Cyclosorus falcatulus (Christ) Copel.
5. Christella parasitica (L.) Lév. --Cyclosorus dentatus sensu Copel. 1960, pro parte. --Cyclosorus benguetensis Copel. --Cyclosorus procurrens (Mett.) Copel. SE Asia (confined here?, or more widely distributed?). Throughout the Philippines. Terrestrial, open sites and degraded habitats generally, along stream embankments. Altitude?
6. Christella subdentata Holttum.
7. Christella subpubescens (Blume) Holttum.; --Cyclosorus subpubescens (Blume) Ching. --Cyclosorus latipinna (Hook. & Baker) Tard.

Coryphopteris Holttum

1. Coryphopteris borealis Holttum, Blumea 23(1): 35 (1976). Conservation status: Critically Endangered (DAO 2017-11).
2. Coryphopteris dura (Copel.) Holttum, Blumea 23(1): 44 (1976); --Lastrea dura (Copel.) Copel.
3. Coryphopteris pubirhachis (Baker) Holttum var. philippinensis Holttum, Blumea 23(1): 37 (1976). --Lastrea viscosa sensu Copel. (1960) non (Baker) Bedd., pro specim. philipp. Photos
4. Coryphopteris squamipes (Copel.) Holttum, Blumea 23(1): 35 (1976); --Lastrea squamipes (Copel.) Copel. Conservation status: Vulnerable (DAO 2017-11).

Excluded

1. Coryphopteris viscosa (J.Sm.) Holttum, Blumea 19: 33. (1971).

Cyclogramma Tagawa = Cyclosorus

Cyclosorus Link

1. Cyclosorus auriculatus (J.Sm.) C.M.Kuo, Taiwania 47(2): 171 (2002); --Cyclogramma auriculata (J.Sm.) Ching. --Thelypteris simulans Ching. Conservation status: Vulnerable (DAO 2017-11 as Cyclogramma auriculata (J.Sm.) Ching).
2. Cyclosorus interruptus (Willd.) H.Ito. --Cyclosorus gongylodes auctt., doubtfully based on Aspidium gongylodes Schkuhr. Photos

 Haplodictyum C.Presl = Pronephrium, Sphaerostephanos

Macrothelypteris Ching

1. Macrothelypteris polypodioides (Hook.) Holttum. --Lastrea leucolepis C.Presl Photos
2. Macrothelypteris torresiana (Gaudich.) Ching; --Lastrea torresiana (Gaudich.) T.Moore. Photos

Mesophlebion Holttum [= Thelypteris?]

1. Mesophlebion crassifolium (Blume) Holttum, Blumea 22: 232 (1975); --Lastrea crassifolia (Blume) Moore.
2. Mesophlebion rufescens Holttum, Blumea 22: 230 (1975).

Metathelypteris Ching

1. Metathelypteris dayi (Bedd.) Holttum. --Thelypteris singalanensis sensu Holttum, non (Baker) Ching.
2. Metathelypteris gracilescens (Blume) Ching; --Lastrea gracilescens (Blume) Moore. --Lastrea calva (Copel.) Copel. Photos
3. Metathelypteris uraiensis (Rosenst.) Ching.

Nannothelypteris Holttum [= Cyclosorus?]

1. Nannothelypteris aoristisora (Harrington) Holttum; --Cyclosorus aoristisorus (Harrington) Copel. Conservation status: Vulnerable (DAO 2017-11).
2. Nannothelypteris camarinensis Holttum. Conservation status: Vulnerable (DAO 2017-11).
3. Nannothelypteris inaequilobata Holttum. --Lastrea nervosa sensu Copel. (1960), pro parte. Conservation status: Vulnerable (DAO 2017-11). Photos
4. Nannothelypteris nervosa (Fée) Holttum; --Lastrea nervosa (Fée) Copel., pro parte. Conservation status: Vulnerable (DAO 2017-11).
5. Nannothelypteris philippina (C.Presl) Holttum. --Lastrea confusa (Copel.) Copel. Conservation status: Vulnerable (DAO 2017-11).

Parathelypteris (H.Ito) Ching emend. Holttum

1. Parathelypteris beddomei (Baker) Ching. Photos
   • var. beddomei; --Lastrea beddomei (Baker) Bedd.
   • var. eugracilis (Copel.) Holttum; --Lastrea eugracilis Copel. 
2. Parathelypteris glanduligera (Kunze) Ching
3. Parathelypteris grammitoides (Christ) Holttum; --Lastrea grammitoides (Christ) Copel. Photos

Phegopteris Fée

1. Phegopteris decursive-pinnata Fée, Mém. Foug., 5. Gen. Filic. 242 (1850) t. 20 A. f. 1. 1850-52. Known only from one collection on Mt. Iraya, Batan I., Batanes (Barcelona 990).

Plesioneuron (Holttum) Holttum

1. Plesioneuron savaiense (Baker) Holttum. --Lastrea quadriaurita (Christ) Copel. Photos

Pneumatopteris Nakai

1. Pneumatopteris costata (Brackenr.) Holttum. --Lastrea cavitensis Copel.
2. Pneumatopteris glabra (Copel.) Holttum; --Cyclosorus glaber Copel. --Cyclosorus viridis Copel. Photos
3. Pneumatopteris laevis (Mett.) Holttum. --Cyclosorus nitidulus sensu Copel. (1960), pro parte. --Cyclosorus microlonchus sensu Copel. (1960), pro parte. Photos
4. Pneumatopteris ligulata (C.Presl) Holttum; --Lastrea ligulata C.Presl --Lastrea foxii (Christ) Copel. Photos 
5. Pneumatopteris lithophila Holttum.
6. Pneumatopteris michaelis Holttum.
7. Pneumatopteris microloncha (Christ) Holttum; --Cyclosorus microlonchus (Christ) Copel.
8. Pneumatopteris nitidula (C.Presl) Holttum; --Cyclosorus nitidulus (C.Presl) Copel. (1960), pro parte. Photos
9. Pneumatopteris truncata (Poir.) Holttum; --Cyclosorus truncatus (Poir.) Farw. Photos

Pronephrium C.Presl

1. Pronephrium amphitrichum Holttum. Photos
2. Pronephrium asperum (C.Presl) Holttum. --Cyclosorus philippinarum (Fée) Copel.
3. Pronephrium bakeri (Harrington) Holttum; --Haplodictyum bakeri (Harrington) Ching; --Cyclosorus bakeri (Harrington) Copel. Photos
4. Pronephrium balabacensis PM Zamora & Co, Nat. Appl. Sc. Bull. Univ. Philipp. 32  (1980) 45, t. 2. Possibly synonymous with Pronephrium merrilliii. The exindusiate condition of the sori mentioned in the original description may actually be due to small, fugaceous indusia.
5. Pronephrium bulusanicum (Holttum) Holttum; --Haplodictyum bulusanicum Holttum. Conservation status: Vulnerable (DAO 2017-11).
6. Pronephrium clemensiae (Copel.) Holttum. Photos
   • var. clemensiae; --Cyclosorus clemensiae (Copel.) Ching.
   • var. degenerum (Christ) Holttum; --Cyclosorus degener (Christ) Copel. 
7. Pronephrium cuspidatum (Blume) Holttum; --Cyclosorus cuspidatus (Blume) Copel.
8. Pronephrium granulosum (C.Presl) Holttum; --Cyclosorus granulosus (C.Presl) Copel. --Cyclosorus subdimorphus Copel. Photos
9. Pronephrium gymnopteridifrons (Hayata) Holttum. --Cyclosorus pustulosus Copel.
10. Pronephrium heterophyllum (C.Presl) Holttum; --Haplodictyum heterophyllum C.Presl Photos
11. Pronephrium hosei (Baker) Holttum; --Cyclosorus hosei (Baker) Copel. Conservation status: Vulnerable (DAO 2017-11).
12. Pronephrium lineatum (Blume) Presl.
13. Pronephrium melanophlebium (Copel.) Holttum forma melanophlebium; --Cyclosorus melanophlebius (Copel.) Copel.
    forma proliferum Co, ined.
14. Pronephrium menisciicarpon (Blume) Holttum. --Cyclosorus latifolius (C.Presl) Copel.
15. Pronephrium merrillii (Christ) Holttum; --Cyclosorus merrillii (Christ) Copel.
16. Pronephrium nitidum Holttum. --Cyclosorus urophyllus sensu Copel. (1960), excl. loc. Malacca. Photos
17. Pronephrium ramosii (Christ) Holttum var. ramosii. --Cyclosorus cuspidatus sensu Copel. (1960), excl. syn.; --Meniscium cuspidatum Blume. Photos
18. Pronephrium rhombeum (Christ) Holttum; --Cyclosorus rhombeus (Christ) Copel. --Cyclosorus lanceolus (Christ) Copel. Photos
19. Pronephrium rubidum (J.Sm. ex Hook.) Holttum; --Cyclosorus rubidus (Hook.) Copel.
20. Pronephrium samarense (Copel.) Holttum; --Cyclosorus samarensis Copel.
21. Pronephrium solsonicum Holttum. Conservation status: Vulnerable (DAO 2017-11).
22. Pronephrium trachyphyllum Holttum.
23. Pronephrium triphyllum (Sw.) Holttum; --Cyclosorus triphyllus (Sw.) Tard.
24. Pronephrium X xiphioides (Christ) Holttum; --Cyclosorus xiphioides (Christ) Copel. --Cyclosorus edanyoi Copel. --Cyclosorus rhombeus sensu Copel. (1960), pro parte. Photos

Pseudocyclosorus Ching [= Cyclosorus?]

1. Pseudocyclosorus tylodes (Kunze) Moore; --Lastrea tylodes (Kunze) Moore. ‘xylodes’ in Copeland’s Fern Flora.

Pseudophegopteris Ching

1. Pseudophegopteris aurita (Hook.) Ching. Photos
2. Pseudophegopteris paludosa (Blume) Ching. --Lastrea pyrrhorhachis sensu Copel., Fern Fl. Philipp. (1960) 330, pro parte. Photos

Sphaerostephanos J.Sm.

1. Sphaerostephanos alcasidii Holttum.
2. Sphaerostephanos acrostichoides (Desv.) Holttum. --Sphaerostephanos smithianus (C.Presl) Holttum. --Cyclosorus boholensis Copel. --Cyclosorus productus sensu Copel. (1960), pro parte.
3. Sphaerostephanos angustifolius (C.Presl) Holttum; --Cyclosorus angustifolius (C.Presl) Copel. Conservation status: Vulnerable (DAO 2017-11).
4. Sphaerostephanos appendiculatus (Blume) Holttum. Photos
5. Sphaerostephanos cartilagidens PM Zamora & Co, Nat. Appl. Sc. Bull. Univ. Philipp.  32 (1980) 43, t. 1. Conservation status: Other Threatened Species (DAO 2017-11).
6. Sphaerostephanos convergens Holttum.
7. Sphaerostephanos dichrotrichoides (Alderw.) Holttum; --Lastrea dichrotrichoides (Alderw.) Copel. Conservation status: Other Threatened Species (DAO 2017-11).
8. Sphaerostephanos diminutus (Copel.) M.G.Price; --Cyclosorus diminutus (Copel.) Copel.; --Pronephrium diminutum (Copel.) Holttum. --Sphaerostephanos lucbanii Holttum
9. Sphaerostephanos diversilobus (C.Presl) Holttum; --Cyclosorus diversilobus (C.Presl) Ching. Photos
10. Sphaerostephanos ellipticus (Rosenst.) Holttum var. ellipticus. --Cyclosorus ellipticus (Rosenst.) Copel. var. glabrior Holttum
11. Sphaerostephanos erectus (Copel.) Holttum; --Cyclosorus erectus Copel.
12. Sphaerostephanos foxworthyi Holttum
13. Sphaerostephanos grandescens Holttum
14. Sphaerostephanos hernaezii Holttum. Conservation status: Vulnerable (DAO 2017-11).
15. Sphaerostephanos heterocarpus (Blume) Holttum. Eight variants of this polymorphic species were enumerated (but no formal names given) by Holttum in the his FM revision. Two are definitely occurring in the Philippines, while another 2 resembles those ascribed to Borneo and New Guinea forms. Thus: -Mindanao form -Luzon form [as Cyclosorus bordenii (Christ) Ching in ‘Fern Flora’]. From Palawan is one collection resembling the Borneo form, while another one from San Ramon, Zamboanga is nearer to the New Guinea form. Photos
16. Sphaerostephanos hirsutus (Kunze ex Mett.) Holttum var. hirsutus. --Cyclosorus adenophorus (C.Chr.) Ching.
17. Sphaerostephanos irayensis (Copel.) Holttum; --Plesioneuron irayense (Copel.) Holttum; --Cyclosorus irayensis Copel. --Sphaerostephanos fenixii Holttum. Conservation status: Other Threatened Species (DAO 2017-11).
18. Sphaerostephanos lastreoides (C.Presl) Holttum. --Cyclosorus duplosetosus Copel. Photos
19. Sphaerostephanos latebrosus (Kunze ex Mett.) Holttum. --Sphaerostephanos todayensis (Christ) Holttum; --Cyclosorus todayensis (Christ) Copel. --Cyclosorus sagittifolioides Copel. --Cyclosorus halconensis Copel. --Cyclosorus polypterus Copel.
20. Sphaerostephanos lobatus (Copel.) Holttum; --Cyclosorus lobatus Copel. --Cyclosorus bordenii sensu Copel. (1960), pro parte.
21. Sphaerostephanos loherianus (Christ) Holttum; --Cyclosorus loherianus (Christ) Copel.
22. Sphaerostephanos magnus (Copel.) Holttum; --Cyclosorus magnus Copel.
23. Sphaerostephanos major (Copel.) Holttum; --Haplodictyum majus Copel. --Cyclosorus dimorphus Copel.
24. Sphaerostephanos norrisii (Rosenst.) Holttum. --Sphaerostephanos elmerorum (Copel.) Holttum; --Cyclosorus elmerorum Copel. --Cyclosorus ellipticus sensu Copel. (1960), pro parte.
25. Sphaerostephanos perglanduliferus (Alderw.) Holttum.
26. Sphaerostephanos polisianus Holttum.
27. Sphaerostephanos polycarpus (Blume) Copel. --Sphaerostephanos asplenioides J.Sm.
28. Sphaerostephanos porphyricola (Copel.) Holttum; --Cyclosorus pophyricola (Copel.) Ching.
29. Sphaerostephanos productus (Kaulf.) Holttum; --Cyclosorus productus (Kaulf.) Ching. --Cyclosorus serratus Copel. --Cyclosorus weberi Copel. --Cyclosorus rigidus Copel. --Cyclosorus leucadenius Copel. --Sphaerostephanos punctatus Holttum. --Cyclosorus megaphyllus sensu Copel. (1960). --Cyclosorus nitidulus sensu Copel. (1960), pro parte.
30. Sphaerostephanos santomasii Holttum.
31. Sphaerostephanos sessilipinna (Copel.) Holttum; --Cyclosorus sessilipinna (Copel.) Ching. --Cyclosorus austrophilippinus (Copel.) Copel.
32. Sphaerostephanos simplicifolius (J.Sm. ex Hook.) Holttum; --Cyclosorus simplicifolius (J.Sm.) Copel.
33. Sphaerostephanos spenceri (Christ) Holttum; --Cyclosorus spenceri (Christ) Copel. Conservation status: Other Threatened Species (DAO 2017-11).
34. Sphaerostephanos stenodontus (Copel.) Holttum; --Cyclosorus stenodontus Copel. Conservation status: Other Threatened Species (DAO 2017-11).
35. Sphaerostephanos subcordatus Holttum.
36. Sphaerostephanos tephrophyllus (Copel.) Holttum; --Cyclosorus tephrophyllus Copel. Conservation status: Other Threatened Species (DAO 2017-11).
37. Sphaerostephanos unitus (L.) Holttum Photos
    • var. unitus; --Cyclosorus unitus (L.) Ching Sri Lanka, S India, across Malesia. Throughout the Philippines. Terrestrial. Degraded habitats generally, along waterways, etc. at low to medium altitudes. Very common.
    • var. mucronatus (Christ) Holttum.
38. Sphaerostephanos urdanetensis (Copel.) Holttum; --Cyclosorus urdanetensis Copel. --Cyclosorus matutumensis Copel.
39. Sphaerostephanos williamsii (Copel.) Holttum; --Lastrea williamsii Copel. Conservation status: Other Threatened Species (DAO 2017-11).

Status?

1. Sphaerostephanos mindorensis Holltum, Kalikasan 4 (1975) 54. Type: M.G.Price 690 (K,  holotype; isotype in PNH). Mindoro: Mt Halcon.  MINDORO: Mt Halcon, in forest, 750m. Known from two collections from the type locality. Not accounted for in Holltum’s Flora Malesiana revision.

Stegnogramma Blume [= Cyclosorus?]

1. Stegnogramma calcicola Co, sp. nov., ined. BOHOL.
2. Stegnogramma dissitifolia Holttum.
3. Stegnogramma gymnocarpa (Copel.) K Iwats. ssp. gymnocarpa; --Lastrea gymnocarpa Copel.
   

Literature

Alston, AHG. 1959. Isoetaceae. Flora Malesiana series 2, 1: 63-64.

Barcelona, JF. Picasa photogalery of Philippine pterophytes and lycophytes.

Barcelona, JF. 2002. Philippine pteridophyte collections as a resource for conservation planning. Proceedings of the “International Symposium on the Fern Flora Worldwide: Threats and Responses” sponsored by the British Pteridological Society and IUCN- Species Survival Commission.” Fern Gaz. 16(6, 7 & 8): 307-312.

Barcelona, JF. 2003. The taxonomy and ecology of the pteridophytes of Mt. Iraya & vicinity, Batan Island, Batanes Province, Northern Philippines, in Pteridology in the New Millenium (Chandra and Srivastava, eds). Kluwer Academy Publishers. The Netherlands, 299-325.

Barcelona, JF. 2003. Preliminary report on the ferns and fern allies (pteridophytes) of Mt. Bali-it, Balbalasang-Balbalan National Park, Kalinga, Northern Luzon, Philippines. Sylvatrop: The Technical Journal of Philippine Ecosystems and Natural Resources 13(1&2): 81-92.

Barcelona, JF. 2004. Collection and conservation status of pteridophytes in Panay Island, Philippines. Philipp. Scient. 41:57-73.

Barcelona, JF. 2005. Noteworthy fern discoveries in the Philippines at the turn of the 21st Century. Fern Gaz.17(3): 139-146.

Barcelona, JF, B. Hernaez, & M. G. Price. 1996. Philippine Schizaea. Asia Life Sciences 5(1): 27-34.

Barcelona, JF. & M. G. Price. 1999. A new Philippine Goniophlebium. Fern Gaz. 15(7). 261-264.

Barcelona, JF., NE Dolotina, GS Madroñero, WG Granert & DD Sopot. 2006. The ferns and fern allies of the karst forests of Bohol Island, Philippines. Amer. Fern J. 96(1): 1-20.

Brownsey PJ. 1987. A review of the fern genus Hypolepis (Dennstaedtiaceae) in the Malesian and Pacific regions. Blumea 32: 227-276

CABI Invasive Species Compendium

Chambers TC, Farrant PA. 2001 Revision of Blechnum (Blechnaceae) in Malesia. Blumea 46: 283-350

Chen, C-W, LT Ngan, A Hidayat, L Evangelista, HP Nooteboom & W-L Chiou. 2014. First insights into the evolutionary history of the Davallia repens complex. Blumea 59: 49-58

Christenhusz, MJM, X-C Zhang, H Schneider. 2011. A linear sequence of extant families and genera of lycophytes and ferns. Phytotaxa 19: 7-54.

DENR Administrative Order. 2017. Updated national list of threatened Philippine plants and their categories. DAO 2017-11

Dong, S-H. 2014. Nomenclatural novelties in the fern genus Tectaria (Tectariaceae). Phytotaxa 178: 225-228.

Ebihara, A, J-Y Dubuisson, K Iwatsuki, S Hennequin, & M Ito. 2006. A taxonomic revision of the Hymenophyllaceae. Blumea 51: 221-280.

Gruezo, WS. 2012. Tmesipteris zamorae: a new species of Tmesipteris Bernh. (Psilotaceae) from the Philippines. Asia Life Sciences 21: 653-670.

Hassler, M & B Schmitt. World ferns: checklist of ferns and lycophytes of the world

Hauke, RI. 1963. A taxonomic monograph of the genus Equisetum subgenus Hippochaete. Beih. Nova Hedwigia 8: 1-123.

Hennipman, E. 1970. Some novelties in Bolbitis from Asia and the Pacific (Filices). Blumea 18: 147-149

Hennipman, E. 1974. The lectotypification of Acrostichum acuminatum Willd. and the reinstatement of the name Photinopteris speciosa (Bl.) Presl. Blumea 22: 51-52

Hennipman, E. 1977. A monograph of the fern genus Bolbitis (Lomariopsidaceae). Leiden Botanical Series 2: 1-329

Hennipman, E. 1987. A new species of Lecanopteris (Polypodiaceae) from the Philippines. Blumea 32: 311-312

Hennipman, E & MC Roos. 1983. Phylogenetic Systematics of the Polypodiaceae (Filicales). Mededelingen van het Botanisch Museum en Herbarium van de Rijksuniversiteit te Utrecht 525: 321-342

Hennipman, E & WLA Hetterscheid. 1984. The emendation of the fern genus Christiopteris, including the transference of two taxa to the microsorioid Polypodiaceae. Mededelingen van het Botanisch Museum en Herbarium van de Rijksuniversiteit te Utrecht 539: 1-10

Holttum, RE. 1959. Keys to the genera of Pteropsida. Flora Malesiana series 2, 1: XV-XXI

Holttum, RE. 1959. Gleicheniaceae. Flora Malesiana series 2, 1: 1-36

Holttum, RE. 1959. Schizaeaceae. Flora Malesiana series 2, 1: 37-61

Holttum, RE. 1959. Cyatheaceae. Flora Malesiana series 2, 1: 65-176

Holttum, RE. 1966. The genera Lomariopsis, Teratophyllum, and Lomagramma in the islands of the Pacific and Australia. Blumea 14: 215-223

Holttum, RE. 1966. The genus Arthropteris J. Sm. in Malesia. Blumea 14: 225-229

Holttum. RE. 1966. Florae Malesianae Precursores XLIV. The fern-genus Elaphoglossum in Malesia, with descriptions of new species. Blumea 14: 317-326

Holttum, RE. 1968. A re-definition of the fern-genus Taenitis Willd. Blumea 16: 87-95

Holttum, RE. 1969. Studies in the family Thelypteridaceae. The genera Phegopteris, Pseudophegopteris, and Macrothelypteris. Blumea 17: 5-32

Holttum, RE. 1971. Studies in the Family Thelypteridaceae. III. A new system of genera in the Old World. Blumea 19: 17-52

Holttum, RE. 1972. Studies in the family Thelypteridaceae IV. The genus Pronephrium Presl. Blumea 20: 105-126

Holttum, RE. 1973. Studies in the family Thelypteridaceae V. The genus Pneumatopteris Nakai. Blumea 21: 293-325

Holttum, RE. 1975. A comparative account of the fern-genera Syngramma J.Sm. and Taenitis Willd., with discussion of their relationships to each other and to other genera. Kew Bulletin 30: 327-343 + pl.

Holttum, RE. 1975. Studies in the family Thelypteridaceae VIII. The genera Mesophlebion and Plesioneuron. Blumea 22: 223-250

Holltum, RE. 1975. Studies in the family Thelypteridaceae. IX. The genus Sphaerostephanos in the Philippines. Kalikasan 4: 47-68.

Holttum, RE. 1976. Studies in the family Thelypteridaceae X. The genus Coryphopteris. Blumea 23: 18-47

Holttum, RE. 1977. Studies in the family Thelypteridaceae. XII. The genus Amphineuron Holttum. Blumea 23: 205-218

Holttum, RE. 1978. Lomariopsis group. Flora Malesiana series 2, 1: 255-330 (Bolbitis treatment by E. Hennipman)

Holttum, RE. 1981. Thelypteridaceae. Flora Malesiana Ser. 2, 1: 331-599

Holttum, RE. 1984. Studies in the fern-genera allied to Tectaria. III. Aenigmopteris and Ataxipteris, two new genera allied to Tectaria Cav., with comments on Psomiocarpa Presl. Blumea 30: 1-11

Holttum, RE. 1985. Studies in the fern genera allied to Tectaria Cav. IV. The genus Ctenitis in Asia, Malesia and the Western Pacific. Blumea 31: 1-38

Holttum, RE. 1991. Tectaria group. Flora Malesiana series 2, 2: 1-132

Holttum, RE. 1991. New taxa in the Tectaria Group (Polypodiaceae) from Malesia. Blumea 35: 547-557

Holttum, RE, U Sen & D Mittra. 1970. Studies in the family Thelypteridaceae II. A comparative study of the type-species of Thelypteris Schmidel, Cyclosorus Link, and Ampelopteris Kunze. Blumea 18: 195-215

Hovenkamp, PH. 1986. A monograph of the fern genus Pyrrosia (Polypodiaceae). Leiden Botanical Series 9: 1-280

Hovenkamp, PH. 1998. An account of the Malay-Pacific species of Selliguea (Polypodiaceae). Blumea 43: 1-108

Hovenkamp, PH. 1998. Lepisorus in Malesia. Blumea 43: 109-115

Hovenkamp, PH, MTM Bosman, E Hennipman, HP Nooteboom, G. Rodl-Linder & MC Roos. 1998. Polypodiaceae. Flora Malesiana series 2, 3: 1-234

Hovenkamp, PH & H Boon-Chuan. 2012. A revision of the fern genus Oleandra (Oleandraceae) in Asia. PhytoKeys 11: 1-37

Hovenkamp, PH & NAP Franken. 1993. An account of the fern genus Belvisia Mirbel (Polypodiaceae). Blumea 37: 511-527

Hovenkamp, PH & F Miyamoto. 2005. A conspectus of the native and naturalized species of Nephrolepis (Nephrolepidaceae) in the world. Blumea 50: 279-322

Karger, DN, S Lehtonen, VB Amoroso, M Kessler. 2012. A new species of Lindsaea (Lindsaeaceae, Polypodiopsida) from Mt. Hamiguitan, Mindanao, Philippines. Phytotaxa 56: 15-20.

Kato, M. 1993. A taxonomic study of the genus Matonia (Matoniaceae). Blumea 38: 167-172

Kato, M. 1998. Matoniaceae. Flora Malesiana series  2, 3: 289-294

Kato, M, Y Yatabe, N Sahashi & N Murakami. 2001. Taxonomic studies of Cheiropleuria (Dipteridaceae). Blumea 46: 513-525

Knapp, R. Picasa albums of Taiwanese pterophytes and gynosperms.

Kramer, KU. 1967. The Lindsaeoid ferns of the Old World II. A revision of Tapeinidium. Blumea 15: 545-556

Kramer, KU. 1967. The Lindsaeoid ferns of the Old World III. Notes on Lindsaea and Sphenomeris in the Flora Malesiana area. Blumea 15: 557-574

Kramer, KU. 1971. Lindsaea Group. Flora Malesiana series 2, 1: 177-254

Laferrière, JE. 1998. Equisetaceae. Flora Malesiana series 2, 3: 287-288

Laferrière, JE. 1998. Cheiropleuriaceae. Flora Malesiana series 2, 3: 285-286

Lehnert, M & FP Coritico. 2018. The genus Dicksonia (Dicksoniaceae - Cyatheales) in western Malesia. Blumea 63: 268-278

Liu, H-M, R-H Jiang, J Guo, P Hovenkamp, LR Perrie, L Shepherd, S Hennequin & H Schneider. 2013. Towards a phylogenetic classification of the climbing fern genus Arthropteris. Taxon 62: 688-700.

Liu, Y-C, CR Fraser-Jenkins, VB Amoroso & W-L Chiou. 2008. Athyrium erythropodum (Woodsiaceae, Pteridophyta), a new Philippine record. Blumea 53: 447-452

Luong, TT, PH Hovenkamp & MSM Sosef. 2015. Revision of the fern genus Orthiopteris (Saccolomataceae) in Malesia and adjacent regions. PhytoKeys 53: 39–71

Masuyama, S & Y Watano. 2010. Cryptic species in the fern Ceratopteris thalictroides (L.) Brongn. (Parkeriaceae). IV. Taxonomic revision. Acta Phytotaxonomica et Geobotanica 61: 75-86.

Murdock, A.G. 2008. A taxonomic revision of the eusporangiate fern family Marattiaceae, with description of a new genus Ptisana. Taxon 57(3): 737-755.

Nooteboom, HP. 1992. Notes on Davalliaceae I. The genera Araiostegia, Davallodes, Leucostegia, and Gymnogrammitis. Blumea 37: 165-187

Nooteboom, HP. 1994. Notes on Davalliaceae II. A revision of the genus Davallia. Blumea 39: 151-214

Nooteboom, HP. 1997. The microsoroid ferns (Polypodiaceae). Blumea 42: 261-395

Nooteboom, HP. 1998. Davalliaceae. Flora Malesiana series 2, 3: 235-276

Panigrahi, G.1993. Parahemionitis, a new genus of Pteridaceae. American Fern Journal 83: 90-92.

Parris, BS. 2009. New genera of Malesian Grammitidacea (Monilophyta). Blumea 54: 217-219

Price, MG. 1990. Philippine fern notes. Contr. Univ. Mich. Herb. 17: 267-278.

Ravensberg, WJ & E Hennipman. 1986. The Pyrrosia species formerly referred to Drymoglossum and Saxiglossum (Filicales, Polypodiaceae). Leiden Botanical Series 9: 281-310

Reid, JD, GM Plunkett, GA Peters. 2006. Phylogenetic relationships in the heterosporous fern genus Azolla (Azollaceae) based on DNA sequence data from three noncoding regions. International Journal of Plant Sciences 167(3): 529-538.

Rödl-Linder. G. 1990. A monograph of the fern genus Goniophlebium (Polypodiaceae). Blumea 34: 277-423

Rödl-Linder, G. 1994. A monograph of the fern genus Thylacopteris (Polypodiaceae). Blumea 39: 351-364

Rolleri, C. 1993. Revision of the genus Christensenia. Amer. Fern Jour. 83(1): 3-19.

Roos, MC. 1985. New combinations in Aglaomorpha Schott (Polypodiaceae). Blumea 31: 153

Ruhfel, B, S Lindsay, CC Davis. 2008. Phylogenetic placement of Rheopteris and Monogramma (Pteridaceae, s.l.): evidence from Rbcl sequence data. Syst. Bot. 33: 37-43.

Salgado, AE. 2017. Does Asplenium falcatum exist in the Philippines? Edinburgh Journal of Botany 74: 1-4.

Saunders, RMK. 1998. Azollaceae. Flora Malesiana series 2, 3: 277-284

Schuettpelz, E, KM Pryer. 2007. Fern phylogeny inferred from 400 Leptosporangiate species and three plastid genes. Taxon 56: 1037-1050.

Schuettpelz, E, H Schneider, L Huiet, MD Windham, KM Pryer. 2007. A molecular phylogeny of the fern family Pteridaceae: Assessing overall relationships and the affinities of previously unsampled genera. Molecular Phylogenetics and Evolution 44:1172-1185.

Smith AR, Pryer KM, Schuettpelz E, Korall P, Schneider H, Wolf PG. 2006. A classification for extant ferns. Taxon 55: 705–731.

Tan, BC. 2013. The Selaginella flora, a good indicator of the Philippine pleistocene island groups. Philippine Journal of Science 142: 211-213.

Tryon. 1941. A revision of the genus Pteridium. Rhodora 43: 1-70.

Wang, LI, Z-Q Wu, QP Xiang, J Heinrichs, H Schneider & XC Zhang. 2010. A molecular phylogeny and a revised classification of tribe Lepisoreae (Polypodiaceae) based on an analysis of four plastid DNA regions. Botanical Journal of the Linnean Society 162: 28-38.

Wardani, W & B Adjie. 2018. The genus Athyrium Roth in Malesia: a checklist with working names and distributions. Sibbaldia 16: 39-48

Wieffering, JH. 1964. A preliminary revision of the Indo-Pacific species of Ophioglossum (Ophioglossaceae). Blumea 12: 321-337

Yesilyurt, JC & H Schneider. 2010. The new fern genus Calciphilopteris (Pteridaceae). Phytotaxa 7: 52-59.

Zhang, LB. 2012. Reducing the fern genus Dryopsis to Dryopteris and the systematics and nomenclature of Dryopteris subgenus Erythrovariae section Dryopsis (Dryopteridaceae). Phytotaxa 71: 17-27

Zhang XC & HP Nooteboom. 1997. Some new combinations in Plagiogyria (Plagiogyriaceae, Pteridophyta). Blumea 42: 483-484

Zhang XC & HP Nooteboom. 1998. A taxonomic revision of Plagiogyriaceae (Pteridophyta). Blumea 43: 401-469

Zhang XC & HP Nooteboom. 1998. Plagiogyriaceae. Flora Malesiana series 2, 3: 295-316

Acknowledgements

Barcelona, J.F. (2011 onwards). Pteridophytes (Lycophytes and Monilophytes). In: Pelser, P.B., J.F. Barcelona & D.L. Nickrent (eds.). Co's Digital Flora of the Philippines. www.philippineplants.org

Copyright © 2011, Co's Digital Flora of the Philippines

Last updated 5 January 2019