Arthropteridaceae = Tectariaceae

Aspleniaceae

Asplenium L.

Last edited by Julie F. Barcelona, 7 February 2021

1. Asplenium aethiopicum (Burm.f.) Bech., Candollea 6 (1935) 22; --Trichomanes aethiopicum Burm.f., Fl. Indica (1768) 32 (err. 28).
   
   • ssp. aethiopicum. Africa, Andaman Isls, Australia, Bangladesh, China, India, Indian Ocean, Java, Lesser Sunda Isls, Middle East, Moluccas, Myanmar, New Caledonia, New Guinea, Pacific, Peninsular Malaysia, Sri Lanka, Sulawesi, ?Thailand, Vietnam, Philippines.
2. Asplenium affine Sw., Schrad. J. Bot. 1800 (2) (1801) 56. --Asplenium spathulinum J.Sm. ex Hook., J. Bot. 3 (1841) 408. --Asplenium gracilifolium Copel., PJS c 4 (1909) 113. Borneo, Cambodia, China, India, Java, Laos, La Reunion, Madagascar, Mauritius, Moluccas, New Guinea, Pacific, Rodrigues Isl, Seychelles, Solomon Isls, Sulawesi, Sumatra, Sri Lanka, ?Tanzania, Thailand, Vietnam, Philippines. Photos
3. Asplenium amboinense Willd., Sp. Pl. 5 (1810) 303. Borneo, Moluccas, New Guinea, Pacific, Sulawesi, Philippines, Palawan, Solomon Isls, Philippines. PALAWAN.
4. Asplenium anguineum Christ, J. Bot. (Morot) sér. 2, 1 (1908) 232. --Asplenium phyllitidis D.Don ssp. malesicum Holttum, Gard. Bull. Sing. 27 (1974) 153. Borneo, Cambodia, China, India, India, Java, Moluccas, Myanmar, New Guinea, Peninsular Malaysia, Solomon Isls, Sumatra, Thailand, Vietnam, Philippines.
   
5. Asplenium apoense Copel., Fragm. Fl. Philipp. (1905) 185. Endemic to the Philippines. MINDANAO: Bukidnon. Photos
6. Asplenium callipteris Fée, Mém. Foug., 5. Gen. Filic. (1850) 193. --Asplenium epiphyticum Copel., Fragm. Fl. Philipp. (1905) 184. Endemic to the Philippines. Photos
7. Asplenium colubrinum Christ, Bull. Herb. Boissier II. 6 (1906) 999. Endemic to the Philippines.
8. Asplenium crinicaule Hance, Ann. Sci. Nat., Bot. sér. 5, 5 (1866) 254. Cambodia, China, India, Laos, Taiwan, Thailand, Tibet, Vietnam, Philippines.
9. Asplenium cuneatum Lam., Encycl. 2 (1786) 309. Africa, Americas, Australia, Caribbean, Comoros, Lesser Sunda Isls, Micronesia, New Guinea, Pacific, Peninsular Malaysia, Solomon Isls, Sulawesi, Madagascar, Micronesia, New Caledonia, Philippines. PALAWAN. Photos
10. Asplenium cymbifolium Christ, Bull. Boiss. 2, 6 (1906) 999. Borneo, Moluccas, New Guinea, ?Samoa, Solomon Isls, Sumatra, Philippines. MINDANAO: Bukidnon, PALAWAN. Photos  
11. Asplenium elmeri Christ, PJS 2 c (1907) Bot. 164. Borneo, Philippines. LUZON. Photos
12. Asplenium exiguum Bedd., Ferns S. India (1863) t.146. Bhutan, China, India, Mongolia, Myanmar, Nepal, Siberia, Taiwan, Thailand, Tibet, Vietnam, Philippines. Photos
13. Asplenium griseum Copel., Polyp. Philipp. (1905) 82. Endemic to the Philippines.
14. Asplenium haenkei Copel., Gen. Fil. (1947) 165. --Triphlebia pinnata (J.Sm. ex Kunze) Baker, Malesia 3 (1886) 42; --Scolopendrium pinnatum J.Sm. ex Kunze, Farnkr. 1. (1843) 124. Moluccas, Philippines. LUZON. Photos
15. Asplenium horridum Kaulf., Enum. Filic. (1824) 173. --?Asplenium caudatum G.Forst., Prodr. (1786) 80. --?Asplenium chamissonianum C.Presl, Tent. Pterid. (1836) 107. ?Bismarck Arch, Lesser Sunda Isls, Moluccas, New Guinea, Pacific, Solomon Isls, Philippines. MINDANAO: Bukidnon. Photos
    
16. Asplenium insiticium Brack., U.S. Expl. Exped., Filic. 16 (1854) 161. Pacific, Philippines. LUZON.
17. Asplenium laciniatum D.Don, Prodr. Fl. Nepal. (1825) 8.
    
    • subsp. fraser-jenkinsii Reichst. ex Pangtey & Khullar, Indian Fern J. 27 ((2011)) 185. --Asplenium semivarians Viane & Reichst., Pteridol. New Millennium (2003) 78. Africa, China, India, Philippines.
18. Asplenium laserpitiifolium Lam., Encycl. 2 (1786) 310. --Asplenium robustum Blume, Enum. Pl. Javae 2 (1828) 189. Australia, Bismarck Arch, Lesser Sunda Isls, Micronesia, Moluccas, New Caledonia, New Guinea, Pacific, ?Ryukyu Isls, Solomon Isls, Sulawesi, Philippines. PALAWAN.
    
19. Asplenium lepturus J.Sm., J. Bot. (Hooker) 3 (1841) 408. --?Asplenium contiguum Kaulf., Enum. Filic. (1824) 172. China, Laos, Sulawesi, Vietnam, Philippines.
20. Asplenium lobulatum Mett., Linnaea 36 (1869) 100. Borneo, New Guinea, Pacific, Solomon Isls, Sulawesi, Philippines. Photos
21. Asplenium longissimum Blume, Enum. Pl. Javae 2 (1828) 178. --Asplenium acutiusculum Blume, Enum. Pl. Javae 2 (1828) 178. --?Asplenium anisodontum C.Presl, Epimel. Bot. (1851) 73. Bangladesh, Cambodia, Chagos Arch, India, Java, Lesser Sunda Isls, Myanmar, Peninsular Malaysia, Singapore, Sulawesi, Thailand, Vietnam, Philippines. Photos  
22. Asplenium macrophyllum Sw., Schrad. J. Bot. 1800 (1801) 52; --Salgado, Edinburgh J. Bot. 74 (2017) 3; --Paclibar & Tadiosa, Philippine J. Syst. Biol. 14 (2020) 9. Borneo, Philippines. LUZON: Quezon.
    
23. Asplenium mantalingahanum P.M.Zamora & Co, Nat. Appl. Sc. Bull. Univ. Philipp. 32 (1980) 46. Endemic to the Philippines. PALAWAN.
24. Asplenium nidus L., Sp. Pl. (1753) 1079. Africa, Australia, Bangladesh, Bismarck Arch, Borneo, Chagos Arch, China, Japan, La Reunion, Lesser Sunda Isls, Madagascar, Mauritius, Micronesia, Moluccas, New Caledonia, Pacific, Peninsular Malaysia, Ryukyu Isls, Singapore, Solomon Isls, Sri Lanka, Sulawesi, Sumatra, Taiwan, Thailand, Tibet, Vietnam, Philippines. MINDANAO: Bukidnon. Photos
    • var. nidus
    • var. curtisorum (Christ) Holtt., Gard. Bull. Sing. 27 (1974) 148; --Asplenium curtisorum Christ, Ann. Jard. Bot. Buitenzorg II. 4 (1904) 39. Borneo, Sulawesi, Philippines.
25. Asplenium nigrescens Blume, Enum. Pl. Javae 2 (1828) 180. --Asplenium militare Copel., PJS 1 (Suppl. 4) (1906) 254. Borneo, ?Java, Moluccas, New Guinea, Solomon Isls, Sulawesi, Philippines. MINDANAO: Bukidnon. Photos
26. Asplenium normale D.Don, Prodr. Fl. Nepal. (1825) 7. Africa, Australia, Bhutan, Cambodia, China, India, Japan, Java, Laos, Lesser Sunda Isls, Madagascar, Moluccas, Myanmar, Nepal, Pacific, Peninsular Malaysia, Ryukyu Isls, South Korea, Sri Lanka, Sulawesi, Sumatra, Taiwan, Thailand, Tibet, Vietnam, Philippines. Photos  
27. Asplenium oxyphyllum J.Sm. ex Kunze, Bot. Zeitung (1846) 441. --Asplenium falcatum auct. non Lam. Indian Ocean, Southeast Asia, West Malesia, China, Hong Kong, Taiwan, Philippines.
    
28. Asplenium pellucidum Lam., Encycl. 2 (1786) 305. Australia, Borneo, Cambodia, Comoros, India, La Reunion, Lesser Sunda Isls, Madagascar, Maldives, Mauritius, Micronesia, Moluccas, Myanmar, New Guinea, Peninsular Malaysia, Rodrigues Isl, Seychelles, Solomon Isls, Southern Marianas, Sri Lanka, Sulawesi, Sumatra, Thailand, Philippines. PALAWAN. Photos  
29. Asplenium persicifolium J.Sm. ex Mett., Aspl. 97 (1859) no. 32. --Asplenium tripinnatifidum Copel., PJS 46 (1931) 215. Borneo, Sulawesi, Philippines. MINDANAO: Bukidnon. Photos
30. Asplenium phyllitidis D.Don, Prodr. Fl. Nepal. (1825) 7; --Asplenium nidus L. var. phyllitidis (D.Don) Alderw., Handb. Malayan Ferns Suppl. (1917) 282. --?Asplenium colubrinum Christ, Bull. Herb. Boissier II. 6 (1906) 999. Bangladesh, China, India, Myanmar, Nepal, Nicobar Isls, Peninsular Malaysia, Philippines, Singapore, Sulawesi, Sumatra, Thailand, Tibet, Vietnam, Philippines. Photos
31. Asplenium scandens J.Sm. ex Mett., Mett. Aspl. 108  (1859) n. 57. Borneo, Micronesia, Moluccas, New Guinea, Solomon Isls, Sulawesi, Philippines. PALAWAN. Photos
32. Asplenium schizocarpum (Copel.) Copel., Gen. Fil. (1947) 165; --Scolopendrium schizocarpum Copel., PJS 1 (Suppl. 2) (1906) 152. Moluccas, New Guinea, Solomon Isls, Sulawesi, Philippines.
33. Asplenium scolopendrioides J.Sm. ex Hook., Icon. Pl. (1854) t. 930. Borneo, Moluccas, Peninsular Malaysia, Philippines. LEYTE.
34. Asplenium setisectum Blume, Enum. Pl. Jav. (1828) 187. Borneo, Java, New Guinea, Peninsular Malaysia, Sulawesi, Philippines. LUZON.
35. Asplenium stantoni Copel., PJS 1 (Suppl. 2) (1906) 151; --Type: Copeland s.n., Luzon: Bontoc, Batilao, 1600m. Endemic to the Philippines. LUZON: Mountain Province.
36. Asplenium steerei Harrington, J. Linn. Soc., Bot. 16 (1877) 28. Moluccas, New Caledonia, New Guinea, Solomon Isls, Taiwan, Philippines. LUZON.
37. Asplenium tenerum G.Forst., Prodr. (1786) 80. Australia, Bonin Isls, Borneo, Cambodia, China, India, Japan, Java, Lesser Sunda Isls, Micronesia, Moluccas, Myanmar, New Guinea, Nicobar Isls, Pacific, Peninsular Malaysia, Ryukyu Isls, Seychelles, Singapore, Solomon Isls, South Korea, Sri Lanka, Sulawesi, Sumatra, Taiwan, Thailand, Vietnam, Philippines. MINDANAO: Bukidnon, PALAWAN. Photos  
38. Asplenium tenuifolium D.Don, Prodr. Fl. Nepal. (1825) 8. Bhutan, China, India, Japan, Myanmar, Nepal, Sri Lanka, Taiwan, Thailand, Vietnam, Philippines.
39. Asplenium thunbergii Kunze, Linnaea 10 (1836) 517. --Asplenium decorum Kunze, Bot. Zeitung (Berlin) 6 (1848) 176. India, Java, Lesser Sunda Isls, Madagascar, Moluccas, Sri Lanka, Sulawesi, Sumatra, Thailand, Vietnam, Philippines. MINDANAO: Bukidnon. Photos
40. Asplenium vittiforme Cav., Descr. (1802) 255; --Coritico & Amoroso, Nature Conservation Research 5 (2020) 81. Borneo, Java, Micronesia, Peninsular Malaysia, Sulawesi, Sumatra, Thailand, Philippines. MINDANAO: Bukidnon, Davao Oriental, Misamis Occidental, North Cotabato, PALAWAN. Conservation status: Vulnerable (DAO 2017-11). Photos
41. Asplenium vulcanicum Blume, Enum. Pl. Jav. (1828) 176. --?Asplenium salignum Blume, Enum. Pl. Javae 2 (1828) 175. Borneo, Java, Lesser Sunda Isls, Micronesia, Moluccas, Peninsular Malaysia, Sulawesi, Philippines.
42. Asplenium yoshinagae Makino
    
    • subsp. austroindicum Fraser-Jenk., Taxon. Revis. Indian Subcontinental Pteridophytes (2008) 160. India, Sri Lanka, Thailand, Philippines.

Possibly present:

1. Asplenium polyodon G.Forst., Prodr. (1786) 80. Australasia, Micronesia, New Caledonia, New Guinea, Pacific, Solomon Isls, ?Philippines. ?PALAWAN. Photos
2. Asplenium powellii Baker, Syn. Fil. (1867) 224. Moluccas, New Guinea, Solomon Isls, ?Philippines.
3. Asplenium serricula Fée, Mém. Foug. 5 (1852) 196. India, Sri Lanka, Taiwan, ?Philippines.

Hymenasplenium Hayata.

Last edited by Julie F. Barcelona, 4 June 2021

1. Hymenasplenium adiantifrons (Hayata) Viane & S.Y.Dong, Fl. China 2-3 (2013) 312; --Asplenium resectum f. adiantifrons Hayata, Ic. Pl. Formos. 4 (1914) 226. --?Asplenium filipes Copel., PJS 3 c (1908) Bot. 304. Taiwan, Philippines.
   
2. Hymenasplenium cheilosorum (Kunze ex Mett.) Tagawa, Acta Phytotax. Geobot. 7 (1938) 84; --Asplenium cheilosorum Kunze ex Mett., Aspl. (1859) 133. Bangladesh, Bhutan, Borneo, China, India, Myanmar, Nepal, Peninsular Malaysia, Ryukyu Isls, Sri Lanka, Taiwan, Thailand, Tibet, Vietnam, Yakushima Isl, Philippines. Photos
3. Hymenasplenium excisum (C.Presl) S.Linds., Thai Forest Bull., Bot. 37 (2009) 69; --Asplenium excisum C.Presl, Epimel. Bot. (1851) 74; --Type: Cuming 110 (BM, K, PRC, syn), Luzon. --Asplenium subresectum Copel., PJS 83 (1954) 97. Australia, Bhutan, Cambodia, China, India, Moluccas, Myanmar, Nepal, New Guinea, Pacific, Peninsular Malaysia, Ryukyu Isls, Solomon Isls, Sri Lanka, Sulawesi, Taiwan, Thailand, Tibet, Vietnam, Philippines. PALAWAN. Photos
4. Hymenasplenium obscurum (Blume) Tagawa, Acta Phytotax. Geobot. 7 (1938) 83; --Asplenium obscurum Blume, Enum. Pl. Javae 2 (1828) 181. Africa, Bhutan, Borneo, China, India, Java, Lesser Sunda Isls, Madagascar, Moluccas, Myanmar, Nepal, New Guinea, Sri Lanka, Sulawesi, Sumatra, Thailand, Vietnam, Philippines. PALAWAN.
5. Hymenasplenium ofeliae (Salgado) Barcelona & Pelser, Blumea 66 (2021) 96; --Asplenium ofeliae Salgado, Amer. Fern J. 93 (2003) 193; --Type: Merrill 700 (US, holo; PRC, iso), Luzon: Benguet Prov., May-1911. Endemic to the Philippines. LUZON: Benguet. Photos 
   
6. Hymenasplenium subnormale (Copel.) Nakaike, New Fl. Jap. Pterid. (1992) 841; --Asplenium subnormale Copel., Fragm. Fl. Philip. (1905) 183. Borneo, China, Moluccas, Peninsular Malaysia, Ryukyu Isls, Sulawesi, Taiwan, Philippines. LUZON, PALAWAN. Photos

1. Hymenasplenium obliquissimum (Hayata) Sugim., Keys Herb. Pl. Jap. Pterid. 356 (1966) 406. China, India, Japan, Myanmar, Nepal, Vietnam, ?Philippines.
2. Hymenasplenium pubirhizoma (Ching & Z.Y.Liu) K.W.Xu & Li Bing Zhang, Phytotaxa 351 (2018) 186; --Asplenium pubirhizoma Ching & Z.Y.Liu, Bull. Bot. Res. Harbin 3 (1983) 7. Borneo, China, India, Japan, Myanmar, Ryukyu Isls, Vietnam, Yakushima Isl, ?Philippines.

1. Asplenium unilaterale (?auct. non) Lam., Encycl. 2 (1786) 305. Note: The name Asplenium unilaterale Lam. has been misapplied to some SE Asian plants (Lin & Viane 2013) and it is presently unclear to which species some of the Philippine plants belong. MINDANAO: Bukidnon. Photos

Athyriaceae

Anisocampium C.Presl

Last edited by Julie F. Barcelona, 2 July 2019

1. Anisocampium cumingianum C.Presl, Abh. Königl. Böhm. Ges. Wiss., ser. 5 6 (1851) 418; --Athyrium cumingianum (C.Presl) Ching, Index Filic., Suppl. 3 (1934) 40. Cambodia, China, India, Java, Laos, Lesser Sunda Isls, Myanmar, Sri Lanka, Sulawesi, Taiwan, Thailand, Philippines. LUZON. Photos

Athyrium Roth

Last edited by Julie F. Barcelona, 7 September 2020

1. Athyrium anisopterum Christ, Bull. Herb. Boissier 6(12) (1898) 962; --Type: A. Henry 10109 (MO, iso). Bhutan, Borneo, China, India, Java, Myanmar, Nepal, Peninsular Malaysia, Sri Lanka, Sumatra, Taiwan, Thailand, Tibet, Vietnam, Philippines. LUZON. Photos
2. Athyrium drepanopterum (Kunze) A.Br. ex Milde, Fil. Eur. (1867) 49; --Polypodium drepanopterum Kunze, Linnaea 23 (1850) 318. Bhutan, China, India, Myanmar, Nepal, Peninsular Malaysia, Sumatra, Thailand, Tibet, Vietnam, Philippines. Photos
3. Athyrium eburneum (J.Sm. ex Mett.) J.Sm., Hist. Fil. (1875) 328; --Asplenium eburneum J.Sm. ex Mett., Cat. Kew Ferns (1857) 47. Bhutan, China, India, Japan, Myanmar, Nepal, Taiwan, Tibet, Vietnam, Philippines.
   
4. Athyrium elmeri Copel., PJS 3 c (1908) 292. Endemic to the Philippines. Photos
5. Athyrium erythropodum Hayata, Icon. Pl. Formosan. 4 (1914) 233. Java, Lesser Sunda Isls, Taiwan, Philippines. LUZON, MINDANAO.
6. Athyrium nakanoi Makino, Bot. Mag. 23 (1909) 247; --Coritico et al., Philippine J. Syst. Biol. 13 (2019) 47. Bhutan, Borneo, China, India, Japan, Nepal, Peninsular Malaysia, Sumatra, Taiwan, Philippines. MINDANAO: Bukidnon. Conservation status: Endangered (Coritico et al. 2019).
   
7. Athyrium nitidulum (Kunze) Milde, Bot. Zeitung (Berlin) 28 (1870) 370. --Athyrium brevipinnulum Copel., PJS 3 c (1908) 291; --Type: Copeland 1939 (MICH, holo). --Athyrium mearnsianum (Copel.) Alderw., Mal. Ferns Suppl. (1917) 279; --Athyrium nigripes var. mearnsianum Copel., PJS 3 c (1908) 291; --Type: Elmer 6543. --Athyrium philippinense Christ, PJS 1 (Suppl. 4) (1906) 254; --Athyrium sarasinorum var. philippinense Christ, Bull. Boiss. 6 (1898) 154. Java, Philippines. Photos
8. Athyrium oppositipennum Hayata, J. Coll. Sci. Imp. Univ. Tokyo 30 (1911) 441. --Athyrium oreopteris Copel., PJS 40 (1929) 302; --Type: Copeland 2305 (MICH, holo; UC, iso). Taiwan, Philippines. LUZON.
9. Athyrium puncticaule (Blume) T.Moore, Index Fil. (1860) 186; --Aspidium puncticaule Blume [Enum. Pl. Jav. (1828) 156. Borneo, India, Japan, Java, Lesser Sunda Isls, Moluccas, Sri Lanka, Sulawesi, Sumatra, Taiwan, Vietnam, Philippines. LUZON, MINDANAO: Bukidnon. Photos
10. Athyrium tozanense (Hayata) Hayata, Icon. Pl. Formosan. 4 (1914) 235; --Asplenium tozanense Hayata, J. Coll. Sci. Imp. Univ. Tokyo 80 (1911) 440. --Athyrium aristulatum Copel., PJS 1 (Suppl. 4) (1906) 253; --Type: Copeland 1880 (MICH, holo). Sulawesi, Sumatra, Philippines. Photos

Possibly present:

1. Athyrium fallaciosum Milde, Fil. Eur. 6 (1867) 54. --Athyrium fauriei (Christ) Makino, Bot. Mag. (Tokyo) 17 (1903) 160; --Nephrodium fauriei Christ, Bull. Boiss. 4 (1896) 671; --Aspidium fauriei (Christ) Christ, Bull. Boiss. 6 (1898) 193. China, Japan, North Korea, South Korea, ?Philippines. ?LUZON.
2. Athyrium minutum Copel., Univ. Calif. Publ. Bot. 18 (1942) 221. Sulawesi, New Guinea, ?Philippines.
3. Athyrium setiferum C.Chr., Index Filic. 3 (1905) 146. Bhutan, Borneo, China, China, India, Japan, Moluccas, Myanmar, Nepal, Peninsular Malaysia, Sri Lanka, Sumatra, Thailand, Tibet, Tibet, ?Philippines. ?LUZON.

Callipteris Bory = Diplazium

Cornopteris Nakai

Last edited by Pieter B. Pelser,  21 July 2019

1. Cornopteris banahaoensis (C.Chr.) K.Iwats. & M.G.Price, South E. Asian Stud. 14 (1977) 564; --Dryopteris banahaoensis C.Chr., Index Filic., Suppl. 1 (1913) 30. --Athyrium nudum Copel., FFP 3 (1958) 391. Bhutan, India, Nepal, Taiwan, Volcano Isls, Yakushima Isl, Philippines. LUZON.
2. Cornopteris decurrenti-alata (Hook.) Nakai, Bot. Mag. (Tokyo) 44 (1930) 8; --Gymnogramma decurrenti-alata Hook., Sp. Fil. 5 (1864) 142. China, India, Japan, Myanmar, Nepal, South Korea, Taiwan, Tibet, Vietnam, Philippines. Photos
3. Cornopteris opaca (D.Don) Tagawa, Acta Phytotax. Geobot. 8 (1939) 92; --Hemionitis opaca D.Don, Prodr. Fl. Nepal. (1825) 13. --Athyrium gymnocarpum Copel., 40 (1929) 301. Bhutan, Borneo, China, India, Japan, Java, ?Laos, Lesser Sunda Isls, Moluccas, Myanmar, Sulawesi, Taiwan, Thailand, Vietnam, Philippines.
4. Cornopteris philippinensis M.Kato, Acta Phytotax. Geobot. 30 (1979) 110. Moluccas, Philippines.

Deparia Hook. & Grev.

Last edited by Julie F. Barcelona, 20 July 2019

1. Deparia boryana (Willd.) M.Kato, Bot. Mag. Tokyo 90 (1977) 36; --Dryoathyrium boryanum (Willd.) Ching, Bull. Fan Mem. Inst. Biol., Bot. 11 (1941) 81; --Ctenitis boryana (Willd.) Copel., Gen. Fil. (1947) 123; --Aspidium boryanum Willd., Sp. Pl. 5 (1810) 285.
   • ssp. austroindica Fraser-Jenk., Taxon. Revis. Indian Subcontinental Pteridophytes (2008) 252. India, ?New Guinea, Peninsular Malaysia, Sri Lanka, ?Philippines.
   • ssp. boryana. Africa, Bhutan, Borneo, China, Fiji, India, Indian Ocean, Japan, Java, Lesser Sunda Isls, Moluccas, Myanmar, Nepal, Peninsular Malaysia, Sri Lanka, Sulawesi, Sumatra, Thailand, Tibet, Vietnam, Philippines.
2. Deparia confluens (Kunze) M.Kato, J. Fac. Sci. Univ. Tokyo, Bot. 13 (1984) 413; --Asplenium confluens Kunze [Bot. Zeit. (Berlin) (1848) 174. --Athyrium grammitoides (C.Presl) Milde, Bot. Zeitung (Berlin) 28 (1870) 353; --Diplazium grammitoides C.Presl, Abh. Kön. Böhm. Ges. Wiss. 5(6) (Epim. Bot.) (1851) 84. Borneo, Java, Lesser Sunda Isls, Ryukyu Isls, Sulawesi, Philippines. Photos
3. Deparia formosana (Rosenst.) Sano, Act. Phytotax. Geobot. 51 (2000) 17; --Diplaziopsis formosana (Rosenst.) M.G.Price, Contrib. Univ. Mich. Herb. 17 (1990) 269; -- Diplazium formosanum Rosenst., Hedw. 56 (1915) 337. China, Taiwan, Yakushima Isl, Vietnam, Philippines.
4. Deparia japonica (Thunb.) M.Kato, Bot. Mag. Tokyo 90 (1977) 37; --Lunathyrium japonicum (Thunb.) Kurata, J. Geobot. 9 (1961) 99; --Athyrium japonicum (Thunb.) Ching, PJS 3 (1908) 290; --Asplenium japonicum Thunb., Syst. Veg. ed. 14 (1784) 934.
   
   • ssp. japonica. Bangladesh, Bhutan, China, East Asia, India, Japan, Lesser Sunda Isls, Myanmar, Nepal, North Korea, Pakistan, Peninsular Malaysia, Ryukyu Isls, South Korea, ?Sri Lanka, Vietnam, Philippines. Photos
5. Deparia lancea (Thunb.) Fraser-Jenkins, New Sp. Syndr. Indian Pteridol. (1997) 101; --Athyrium lanceum (Thunb.) Milde, nom illeg, auct. non Moore; --Copel., PJS 7 c (1912) 55; --Diplazium lanceum (Thunb.) C.Presl, Tent. Pterid. (1836) 113, nom. illeg., auct. non Bory; --Micropodium lanceum (Thunb.) J.Smith, Bot. Zeitung (Berlin) 28 (1870) 353; --Triblemma lancea (Thunb.) Ching., Acta Phytotax. Sin. 16 (1978) 24; --Asplenium lanceum Thunb., Fl. Jap. (1784) 333. --Asplenium subsinuatum Wall. ex Hook. & Grev., Icon. Fil. (1827) t.27; --Diplazium subsinuatum (Wall. ex Hook. & Grev.) Tagawa, Col. Ill Jap. Pterid. (1959) 203; --Price, Kalikasan 1 (1972) 42. --Athyrium dubium (D.Don) Ohwi, Bull. Natl. Sci. Mus. 3 (1956) 99, nom. illeg., auct. non Ching; --Copel., FFP 3 (1961) 412; --Scolopendrium dubium D.Don, Prodr. Fl. Nepal. (1825) 9. --Diplazium simplicifolium Kodama, Icon. Pl. Koisik. 1 (1913) pl.68. Bangladesh, Bhutan, Borneo, ?Cambodia, China, India, Japan, Laos, Myanmar, Nepal, Ryukyu Isls, South Korea, Sri Lanka, Taiwan, Thailand, Vietnam, Philippines. BATANES, LUZON, PANAY. Photos
6. Deparia petersenii (Kunze) M.Kato, Bot. Mag. (Tokyo) 90 (1977) 37; --Asplenium petersenii Kunze, Analecta Pteridogr. (1837) 24.
   
   • ssp. petersenii. Bhutan, Bonin Isls, Borneo, China, India, Indian Ocean, Japan, Lesser Sunda Isls, Moluccas, Myanmar, Nepal, New Guinea, Pakistan, Ryukyu Isls, Solomon Isls, South Korea, Sri Lanka, Sulawesi, Taiwan, Tibet, Vietnam, Volcano Isls, Philippines. Photos
7. Deparia subfluvialis (Hayata) M.Kato, J. Fac. Sci. Univ. Tokyo, Sect. 3, Bot. 13 (1984) 389; --Dryopteris subfluvialis Hayata, Icon. Pl. Formosan. 5 (1915) 288. Taiwan, Thailand, Philippines.

Diplazium Sw.

1. Diplazium atratum Christ, PJS 2 c (1907) 163 1907; --Athyrium atratum (Christ) Copel., PJS 3 c (1908) 293. Endemic to the Philippines. PALAWAN.
2. Diplazium bantamense Blume, Fil. Jav. (18560 190; --Athyrium bantamense (Blume) Milde, Bot. Zeitung (Berlin) 28 (1870) 353; --Asplenium bantamense Baker, Syn. Fil. (1867) 23. ?Bangladesh, Borneo, Java, Lesser Sunda Isls, Moluccas, New Guinea, Peninsular Malaysia, Solomon Isls, Sulawesi, Sumatra, Thailand, Vanuatu, Vietnam, Philippines. PALAWAN.
3. Diplazium bolsteri Copel., PJS 1 (Suppl. 4) (1906) 252; --Athyrium bolsteri (Copel.) Copel., PJS 3 c (1908) 296. Endemic to the Philippines. MINDANAO.
4. Diplazium brachysoroides Copel., PJS 2 c (1907) 127; --Athyrium brachysoroides (Copel.) Copel., PJS 3 (1908) 296. Endemic to the Philippines. MINDORO.
5. Diplazium calliphyllum (Copel.) M.G.Price, Contrib. Univ. Mich. Herb. 17 (1990) 270; --Athyrium calliphyllum Copel., PJS 40 (1929) 303. Moluccas, Philippines. MINDANAO. Conservation status: Other Threatened Species (DAO 2017-11).
6. Diplazium ciliatum M.G.Price, Contr. Univ. Michigan Herb. 17 (1990) 270. Endemic to the Philippines.
   
7. Diplazium cordifolium Blume, Enum. Pl. Jav. (1828) 190; --Athyrium cordifolium (Blume) Copel., PJS 3 c (1908) 300. --Athyrium parvum Copel., PJS 81 (1952) 41. MINDANAO: Bukidnon. Photos
   • var. cordifolium. Australia, Borneo, Java, ?Lesser Sunda Isls, Moluccas, New Guinea, New Guinea, Peninsular Malaysia, Singapore, Solomon Isls, Sulawesi, Sumatra, Thailand, Vietnam, Philippines. PALAWAN.
   • var. pariens (Copel.) C.Chr., Gard. Bull. S. S. 7 (1934) 274; --Diplazium pariens (Copel.) C.Chr., Index Filic. (1905) 663; --Athyrium pariens Copel., PJS 3 (1908) 299. Borneo, Moluccas, Philippines. MINDANAO: Bukidnon, Davao Oriental, North Cotabato (Mt Apo), NEGROS.
8. Diplazium costulisorum (Copel.) C.Chr., Index Filic., Suppl. 1906-1912 (1913) 26; --Coritico & Amoroso, Nature Conservation Research 5 (2020) 81; --Athyrium costulisorum Copel., LPB 3 (1910) 815. Moluccas, Sulawesi, Philippines. MINDANAO: Bukidnon, Davao (Mt Apo), North Cotabato. Conservation status: Endangered (DAO 2017-11).
9. Diplazium cumingii (C.Presl) C.Chr., Index Filic. (1905) 230; --Athyrium cumingii (C.Presl) Milde, Bot. Zeitung (Berlin) 28 (1870) 353; --Ochlogramma cumingii C.Presl, Abh. Kön. Böhm. Ges. Wiss. 5(6) (Epim. Bot.) (1851) 94. Borneo, Solomon Isls, Sulawesi, Philippines. LUZON. Photos
10. Diplazium deltoideum (C.Presl) C.Presl, Tent. Pterid. (1836) 114; --Athyrium deltoideum (C.Presl) Milde, Bot. Zeitung (Berlin) 28 (1870) 353; --Asplenium deltoideum C.Presl, Rel. Haenk. 1 (1825) 47. Sulawesi, Philippines. LUZON.
11. Diplazium dilatatum Blume, Enum. Pl. Javae 2 (1828) 194; --Athyrium dilatatum (Blume) Milde, Bot. Zeitung (Berlin) 28 (1870) 353. --Diplazium davaoense Copel., PJS 1 (Suppl. 2) (1906) 151; --Athyrium davaoense (Copel.) Copel., PJS 3 c (1908) 295. --Diplazium dolichosorum Copel., PJS 1 (Suppl. 2) (1906) 151; --Athyrium dolichosorum (Copel.) Copel., PJS 3 c (1908) 294. Australia, Bangladesh, Bismarck Arch, Borneo, ?Cambodia, China, India, Japan, Java, Java, Laos, Lesser Sunda Isls, Moluccas, Myanmar, Nepal, New Caledonia, New Guinea, Pacific, Peninsular Malaysia, Solomon Isls, Sulawesi, Sumatra, Taiwan, Thailand, Vietnam, Philippines. MINDANAO: Bukidnon, PALAWAN. Photos
12. Diplazium doederleinii (Luerss.) Makino, Bot. Mag. (Tokyo) 13 (1899) 15; --Asplenium doederleinii Luerss., Engl. Jahrb. 4 (1883) 355. --Athyrium irayense Copel., PJS 81 (1952) 39. Bhutan, China, India, Japan, Myanmar, Nepal, Ryukyu Isls, Taiwan, Thailand, Vietnam, Philippines.
13. Diplazium egenolfioides M.G.Price, Brit. Fern Gaz. 10 (1973) 258. LUZON: Luzon (Mt Makiling, several collections from this type locality), Nueva Vizcaya (Sta Fe, mtns N of Imugan, c. 1200m, L.L.Co 1640 (MICH, PNH, PUH)). Endemic to the Philippines. Conservation status: Endangered (DAO 2017-11).
14. Diplazium esculentum (Retz.) Sw., J. Bot. (Schrader) 1801 (1803) 312; --Athyrium esculentum (Retz.) Copel., PJS 3 (1908) 295; --Hemionitis esculenta Retz., Observ. Bot. 6 (1791) 38. --Asplenium manilense Spreng., Syst. Veg. (ed. 16) 4 (1827) 89. Andaman Isls, Australia, Bangladesh, Bhutan, Bismarck Arch, Borneo, Cambodia, China, India, Japan, Java, Laos, Lesser Sunda Isls, Moluccas, Myanmar, Nepal, New Guinea, Nicobar Isls, Pakistan, Peninsular Malaysia, Ryukyu Isls, Singapore, Solomon Isls, Sri Lanka, Sulawesi, Sumatra, Taiwan, Thailand, Tibet, Vietnam, Philippines. Photos
15. Diplazium forbesii (Baker) C.Chr., Index Filic. (1905) 232; --Asplenium forbesii Baker, Ann. Bot. 4 (1891) 312. Borneo, Java, Sumatra, Philippines. PALAWAN. Photos
16. Diplazium fraxinifolium C.Presl, Reliq. Haenk. 1 (1825) 49; --Athyrium fraxinifolium (C.Presl) Milde, Bot. Zeitung (Berlin) 28 (1870) 353; --Type: Haenke s.n. (PRC, lecto), Luzon. Borneo, India, New Guinea, Peninsular Malaysia, Sulawesi, ?Vietnam, Philippines. LUZON, MINDANAO: Bukidnon, PALAWAN. Photos
17. Diplazium fructuosum Copel., PJS 1 (Suppl. 2) (1906) 150; --Athyrium fructuosum (Copel.) Copel., PJS 3 c (1908) 294. Endemic to the Philippines. MINDANAO: Bukidnon.
18. Diplazium fuliginosum (Hook.) M.G.Price, Brit. Fern Gaz. 10 (1973) 260; --Asplenium fuliginosum Hook., Sp. Fil. 3 (1860) 120. --Diplazium longissimum (Copel.) C.Chr., Ind. Suppl. 3 (1934) 74; --Athyrium longissimum Copel., PJS 38 (1929) 139. Bismarch Arch, Borneo, New Guinea, Philippines. LEYTE.
19. Diplazium geophilum (Copel.) Alderw., Bull. Jard. Bot. Buitenzorg Ser. 2, 7 (1912) 12; --Athyrium geophilum Copel., PJS 3c (1908) 296. --Athyrium ramosii Copel., PJS 38 (1929) 140. Moluccas, Sulawesi, Philippines. LUZON, NEGROS.
20. Diplazium griffithii T.Moore, Index Fil. (1861) 330.. --Athyrium platyphyllum Copel., PJS 3 (1908) 292. --Diplazium mearnsii Hieron., Hedwigia 59 (1917) 338. China, India, Japan, Ryukyus Isls, Vietnam, Philippines. Photos
21. Diplazium irigense (Copel.) M.G.Price, Contrib. Univ. Mich. Herb. 17 (1990) 273; --Athyrium irigense Copel., PJS 81 (1952) 38. Moluccas, Philippines. LUZON.
22. Diplazium lomariaceum (Christ) M.G.Price, Gard. Bull. Sing. 36 (1983) 27; --Asplenium lomariaceum Christ, Verh. Naturf. Ges. Basel 11 (1895) 229. --Diplazium altum (Copel.) C.Chr., Index Filic., Suppl. 3 (1934) 72; --Athyrium altum Copel., PJS 38 (1929) 138. --Diplazium merrillii Copel., PJS 2 c (1907) 128; --Athyrium merrillii (Copel.) Copel., PJS 3 (1908) 300. Borneo, Moluccas, New Guinea, Sulawesi, Sumatra, Philippines. MINDANAO: Bukidnon. Photos
23. Diplazium magnificum (Copel.) M.G.Price, 1982; --Athyrium magnificum Copel., PJS 81 (1952) 39. Moluccas, Philippines. NEGROS.
24. Diplazium maximum (D.Don) C.Chr., Index Filic. 4 (1905) 235; --Asplenium maximum D.Don, Prodr. Fl. Nepal. (1825) 8. --Diplazium macrosorum (Copel.) M.G.Price, Contrib. Herb. Univ. Mich. 17 (1990) 273; --Athyrium macrosorum Copel., LPB 3 (1910) 815. ?Afghanistan, Bhutan, China, India, Japan, Myanmar, Nepal, Pakistan, Sulawesi, Tibet, Vietnam, Philippines. MINDANAO. Conservation status: Other Threatened Species (DAO 2017-11).
    
25. Diplazium melanopodium Fée, Mém. Foug. 8 (1857) 85; --Athyrium melanopodium (Fée) Copel., PJS 81 (1952) 40. Endemic to the Philippines. Photos
26. Diplazium mollifrons (C.Chr.) M.G.Price, Contr. Univ. Michigan Herb. 16 (1987) 195; --Athyrium mollifrons C.Chr., LPB 9 (1933) 3153. Endemic to the Philippines. LUZON.
27. Diplazium muricatum (Mett.) Alderw., Malayan Ferns (1909) 829; --Athyrium muricatum (Mett.) Copel., PJS 7 c (1912) 61; --Asplenium muricatum Mett., Ann. Mus. Bot. Lugd.-Bat. 2 (1866) 239. Java, Philippines.
28. Diplazium oligosorum Copel., PJS 2 c (1907) 128; --Athyrium oligosorum (Copel.) Copel., PJS 3 c (1908) 293. Endemic to the Philippines. MINDORO.
29. Diplazium pallidum (Blume) T.Moore, Index Filicum (1861) 333; --Athyrium pallidum Blume, Enum. Fil. (1828) 177. --Diplazium crenato-serratum (Blume) T.Moore, Index Fil. 121 (1859) 325; --Athyrium crenato-serratum (Blume) Milde, Bot. Zeitung (Berlin) 28 (1870) 353; --Asplenium crenatoserratum Blume, Enum. Fil. (1828) 177. Australia, Java, Lesser Sunda Isls, Moluccas, New Guinea, Peninsular Malaysia, Solomon Isls, Sulawesi, Sumatra, Philippines. MINDANAO: Bukidnon, PALAWAN. Photos
30. Diplazium petiolare C.Presl, Abh. Kön. Böhm. Ges. Wiss. 5(6) (Epim. Bot.) (1851) 86; --Athyrium petiolare (C.Presl) Milde, Bot. Zeitung (Berlin) 28 (1870) 354. Borneo, Sumatra, Philippines. PALAWAN.
31. Diplazium phanerotis Kunze, Bot. Zeitung (Berlin) 4 (1846) 443; --Hovenkamp, Gard. Bull. Singapore 71 (2019) 63; --Type: from Java. --Diplazium cultratum C.Presl, Epimel. Bot. (1851) 84; --Athyrium cultratum (C.Presl) Milde, Bot. Zeitung (Berlin) 28 (1870) 353. Java, ?Moluccas, Peninsular Malaysia, Singapore, Philippines. BILIRAN, LUZON: Camarines Norte, Laguna, Quezon. Photos
    
32. Diplazium pinatubicum M.G.Price, Contrib. Herb. Univ. Michigan 17 (1990) 273. Endemic to the Philippines.
33. Diplazium polypodioides Blume, Enum. Pl. Jav. (1828) 194. --Athyrium blumei (Bergsm.) Copel., PJS 3 (1908) 294; --Asplenium blumei Bergsm., Cat. hort. Rheno-Traj. (1857). --Diplazium ophiodontum (Copel.) C.Chr., Index Filic., Suppl. Tertium pro Annis 1917-1933 (1934) 75; --Athyrium ophiodontum Copel., PJS 46 (1931) 214. --Diplazium asperum Blume, Enum. Pl. Javae 2 (1828) 195; --Athyrium asperum (Blume) Milde, Bot. Zeitung (Berlin) 28 (1870) 353. ?Bangladesh, Borneo, Cambodia, China, India, Java, Laos, Lesser Sunda Isls, Moluccas, ?Myanmar, New Guinea, Peninsular Malaysia, Seychelles, Singapore, ?Solomon Isls, Sri Lanka, Sulawesi, Sumatra, Thailand, Vietnam, Philippines.
34. Diplazium proliferum (Lam.) Thouars, Esquisse Fl. Tristan D'Acugna (1808) 35; --Callipteris prolifera (Lam.) Bory, Voy. Îles Afrique 1 (1804) 283; --Asplenium proliferum Lam., Encycl. 2 (1786) 307. --Diplazium accedens Blume, Enum. Pl. Javae 2 (1828) 192. Africa, Andaman Isls, Australia, Bismarck Arch, Borneo, Indian Ocean, Java, Lesser Sunda Isls, Moluccas, New Guinea, Pacific, Peninsular Malaysia, Solomon Isls, Sulawesi, Sumatra, Taiwan, Thailand, Philippines. POLILLO: PNBG 184 Sotalbo (PUH). Photos
35. Diplazium propinquum (Copel.) Alderw., Mal. Ferns Suppl. (1917) 259; --Athyrium propinquum Copel., LPB 5 (1914) 1683. Borneo, Philippines. MINDANAO: Agusan. Conservation status: Endangered (DAO 2017-11).
36. Diplazium pseudocyatheifolium Rosenst., Meded. Rijks-Herb. 31 (1917) 4; --Athyrium pseudocyatheifolium (Rosenst.) Copel., FFP 3 (1960) 395. --Athyrium ebenirachis Copel., LPB 9 (1920) 3110. Endemic to the Philippines. Photos
37. Diplazium pseudosylvaticum Panigrahi, Phytologia 31 (1975) 255. --Diplazium allantoideum M.G.Price, Contr. Univ. Michigan Herb. 16 (1987) 195. Java, Singapore, Sulawesi, Philippines.
    
38. Diplazium sibuyanense (Copel.) Alderw., Bull. Buit. 2, 11 (1913) 9; --Athyrium sibuyanense Copel., LPB 4 (1911) 1152. Endemic to the Philippines. SIBUYAN. Conservation status: Other Threatened Species (DAO 2017-11). Photos
39. Diplazium sorzogonense (C.Presl) C.Presl, Tent. Pterid. (1836) 114; --Athyrium sorzogonense (C.Presl) Milde, Bot. Zeitung (Berlin) 28 (1870) 354; --Type: Haenke s.n. (PRC, lecto), Luzon. Borneo, Java, Moluccas, New Guinea, Northern Marianas, Peninsular Malaysia, Singapore, Sulawesi, Sumatra, Thailand, Vietnam, Philippines. MINDANAO: Bukidnon, PALAWAN. Photo
40. Diplazium sylvaticum (Bory) Sw., Syn. Fil. (1806) 92; --Athyrium sylvaticum (Bory) Milde, Bot. Zeit. (Berlin) (1870) 354; --Callipteris sylvatica Bory, Voy. Îles Afrique 1 (1804) 282. --Diplazium woodwardioides (C.Presl) Holttum, Novit. Bot. 1968 (1969) 30; --Brachysorus woodwardioides C.Presl, Abh. Kön. Böhm. Ges. Wiss. 5(6) (Epim. Bot.) (1851) 70. --Allantodia pinnata Blanco, Fl. Filip., ed. 2 (1845) 571. Australia, Bangladesh, Borneo, India, Java, Lesser Sunda Isls, Mauritius, Moluccas, Myanmar, New Guinea, Pacific, Palau Isls, Peninsular Malaysia, Sri Lanka, ?Sulawesi, Sumatra, Thailand, Philippines.
    
41. Diplazium symmetricum (Copel.) M.G.Price, Gard. Bull. Sing. 36 (1983) 26; --Athyrium symmetricum Copel., PJS 81 (1861) 41; --Type: Edano PNH 15148 (MICH, lecto), Samar: Mt Cabayanan. Endemic to the Philippines. SAMAR.
42. Diplazium tenuifolium (Copel.) Lellinger, Am. Fern J. 74 (1954) 57; --Athyrium tenuifolium Copel., PJS 40 (1929) 302. Solomon Isls, Philippines. MINDANAO.
43. Diplazium tomentosum Blume, Enum. Fil. (1828) 192. Borneo, Java, Moluccas, Myanmar, Peninsular Malaysia, Singapore, Sumatra, Thailand, Vietnam, Philippines. MINDANAO.
44. Diplazium whitfordii Copel., PJS 1 (Suppl. 2) (1906) 150; --Athyrium whitfordii (Copel.) Copel., PJS 3 c (1908) 296. Endemic to the Philippines. LUZON.
45. Diplazium williamsii Copel., PJS 1 (Suppl. 2) (1906) 150; --Athyrium williamsii (Copel.) Copel., PJS 3 c (1908) 297. --Athyrium banahaoense Copel., PJS 38 (1929) 139; --Diplazium banahaoense (Copel.) C.Chr., Index Filic., Suppl. Tertium pro Annis 1917-1933 (1934) 72. Endemic to the Philippines. LUZON, MINDANAO.
46. Diplazium xiphophyllum (Baker) Copel., Index Filic. (1905) 241; --Athyrium xiphophyllum (Baker) Copel., Sarawak Mus. Journ. 2 (1917) 379; --Asplenium xiphophyllum Baker, J. Bot. (1879) 40. Borneo, Java, Moluccas, Peninsular Malaysia, Sumatra, Thailand, Philippines. PALAWAN.

Possibly present:

1. Diplazium dameriae Pic.Serm., Webbia 45 (1991) 357. Andaman Isls, Australia, Bismarck Arch, Java, Moluccas, New Guinea, Pacific, Peninsular Malaysia, Solomon Isls, ?Sulawesi, ?Philippines.
2. Diplazium decompositum (Copel.) Parris, Kew Bull. 41 (1986) 69; --Athyrium decompositum Copel., Univ. Calif. Publ. Bot. 18 (1942) 222. Moluccas, New Guinea, ?Philippines. ?MINDANAO.
3. Diplazium integrifolium Blume, Enum. Fil. (1828) 190. Borneo, India, Java, Peninsular Malaysia, Sumatra, Thailand, Vietnam, ?Philippines.
4. Diplazium lobbianum Moore, Ind. Fil. (1861) 331. Java, New Guinea, ?Vietnam, ?Philippines.
5. Diplazium petri Tardieu, Aspl. du Vietnam 67 (1932) t.9. China, Laos, Ryukyu Isls, Taiwan, Thailand, Vietnam, ?Philippines.
6. Diplazium vestitum C.Presl, Epimel. Bot. (1851) 87; --Athyrium vestitum (C.Presl) Milde, Bot. Zeitung (Berlin) 28 (1870) 354. Borneo, Moluccas. ?Philippines. Conservation status: Other Threatened Species (DAO 2017-11).

Dryoathyrium Ching = Deparia

Lunathyrium Koidz. = Deparia

Blechnaceae

Austroblechnum Gasper & V.A.O.Dittrich

1. Austroblechnum melanocaulon (Brack.) Gasper & V.A.O.Dittrich, Phytotaxa 275 (2016) 203; --Blechnum melanocaulon (Brack.) T.C.Chambers & P.A.Farrant, Blumea 46 (2001) 318; --Lomaria melanocaulon Brack., U. S. Expl. Exp. 16 (1854) 122. Andaman Isls, Bismarck Arch, Borneo, Fiji, India, Java, New Guinea, Nicobar Isls, Peninsular Malaysia, Solomon Isls, Sulawesi, Sumatra, Vanuatu, Philippines. MINDANAO: North Cotabato, NEGROS. Photos
   • ssp. melanocaulon. Bismarck Arch, Borneo, Fiji, New Guinea, Sulawesi, Philippines.
   • ssp. pallens (T.C.Chambers & P.A.Farrant) ined.; --Blechnum melanocaulon ssp. pallens T.C.Chambers & P.A.Farrant, Blumea 46 (2001) 321. Andaman Isls, India, Java, Nicobar Isls, Peninsular Malaysia, Solomon Isls, Sumatra, Vanuatu, Philippines. NEGROS.
2. Austroblechnum patersonii (R.Br.) Gasper & V.A.O.Dittrich, Phytotaxa 275 (2016) 203; --Blechnum patersonii (R.Br.) Mett., Fil. Lips. (1856) 64; --Stegania patersonii R.Br., Prodr. (1810) 152. Australia, Lesser Sunda Isls, New Guinea, Pacific, Philippines. MINDANAO: Bukidnon.

Blechnopsis C.Presl

1. Blechnopsis finlaysoniana (Wall. ex Hook. & Grev.) C.Presl, Abh. Kön. Böhm. Ges. Wiss. 5(6) (Epim. Bot.) (1851) 116; --Blechnum finlaysonianum Wall. ex Hook. & Grev., Icon. Fil. (1831) t.225. Andaman Isls, Borneo, Java, Moluccas, Moluccas, New Guinea, Nicobar Isls, Peninsular Malaysia, Singapore, Sulawesi, Sumatra, Thailand, Philippines. PALAWAN. Photos
2. Blechnopsis orientalis (L.) C.Presl, Abh. Kön. Böhm. Ges. Wiss. 5(6) (Epim. Bot.) (1851) 117; --Blechnum orientale L., Sp. Pl. (1753) 1077. Andaman Isls, Australia, Bangladesh, Bhutan, Bismarck Arch, Borneo, Cambodia, China, India, Japan, Laos, Lesser Sunda Isls, Moluccas, Myanmar, Nepal, New Caledonia, New Guinea, Nicobar Isls, Pacific, Palau Isls, Peninsular Malaysia, Ryukyu Isls, Singapore, Solomon Isls, Sri Lanka, Sulawesi, Sumatra, Taiwan, Thailand, Tibet, Vietnam, Philippines. MINDANAO: Bukidnon, PALAWAN. Photos

Blechnum L. = Austroblechnum, Blechnopsis, Diploblechnum, Oceaniopteris, Parablechnum (Philippines)

Brainea J.Sm.

1. Brainea insignis (Hook.) J.Sm., Cat. Ferns Gard. Kew (1856) 5; --Bowringia insignis Hook., J. Bot. Kew Gard. Misc. 5 (1853) 237. Cambodia, China, India, Laos, Lesser Sunda Isls, Myanmar, Peninsular Malaysia, Sumatra, Taiwan, Thailand, Vietnam, Philippines. LUZON: Laguna (photos), MINDORO. Conservation status: Endangered (DAO 2017-11). Photos

Cranfillia Gasper & V.A.O.Dittrich

1. Cranfillia vulcanica (Blume) Gasper & V.A.O.Dittrich, Phytotaxa 275 (2016) 208; --Blechnum vulcanicum (Blume) Kuhn, Ann. Lugd.-Bat. 4 (1869) 284; --Lomaria vulcanica Blume, Enum. Fil. (1828) 202. Bismarck Arch, Borneo, Java, Lesser Sunda Isls, Moluccas, New Guinea, Pacific, Peninsular Malaysia, Solomon Isls, Sulawesi, Sumatra, Philippines. LUZON, MINDANAO: Bukidnon. Photos

Possibly present:

1. Cranfillia fluviatilis (R.Br.) Gasper & V.A.O. Dittrich, Phytotaxa 275 (2016) 207; --Stegania fluviatilis R.Br., Prod. (1810) 152. Australia, Borneo, Sulawesi, New Guinea, New Zealand, Pacific, Sumatra, ?Philippines. Photos

Diploblechnum Hayata

1. Diploblechnum fraseri (A.Cunn.) De Vol in H.L.Li et al., Fl. Taiwan 1 (1975) 153; --Coritico & Amoroso, Nature Conservation Research 5 (2020) 81; --Blechnum fraseri (A.Cunn.) Luerss., Flora (1876) 292; --Lomaria fraseri A.Cunn., Comp. Bot. Mag. 2 (1836) 364. Bismarck Arch, Borneo, Moluccas, New Guinea, Pacific, Peninsular Malaysia, Sulawesi, Sumatra, Taiwan, Philippines. LUZON, MINDANAO: Bukidnon, Misamis Occidental, MINDORO. Conservation status: Vulnerable (DAO 2017-11 as Blechnum fraseri). Photos

Oceaniopteris Gasper & Salino

1. Oceaniopteris egregia (Copel.) Gasper & Salino, Phytotaxa 275 (2016) 214; --Coritico & Amoroso, Nature Conservation Research 5 (2020) 81; --Blechnum egregium Copel., Fragm. Fl. Philip. (1905) 187. Borneo, New Guinea, Solomon Isls, Philippines. MINDANAO: Bukidnon, Davao Oriental, Misamis Occidental, North Cotabato, PALAWAN. Conservation status: Vulnerable (DAO 2017-11 as Blechnum egregium). Photos

Parablechnum L.

1. Parablechnum revolutum (Alderw.) Gasper & Salino, Phytotaxa 275 (2016) 217; --Lomaria revoluta Alderw., Nova Guinea 14 (1924) 31. New Guinea, Philippines. MINDANAO.
   
2. Parablechnum vestitum (Blume) Gasper & Salino, Phytotaxa 275 (2016) 217; --Blechnum vestitum (Blume) Kuhn, Ann. Lugd.-Bat. 4 (1869) 284; --Lomaria vestita Blume, Enum. Fil. (1828) 203. ?Blechnum capense auct. non (L.) Schltdl.; --Coritico et al., PJS 149 (2020) 776. Bismarck Arch, Borneo, Java, ?Lesser Sunda Isls, Moluccas, Pacific, Peninsular Malaysia, Sulawesi, Sumatra, Philippines. MINDANAO: Bukidnon, PALAWAN. Photos

Stenochlaena J.Sm.

1. Stenochlaena areolaris (Harr.) Copel., PJS 2 c (1908) 406; --Lomaria areolaris Harr., J. Linn. Soc. 16 (1877) 28. New Guinea, Philippines. LUZON.
2. Stenochlaena cumingii Holttum, Amer. Fern J. 61 (1971) 122. Moluccas, New Guinea, Sulawesi, Philippines.
3. Stenochlaena milnei Underw., Bull. Torr. Bot. Club 33 (1906) 38. --Stenochlaena juglandifolia C.Presl, Epimel. Bot. (1851) 164. Bismarck Arch, Moluccas, New Guinea, Solomon Isls, Philippines. Photos
4. Stenochlaena palustris (Burm.f.) Bedd., Ferns Brit. Ind. Suppl. (1876) 26; --Polypodium palustre Burm.f., Fl. Indica (1768) 234. --Stenochlaena laurifolia C.Presl, Abh. Kön. Böhm. Ges. Wiss. 5(6) (Epim. Bot.) (1851) 164. Australia, Bangladesh, Bismarck Arch, Borneo, Cambodia, China, India, Java, Laos, Lesser Sunda Isls, Moluccas, Nepal, New Guinea, Nicobar Isls, Pacific, Palau Isls, Peninsular Malaysia, Singapore, Solomon Isls, Sri Lanka, Sulawesi, Sumatra, Thailand, Vietnam, Philippines. PALAWAN. Photos

Woodwardia Sm.

1. Woodwardia prolifera Hook. & Arn., Bot. Beechey Voy. (1841) 275; --Woodwardia orientalis var. prolifera (Hook. & Arn.) Ching. China, Japan, Ryukyu Isls, Taiwan, Philippines. LUZON: Bataan. Photos
2. Woodwardia unigemmata (Makino) Nakai, Bot. Mag. (Tokyo) 39 (1925) 103; --Woodwardia radicans var. unigemmata Makino, Jap. Bot. 2 (1918) 7. Bhutan, China, India, Jammu & Kashmir, Japan, Java, Myanmar, Nepal, New Guinea, Pakistan, Taiwan, Tibet, Vietnam, Philippines. LUZON.  Photos

Cibotiaceae 

Cibotium Kaulf.

1. Cibotium barometz (L.) J.Sm., London J. Bot. 1 (1842) 437; --Polypodium barometz L., Sp. Pl. 2 (1753) 1092. Assam to S China, W Malesia. LUZON: Mountain Province (near Bontoc). Known in the Philippines by a single collection, CAHUP 16635 Pancho. Conservation status: Endangered (DAO 2017-11). Photos
2. Cibotium cumingii Kunze, Farrnkräuter 1 (1841) 64. Borneo, Philippines. LUZON, MINDANAO, MINDORO. Conservation status: Endangered (DAO 2017-11). Photos

Cyatheaceae

Cyathea Sm.

1. Cyathea acuminata Copel., PJS 81 (1952) 15; --Alsophila acuminata (Copel.) R.M.Tryon, Contr. Gray Herb. 200 (1970) 32. Endemic to the Philippines. PANAY, SAMAR. Conservation status: Endangered (DAO 2017-11).
2. Cyathea apoensis Copel., LPB 3 (1910) 802; --Alsophila apoensis (Copel.) R.M.Tryon, Contr. Gray Herb. 200 (1970) 32; --Coritico & Amoroso, Nature Conservation Research 5 (2020) 81. --Cyathea lobata Copel., PJS 81 (1952) 15. Endemic to the Philippines. LUZON: Aurora, Laguna, Quezon, MINDANAO: Davao, Misamis Occidental, North Cotabato, MINDORO, NEGROS. Conservation status: Endangered (DAO 2017-11).
3. Cyathea atropurpurea Copel., PJS 3 (1909) 354; --Gymnosphaera atropurpurea (Copel.) Copel., Gen. Fil. (1947) 98; --Alsophila atropurpurea (Copel.) C.Chr., Index Filic., Suppl. 1 (1913) 4; --Coritico & Amoroso, Nature Conservation Research 5 (2020) 81. Endemic to the Philippines. MINDANAO: Agusan del Norte, Bukidnon, Davao, Misamis Occidental, North Cotabato, MINDORO. Conservation status: Endangered (DAO 2017-11).
4. Cyathea biliranensis Copel., PJS 84 (1955) 162. Endemic to the Philippines.
5. Cyathea binuangensis Alderw., Bull. Jard. Bot. Buitenzorg, sér. 3, 2 (1920) 136; --Sphaeropteris binuangensis (Alderw.) R.M.Tryon, Contr. Gray Herb. 200 (1970) 21. Endemic to the Philippines. LUZON, TAWI-TAWI. Conservation status: Endangered (DAO 2017-11).
6. Cyathea callosa Christ, Bull. Herb. Boissier, sér. 2, 6 (1906) 1008; --Alsophila callosa (Christ) R.M.Tryon, Contr. Gray Herb. 200 (1970) 33. --Cyathea foxworthyi Copel., PJS 3 c (1909) 355. --Cyathea camiguinensis Copel., PJS 81 (1952) 16, as 'camaguinensis'. Endemic to the Philippines. CAMIGUIN, CATANDUANES, LUZON: Bataan, Benguet, Camarines Sur, Kalinga, Laguna, Quezon, PALAWAN. Conservation status: Vulnerable (DAO 2017-11).
7. Cyathea caudata (J.Sm.) Copel., PJS 1 Suppl 2 (1906) 144; --Alsophila caudata J.Sm. ex Hook., Sp. Fil. 1 (1844) 52. --Cyathea arborescens Copel., PJS 84 (1955) 162. --Cyathea brevipes Copel., PJS 81 (1952) 18. --Cyathea dupaxensis Copel., 46 (1931) 211. Endemic to the Philippines. LUZON, MINDANAO, MINDORO, NEGROS, PALAWAN. Conservation status: Vulnerable (DAO 2017-11).
8. Cyathea christii Copel., PJS 1 (Suppl. 2) (1906) 144; --Alsophila hermannii R.M.Tryon, Contr. Gray Herb. 290 (1970) 34; --Coritico & Amoroso, Nature Conservation Research 5 (2020) 81. Endemic to the Philippines. MINDANAO: Bukidnon, Davao Oriental, North Cotabato, NEGROS. Conservation status: Endangered (DAO 2017-11).
9. Cyathea cinerea Copel., LPB 5 (1913) 1681; --Alsophila cinerea (Copel.) R.M.Tryon, Contr. Gray Herb. 290 (1970) 33. Endemic to the Philippines. MINDANAO: Agusan, Misamis Occidental. Conservation status: Vulnerable (DAO 2017-11).
10. Cyathea contaminans (Wall. ex Hook.) Copel., PJS 4 c (1909) Bot. 60; --Polypodium contaminans Wall. ex Hook. --Sphaeropteris glauca (Blume) R.M.Tryon, Contr. Gray Herb. 200 (1970) 21; --Coritico & Amoroso, Nature Conservation Research 5 (2020) 81; --Chnoophora glauca Blume, Enum. Pl. Javae (1828) 243. --Cyathea clementis Copel., PJS 4 c (1909) 59. New Guinea to India. Throughout the Philippines. MINDANAO: Bukidnon, Davao Oriental, Misamis Occidental, North Cotabato. Usually common in open places and light forests from near sea-level to 1300m. Conservation status: Endangered (DAO 2017-11). Photos
11. Cyathea curranii Copel., PJS 3 c (1909) 356; --Sphaeropteris curranii (Copel.) R.M.Tryon, Contr. Gray Herb. 200 (1970) 22. Endemic to the Philippines. LUZON: Quezon. Conservation status: Critically Endangered (DAO 2017-11). Photos
12. Cyathea edanoi Copel., PJS 46 (1931) 211; --Alsophila edanoi (Copel.) R.M.Tryon, Contr. Gray Herb. 200 (1970) 33. Endemic to the Philippines. LUZON: Cagayan, Quezon. Conservation status: Endangered (DAO 2017-11). Photos  
13. Cyathea elmeri (Copel.) Copel., PJS 4 c (1909) 54; --Sphaeropteris elmeri (Copel.) R.M.Tryon, Contr. Gray Herb. 200 (1970) 22; --Coritico & Amoroso, Nature Conservation Research 5 (2020) 81. --Cyathea argyrolepis Copel., PJS 81 (1952) 17. --Cyathea dimorphotricha Copel., LPB 5: (1913) 1681. Moluccas, Sulawesi, Philippines. BILIRAN, LEYTE, MINDANAO: Bukidnon, Davao Oriental, Misamis Occidental, North Cotabato, NEGROS, PALAWAN. Conservation status: Vulnerable (DAO 2017-11).
14. Cyathea fenicis Copel., PJS 3 c (1909) 354; --Alsophila fenicis (Copel.) C.Chr., Index Filic., Suppl. 1 (1913) 5. Moluccas, Taiwan, Philippines. LUZON. Conservation status: Vulnerable (DAO 2017-11).
15. Cyathea ferruginea Christ, PJS C 2 (1907) 181; --Alsophila ferruginea (Christ) R.M.Tryon, Contr. Gray Herb. 200 (1970) 33. --Cyathea ferrugineoides Copel., PJS 81 (1952) 15. Endemic to the Philippines. BALABAC, NEGROS, PALAWAN. Conservation status: Endangered (DAO 2017-11).
16. Cyathea fuliginosa (Christ) Copel., PJS 4 c (1909) 43; --Alsophila fuliginosa Christ, Bull. Herb. Boissier 6 (1898) 138; --Coritico & Amoroso, Nature Conservation Research 5 (2020) 81. --Cyathea bicolor Copel., LPB 3 (1910) 804. --Cyathea dura Copel., PJS 81 (1952) 13. --Cyathea lanaensis Christ., PJS 3 (1908) 271. --Cyathea lepidigera Copel., PJS 81 (1952) 16. --Cyathea mindanensis (Christ) Copel., PJS 4 (1909) 34; --Alsophila mindanensis Christ, Nat. Pfl. 1 (1899) 136. --Cyathea squamicosta Copel., PJS 46 (1931) 212. --Cyathea sulitii Copel., PJS 81 (1952) 18. --Cyathea warihon Copel., LPB 5 (1913) 1680. Endemic to the Philippines. BILIRAN, LEYTE, LUZON, MINDANAO: Bukidnon, North Cotabato, NEGROS. Conservation status: Vulnerable (DAO 2017-11).
17. Cyathea halconensis Christ, PJS 3 c (1908) 270; --Alsophila halconensis (Christ) R.M.Tryon, Contr. Gray Herb. 200 (1970) 34. --Cyathea mearnsii Copel., PJS 3 (1909) 356. --Cyathea melanophlebia Copel., PJS 38 (1929) 131. Endemic to the Philippines. LUZON: Benguet, Mountain Province, Nueva Vizcaya, MINDORO. Conservation status: Vulnerable (DAO 2017-11).
18. Cyathea heterochlamydea Copel., LPB 2 (1908) 418; --Alsophila heterochlamydea (Copel.) R.M.Tryon, Contr. Gray Herb. 200 (1970) 34; --Coritico & Amoroso, Nature Conservation Research 5 (2020) 81. --Cyathea merrillii Copel., 46 (1931) 212. Endemic to the Philippines. LUZON, MINDANAO: North Cotabato, NEGROS, PANAY. Conservation status: Vulnerable (DAO 2017-11). Photos  
19. Cyathea integra J.Sm. ex Hook., Icon. Pl. (1844) 638; --Sphaeropteris integra (J.Sm.) R.M.Tryon, Contr. Gray Herb. 200 (1970) 22. --Cyathea arguta Copel., PJS 38 (1929) 133. --Cyathea breviloba Copel., PJS 81 (1952) 13. --Cyathea bulusanensis Copel., LPB 9 (1920) 3109. Moluccas, Philippines. BILIRAN, CATANDUANES, LUZON, MINDANAO, MINDORO, NEGROS, PANAY, POLILLO, SAMAR. Photos  
20. Cyathea latebrosa (Wall. ex Hook.) Copel., PJS 4 c (1909) 52; --Alsophila latebrosa Wall. ex Hook., Sp. Fil. 1 (1844) 37. Borneo, Cambodia, China, Laos, Lesser Sunda Isls, Peninsular Malaysia, Singapora, Sumatra, Thailand, Vietnam, Philippines.
21. Cyathea latipinnula Copel., LPB 4 (1911) 1149; --Alsophila latipinnula (Copel.) R.M.Tryon, Contr. Gray Herb. 200 (1970) 34. Endemic to the Philippines. SIBUYAN. Conservation status: Critically Endangered (DAO 2017-11).
22. Cyathea lepifera (J.Sm. ex Hook.) Copel., PJS 4 (1909) 40; --Sphaeropteris lepifera (Hook.) R.M.Tryon, Contr. Gray Herb. 200 (1970) 21; --Coritico & Amoroso, Nature Conservation Research 5 (2020) 81. --Alsophila lepifera J.Sm. ex Hook., Sp. Fil. 1 (1844) 54. --Cyathea calocoma (Christ) Copel., PJS 4 (1909) 53; --Alsophila calocoma Christ, PJS 2 (1907) 182.  --Cyathea pteridioides Copel., PJS 56 (1935) 98. China, Hong Kong, New Guinea, Ryukyus Isls, Taiwan, Philippines. BABUYAN, LUZON, MINDANAO: Bukidnon, Misamis Occidental, North Cotabato, MINDORO, PANAY. Conservation status: Endangered (DAO 2017-11).
23. Cyathea loheri Christ, Bull. Herb. Boiss. 2, 6 (1906) 1007; --Alsophila loheri (Christ) R.M.Tryon, Contr. Gray Herb. 200 (1970) 32. --Cyathea campbellii Copel., PJS 38 (1929) 132. --Cyathea fructuosa Copel., LPB 2 (1908) 419. --Cyathea indusiosa Copel., PJS 81 (1952) 14. --Cyathea matthewii (Christ) Domin, Pteridophyta (1929) 263; --Alsophila matthewii Christ, J. Linn. Soc., Bot. 39 (1909) 213. --Cyathea mitrata Copel., PJS 3 (1909) 354. Borneo, Taiwan, Philippines. LUZON, MINDANAO: Bukidnon, NEGROS. Photos
24. Cyathea lurida (Blume) Copel., PJS 4 (1901) 45; --Alsophila lurida (Blume) Hook., Sp. Fil. 1 (1844) 55; --Chnoophora lurida Blume, Enum. Pl. Javae (1828) 244. --Gymnosphaera melanorachis (Copel.) Copel., Gen. Fil. (1947) 98; --Alsophila melanorachis Copel., PJS 2 (1907) 146. Java, Peninsular Malaysia, Sulawesi, Sumatra, Philippines. MINDANAO: Bukidnon, MINDORO.
25. Cyathea masapilidensis Copel., PJS 81 (1952) 17; --Alsophila masapilidensis (Copel.) R.M.Tryon, Contr. Gray Herb. 200 (1970) 34. Endemic to the Philippines. LUZON: Benguet, ?MINDANAO: ?Bukidnon. Conservation status: Endangered (DAO 2017-11).
26. Cyathea microchlamys Holttum, Kew Bull. 16 (1962) 58; --Alsophila microchlamys (Holtt.) R.M.Tryon, Contr. Gray Herb. 200 (1970) 35. Endemic to the Philippines. CATANDUANES. Conservation status: Critically Endangered (DAO 2017-11).
27. Cyathea negrosiana Christ, PJS 2 (1907) 181; --Alsophila negrosiana (Christ) R.M.Tryon, Contr. Gray Herb. 200 (1970) 35; --Coritico & Amoroso, Nature Conservation Research 5 (2020) 81. Endemic to the Philippines. BILIRAN, LEYTE, MINDANAO: Misamis Occidental, NEGROS. Conservation status: Vulnerable (DAO 2017-11).
28. Cyathea obliqua Copel., LPB 4 (1911) 1150; --Gymnosphaera obliqua (Copel.) Copel., Gen. Fil. (1947) 98; --Sphaeropteris obliqua (Copel.) R.M.Tryon, Contr. Gray Herb. 200 (1970) 22. Endemic to the Philippines. SIBUYAN. Conservation status: Critically Endangered (DAO 2017-11). Photos
29. Cyathea philippinensis Baker, Ann. Bot. 5 (1891) 186; --Copel., FFP 2 (1960) 205; --Holttum, Fl. Males. ser. 2, 1 (1963) 149; --Sphaeropteris philippinensis (Baker) R.M.Tryon, Contr. Gray Herb. 200 (1970) 23; --Type: From a cultivated plant originating from the Philippines, ex Hort. Veitch., Feb-1878, cult. Kew (K). --Cyathea bicolana Copel., LPB 9 (1920) 3108; --Copel., FFP 2 (1960) 204; --Type: Elmer 16528  (L, U, US, iso), Luzon: Sorsogon Prov., Mt Bulusan, Jul-1916. --Cyathea bontocensis Copel., PJS 46 (1931) 209; FFP 2 (1960) 205; --Type: Vanoverbergh 813, Luzon: Mountain Province. --Cyathea heteroloba Copel., PJS 38 (1929) 134; FFP 2 (1960) 204; --Type: BS 45633 Ramos & Edaño (NY, iso), Luzon: Quirino Prov., Mt Alzapan, 1500m, May/Jun-1925. --Cyathea ramosiana Alderw., Bull. Jard. Bot. Buitenzorg Ser. 2i, 2 (1920) 137; --Copel., FFP 2 (1960) 206; --Type: BS 23411 Ramos (L, US, iso), Luzon: Sorsogon Prov., Jul/Aug-1915. Endemic to the Philippines. LUZON: Mountain Province, Quirino (Mt Alzapan), Aurora (Mt Camatis), Quezon (Mt Banahaw), Camarines Sur (Mt Isarog, apparently common), Sorsogon (Mt Bulusan), MINDORO: Mindoro Oriental (Mt Halcon). Montane forests, 800-1500m. Conservation status: Vulnerable (DAO 2017-11). Photos  
30. Cyathea polypoda Baker in Stapf, Trans. Linn. Soc. Lond. Bot. 4 (1894) 250; --Sphaeropteris polypoda (Baker) R.M.Tryon, Contr. Gray Herb. 200 (1970) 23. --Gymnosphaera dinagatensis Copel., PJS 81 (1952) 19. --Gymnosphaera glabra sensu Copel., Fern Fl. Philip. 2 (1960) 235, non Blume. Borneo, Peninsular Malaysia, Philippines. MINDANAO: Bukidnon, PANAY.
31. Cyathea recommutata Copel., PJS 4 (1909) 36; --Alsophila commutata Mett., Ann. Mus. Bot. Lugduno-Batavi 1 (1863) 53; --Type: from Malaysia. Borneo, Malay Peninsula, Sumatra, Philippines. MINDANAO: Bukidnon. Montane forests c. 1600-1700m. Photos
32. Cyathea robinsonii Copel., PJS 6 (1911) 145; --Sphaeropteris robinsonii (Copel.) R.M.Tryon, Contr. Gray Herb. 200 (1970) 23. --Cyathea pseudalbizzia Copel., PJS 38 (1929) 135. Endemic to the Philippines. LUZON: Aurora, Ilocos Norte, Isabela, Quezon, MINDANAO: Bukidnon, TAWI-TAWI. Conservation status: Vulnerable (DAO 2017-11).
33. Cyathea rufopannosa Christ, PJS 2 (1907) 180; --Alsophila rufopannosa (Christ) R.M.Tryon, Contr. Gray Herb. 200 (1970) 36; --Coritico & Amoroso, Nature Conservation Research 5 (2020) 81. Endemic to the Philippines. MINDANAO: Bukidnon, Misamis Occidental, Zamboanga. Conservation status: Endangered (DAO 2017-11).
34. Cyathea setulosa Copel., PJS 81 (1952) 14; --Alsophila setulosa (Copel.) R.M.Tryon, Contr. Gray Herb. 200 (1970) 36. Endemic to the Philippines. LUZON: Isabela, Kalinga, Quezon. Conservation status: Vulnerable (DAO 2017-11).
35. Cyathea sibuyanensis Copel., LPB 4 (1911) 1150; --Sphaeropteris sibuyanensis (Copel.) R.M.Tryon, Contr. Gray Herb. 200 (1970) 23. Endemic to the Philippines. SIBUYAN. Conservation status: Critically Endangered (DAO 2017-11). SIBUYAN. Photos
36. Cyathea suluensis Baker, J. Bot. 17 (1879) 65; --Sphaeropteris suluensis (Baker) R.M.Tryon, Contr. Gray Herb. 200 (1970) 23. --Cyathea sessilipinnula Copel., PJS 38 (1929) 134. ?Moluccas, Philippines. BASILAN, MINDANAO.
37. Cyathea trichophora Copel., PJS 6 c (1911) Bot. 363; --Gymnosphaera trichophora (Copel.) Copel., Gen. Fil. (1947) 99; --Sphaeropteris trichophora (Copel.) R.M.Tryon, Contr. Gray Herb. 200 (1970) 23. Borneo, Philippines. Widespread, incl. PALAWAN.
38. Cyathea tripinnata Copel., PJS 1 (1906) Suppl. 251; --Sphaeropteris tripinnata (Copel.) R.M.Tryon, Contr. Gray Herb. 200 (1970) 23. --Cyathea densisora Alderw., Bull. Jard. Bot. Buitenzorg, sér. 3, 2 (1920) 138. --Cyathea leucostegia Copel., PJS 38 (1929) 130. --Cyathea leytensis Copel., PJS 38 (1929) 131. Borneo, Java, Moluccas, Peninsular Malaysia, Sulawesi, Philippines. Widespread. MINDANAO: Bukidnon.
39. Cyathea zamboangana Copel., PJS 30 (1926) 325; --Sphaeropteris zamboangana (Copel.) R.M.Tryon, Contr. Gray Herb. 200 (1970) 24. --Cyathea urdanetensis Copel., PJS 38 (1929) 132. Endemic to the Philippines. MINDANAO: Agusan, Zamboanga, NEGROS, ?SAMAR. Conservation status: Endangered (DAO 2017-11). Photos

Possibly present:

1. Cyathea squamulata (Blume) Copel., PJS 4 c (1909) Bot. 37; --Gymnosphaera squamulata Blume, Enum. Pl. Javae (1828) 243; --Sphaeropteris squamulata (Blume) R.M.Tryon, Contr. Gray Herb. 200 (1970) 23. Borneo, Java, Moluccas, Peninsular Malaysia, Singapore, Sumatra, ?Philippines.

Gymnosphaera Blume = Cyathea

Cystopteridaceae

Acystopteris Nakai

1. Acystopteris tenuisecta (Blume) Tagawa, Acta Phytotax. Geobot. 7 (1938) 73; --Cystopteris tenuisecta (Blume) Mett., Ann. Lugd. Bat. 1 (1864) 241; --Aspidium tenuisectum Blume, Enum. Pl. Javae 2 (1828) 170. --Ctenitis tenerifrons (Christ) Copel., Gen. Fil. (1947) 125; --Hypolepis tenerifrons Christ, PJS 3 c (1908) 274. Bhutan, Borneo, China, India, Japan, Java, Moluccas, Myanmar, Nepal, Nepal, New Guinea, Peninsular Malaysia, Sulawesi, Taiwan, Thailand, Tibet, Vietnam, Yakushima Isl, Philippines.

Currania Copel. = Gymnocarpium

Cystopteris Bernhardi = Acystopteris (Philippines)

Gymnocarpium Newman

1. Gymnocarpium oyamense (Baker) Ching, Contr. Biol. Lab. Sci. Soc. China 9 (1933) 40; --Polypodium oyamense Baker, J. Bot. 15 (1877) 366. --Currania gracilipes Copel., PJS 4 (1909) 112. Bhutan, China, India, Japan, Moluccas, Nepal, New Guinea, Taiwan, Tibet, Philippines. LUZON. Conservation status: Vulnerable (DAO 2017-11).

Davalliaceae

Araiostegia Copel. = Davallia

Davallia J.Sm.

1. Davallia angustata Wall. ex Hook. & Grev., Ic. Fil. (1831) t.231; --Humata angustata (Wall. ex Hook. & Grev.) J.Sm., J. Bot. 3 (1841) 416. --Humata microsora Alderw., PJS 7 c (1912) 55. Borneo, ?Cambodia, Palau Isls, Peninsular Malaysia, Singapore, Sulawesi, Sumatra, Thailand, Vietnam, Philippines. MINDANAO, PALAWAN.
2. Davallia brevipes Copel., PJS 1 (Suppl. 2) (1906) 147; --Hovenkamp & de Joncheere, Blumea 33 (1988) 408. Bismarck Arch, Moluccas, New Guinea, Solomon Isls, Sulawesi, Philippines. MINDANAO.
3. Davallia denticulata (Burm.f.) Mett. ex Kuhn, Fil. Deck. (1867) 27; --Adiantum denticulatum Burm.f., Fl. Ind. (1768) 236. Andaman Isls, Australia, Bismarck Arch, Borneo, Cambodia, China, India, Laos, Lesser Sunda Isls, Myanmar, New Guinea, Nicobar Isls, Peninsular Malaysia, Singapore, Sri Lanka, Sulawesi, Sumatra, Thailand, Vietnam, Philippines. MINDANAO: Bukidnon, PALAWAN. Photos
4. Davallia divaricata Blume, Enum. Fil. (1828) 237. Borneo, Cambodia, China, Hong Kong, India, Java, Laos, Lesser Sunda Isls, Moluccas, Myanmar, New Guinea, Peninsular Malaysia, Sulawesi, Sumatra, Thailand, Vietnam, Philippines.
5. Davallia embolostegia Copel., PJS 1 (Suppl. 2) (1906) 147. Borneo, Micronesia, Moluccas, Palau Isls, ?Samoa, Sulawesi, Sumatra, Thailand, Philippines. Conservation status: Other Threatened Species (DAO 2017-11). Photos
6. Davallia epiphylla (G.Forst.) Spreng., Schrad., J. Bot. 1799 (1799) 271; --Trichomanes epiphyllum G.Forst., Prod. (1786) 85. --Davallia denticulata var. elata (G.Forst.) Mett. ex Kuhn, Miq., Ann. Mus. Bot. Lugd.-Bat. 4 (1869) 288. Bismarck Arch, Moluccas, New Caledonia, New Guinea, Pacific, Solomon Isls, Sulawesi, Philippines. Conservation status: Other Threatened Species (DAO 2017-11; as Davallia denticulata var. elata).
   
7. Davallia falcinella C.Presl, Reliq. Haenk. 1 (1825) 66; --Trogostolon falcinellus (C.Presl) Copel., PJS 84 (1927) 251. Endemic to the Philippines. LEYTE, LUZON, MINDANAO, NEGROS, SAMAR.
8. Davallia heterophylla Sm., Mém. Acad. Turin 5 (1793) 415; --Humata heterophylla (Sm.) Desv., Prodr. (1825) 323. --Humata ophioglossa Cav., Descr. Pl. 1 (1802) 273. Bismarck Arch, Borneo, Cambodia, Java, Lesser Sunda Isls, Moluccas, New Guinea, Nicobar Isls, Pacific, Peninsular Malaysia, Singapore, Solomon Isl, Sulawesi, Sumatra, Thailand, Vietnam, Philippines. PALAWAN. Photos
9. Davallia hymenophylloides (Blume) Kuhn, Ann. Mus. Bot. Lugd.-Bat. 4 (1869) 286; --Araiostegia hymenophylloides (Blume) Copel., PJS 34 (1927) 241; --Aspidium hymenophylloides Blume, Enum. Pl. Jav. (1828) 172. Borneo, India, Java, Laos, Lesser Sunda Isls, Peninsular Malaysia, Sri Lanka, Sulawesi, Sumatra, Thailand, Philippines. LUZON, MINDANAO: Bukidnon, MINDORO. Photos
10. Davallia lorrainii Hance, Ann. Sci. Nat. 5, 5 (1866) 254; --Davallia trichomanoides var. lorrainii (Hance) Holttum, Rev. Fl. Malaya ed. 1, 2 (1955) 361. Borneo, Cambodia, India, Myanmar, Peninsular Malaysia, Sulawesi, Sumatra, Thailand, Vietnam, Philippines. Conservation status: Other Threatened Species (DAO 2017-11).
    
11. Davallia pectinata Sm., Mém. Acad. Turin 5 (1793) 415; --Humata pectinata (Sm.) Desv., Mem. Soc. Linn. Paris 6, 2 (1827) 323. Australia, Bismarck Arch, Borneo, ?Cambodia, Java, Moluccas, New Caledonia, New Guinea, Nicobar Isls, Pacific, Palau Isls, Peninsular Malaysia, Singapore, Solomon Isls, Sulawesi, Sumatra, Taiwan, Thailand, Vietnam, Philippines. PALAWAN. Photos
12. Davallia pubescens C.W.Chen, Sol Amazing (2017) 450; --Davallodes hirsutum (J.Sm.) Copel., PJS 3 c (1908) 33, nom. illeg., auct. non Sw.; --Leucostegia hirsuta J.Sm., J. Bot. 3 (1841) 416. --Davallodes grammatosorum Copel., PJS 3c (1908) 34. --Davallodes gymnocarpum Copel., PJS 3c (1908) 34. --Davallodes laxum Copel., PJS 34 (1927) 246. --Davallodes congestum Copel., PJS 34 (1927) 247. Bismarck Arch, Borneo, New Guinea, Solomon Isls, Sulawesi, Sumatra, Philippines. MINDANAO: Bukidnon. Photos
    
13. Davallia pusilloides (Copel.) Parris, Fern Gaz. 20 (2018) 305; --Humata pusilloides Copel. ?ex Alderw., Bull. Jard. Bot. Buitenzorg Ser. 2, 28 (1918) 26. Borneo, Philippines. MINDANAO. 
    
14. Davallia repens (L.f.) Kuhn, Fil. Deck. (1867) 27, nom. cons.; --Humata repens (L.f.) Diels in Engl. & Prant, Nat. Pflanzenfam. 14 (1899) 209; --Adiantum repens L.f., Suppl. Pl. (1781) 446. --Humata intermedia C.Chr., Index Filic. (1905) 353. --Humata obtusata Alderw., Bull. Jard. Bot. Buitenzorg Ser. 2, I (1911) 8. --Humata vestita (Blume) T.Moore, Index (1857) 92. Africa, Andaman Isls, Australia, Borneo, Cambodia, China, India, Indian Ocean, Japan, Java, Laos, Lesser Sunda Isls, Moluccas, Myanmar, New Guinea, Pacific, Palau Isls, Peninsular Malaysia, Ryukyu Isl., Singapore, Sri Lanka, Sulawesi, Sumatra, Taiwan, Thailand, Vietnam, Philippines. MINDANAO: Bukidnon, PALAWAN. Photos
15. Davallia sessilifolia Blume, Enum. Fil. (1828) 231. Borneo, Java, Lesser Sunda Isls, Moluccas, New Guinea, Pacific, Solomon Isls, Sulawesi, Sumatra, Philippines. LUZON.
16. Davallia solida (G.Forst.) Sw., Schrad. J. Bot. 1800 (1801) 87; --Coritico & Amoroso, Nature Conservation Research 5 (2020) 81. --Davallia elmeri Copel., LPB 9 (1920) 3107. --Davallia robinsonii Copel., PJS 30 (1926) 326. Photos
    
    • var. solida. Andaman Isls, Australia, Bismarck Arch, Borneo, Cambodia, China, India, Java, Lesser Sunda Isls, Moluccas, Myanmar, New Caledonia, New Guinea, Nicobar Isls, Pacific, Palau Isls, Peninsular Malaysia, Singapore, Solomon Isls, Sri Lanka, Sulawesi, Sumatra, Taiwan, Thailand, Vietnam, Philippines. Epiphytic or epilithic, sea-level to c. 600m. MINDANAO: Bukidnon, Davao Oriental, Misamis Occidental, North Cotabato. Conservation status: Other Threatened Species (DAO 2017-11).
17. Davallia trichomanoides Blume, Enum. Fil. (1828) 238. Bhutan, China, India, Japan, Java, Laos, Lesser Sunda Isls, Moluccas, Myanmar, New Guinea, North Korea, Palau Isls, Peninsular Malaysia, Ryukyu Isls, South Korea, Sulawesi, Sumatra, Taiwan, Thailand, Vietnam, Philippines. Photos
18. Davallia wagneriana Copel., Fragm. Fl. Philipp. (1905) 180. Borneo, Moluccas, Sulawesi, Sumatra, Philippines. MINDANAO: Bukidnon. Photos

Davallodes (Copel.) Copel. = Davallia

Humata Cav. = Davallia

Trogostolon Copel. = Davallia falcinella

Dennstaedtiaceae 

including Hypolepidaceae, Monachosoraceae, and Pteridiaceae  

Dennstaedtia Bernh.

1. Dennstaedtia articulata Copel., LPB 2 (1908) 396. Endemic to the Philippines. Conservation status: Endangered (DAO 2017-11).
2. Dennstaedtia dennstaedtioides (Copel.) Copel., PJS 2 c (1907) 126; --Microlepia dennstaedtioides Copel., PJS 1 (Suppl. 2) (1906) 148. Sulawesi, Philippines. LUZON, MINDANAO.
3. Dennstaedtia elmeri Copel., LPB 1 (1907) 233. Endemic to the Philippines. LUZON.
4. Dennstaedtia fusca Copel., PJS 81 (1952) 4. Endemic to the Philippines. PANAY. Conservation status: Endangered (DAO 2017-11).
5. Dennstaedtia glabrata (Ces.) C.Chr., Index Filic. (1905) 217; --Dicksonia glabrata Ces., Rend. Ac. Napoli 16 (1877) 24. ?Bismarck Arch, Fiji, New Guinea, Solomon Isls, Sulawesi, Philippines. MINDANAO, NEGROS.
6. Dennstaedtia hooveri Christ., PJS 2 c (1907) 169. Moluccas, Philippines. MINDANAO. Photos  
7. Dennstaedtia macgregorii Copel., PJS 81 (1952) 4. Endemic to the Philippines. LUZON.
8. Dennstaedtia merrillii Copel., PJS 2 c (1907) 126. Endemic to the Philippines. MINDORO.
9. Dennstaedtia philippinensis Copel., LPB 9 (1920) 3107. Endemic to the Philippines. LUZON.
10. Dennstaedtia resinifera (Blume) Mett. ex Kuhn, Ann. Mus. Bot. Lugduno-Batavi 4 (1869) 290; --Cheilanthes resinifera Blume, Enum. Fil. (1828) 138. --Dennstaedtia cuneata (J.Sm.) T.Moore, Index Fil. (1857) 97; --Sitolobium cuneatum J.Sm., J. Bot. 3 (1841) 418. Borneo, Java, Moluccas, Philippines.
11. Dennstaedtia rufidula C.Chr. in C.Chr. & Holltum, Gard. Bull. Sing. 7 (1934) 226. Borneo, Philippines.
12. Dennstaedtia scandens (Blume) T.Moore, Index Fil. (1861) 307; --Dicksonia scandens Blume, Enum. Pl. Javae (1828) 240. Lesser Sunda Isls, Moluccas, New Guinea, Solomon Isls, Sulawesi, Tahiti, Taiwan, Philippines. MINDANAO: Bukidnon. Photos
13. Dennstaedtia smithii (Hook.) T.Moore, Index Fil. (1861) 308; --Dicksonia smithii Hook., Sp. Fil. 1 (1844) 80. Sulawesi, Taiwan, Philippines.
14. Dennstaedtia williamsii Copel., PJS 1 (Suppl. 2) (1906) 148. Endemic to the Philippines. MINDANAO.
15. Dennstaedtia zeylanica (Sw.) Zink ex Fraser-Jenk. & Kandel, Ferns and Fern-allies of Nepal 1 (2015) 161; --Dicksonia zeylanica Sw., Schrad. Journ. 1800 (1801) 91. --Dennstaedtia scabra (Wall. ex Hook.) T.Moore, Index Fil. (1861) 307; --Dicksonia scabra Wall. ex Hook., Sp. Fil. 1 (1844) 80. Bhutan, Borneo, China, India, Japan, Laos, Myanmar, Nepal, North Korea, Peninsular Malaysia, South Korea, Sri Lanka, Sulawesi, Sumatra, Taiwan, Thailand, Tibet, Vietnam, Philippines. Photos

Histiopteris (Agardh) J.Sm.

1. Histiopteris incisa (Thunb.) J.Sm., Hist. Fil. (1875) 295. --Histiopteris montana Copel., Fragm. Fl. Philipp. (1905) 193. Africa, Americas, Australia, Bhutan, Borneo, Cambodia, China, India, Indian Ocean, Japan, Lesser Sunda Isls, Moluccas, Myanmar, Nepal, New Caledonia, Peninsular Malaysia, Ryukyu Isls, Singapore, Solomon Isls, Sri Lanka, Sulawesi, Taiwan, Thailand, Tibet, Vietnam, Philippines. MINDANAO: Bukidnon. Photos  

Hypolepis Bernh.

1. Hypolepis alpina (Blume) Hook., Sp. Fil. 2 (1852) 63; --Cheilanthes alpina Blume, Enum. Pl. Javae 2 (1828) 138. Borneo, Java, New Guinea, Pacific, Ryukyu Isls, Taiwan, Philippines.
2. Hypolepis bamleriana Rosenst., Feddes Repert. 10 (1912) 325. Borneo, Micronesia, Moluccas, New Guinea, Sulawesi, Philippines.
3. Hypolepis malesiana Brownsey, Blumea 32 (1987) 248. New Guinea, ?Sumatra, Philippines.
4. Hypolepis polypodioides (Blume) Hook., Sp. Fil. 2 (1852) 63; --Cheilanthes polypodioides Blume, Enum. Pl. Javae 2 (1828) 139. Bangladesh, Bhutan, Cambodia, China, India, Laos, Myanmar, Nepal, Peninsular Malaysia, Taiwan, Tibet, Vietnam, Philippines. MINDANAO: Bukidnon.
5. Hypolepis resistens (Kunze) Hook., Sp. Fil. 2. (1852) 64; --Cheilanthes resistens Kunze, Linn. 24 (1851) 275. --Hypolepis glandulifera Brownsey & Chinnock, J. Adelaide Bot. Gard. 10 (1987) 16. Australia, Bhutan, Borneo, China, India, Myanmar, New Caledonia, New Guinea, Peninsular Malaysia, Sri Lanka, Vietnam, Philippines.
   
6. Hypolepis rugosula J.Sm., Companion Bot. Mag. 72 (1846) 8.
   
   • ssp. rudis (Kunze) Schwartsb., Acta Bot. Brasil. 28 (2014) 216; --Cheilanthes rudis Kunze, Bot. Zeit. (Berlin) (1848) 213; --Type: Cuming 140 (FI, K, RB), Luzon. Endemic to the Philippines. LUZON.
7. Hypolepis tenuifolia (G.Forst.) Bernh. ex C.Presl, Neues J. Bot. 1 (1805) 34; --Lonchitis tenuifolia G.Forst., Fl. Ins. Austr. (1786) 80. Australia, China, Lesser Sunda Isls, New Caledonia, New Caledonia, New Guinea, Pacific, Sulawesi, Taiwan, Vietnam, Philippines.

Possibly present:

1. Hypolepis pallida (Blume) Hook., Sp. Fil. 2 (1852) 64; --Cheilanthes pallida Blume, Enum. Fil. (1828) 139. Bonin Isl, ?Borneo, Peninsular Malaysia, Sumatra, Taiwan, ?Vietnam, ?Philippines. ?NEGROS. Doubtfully occurring in the Philippines (Negros: Amlan, M.G.Price 2479).
2. Hypolepis punctata (Thunb. ex Murray) Mett. ex Kuhn, Fil. Afr. (1868) 120; --Polypodium punctatum Thunb., Fl. Jap. (1784) 337. China, Ryukyus Isls, South Korea, Taiwan, ?Philippines. ?LUZON: ?Ilocos Norte (Mt  Bubungkayo, Iwatsuki et al. P375 (L)). Doubtfully occurring in the Philippines.

Microlepia C.Presl

1. Microlepia matthewii Christ, Not. syst. 1 (1909) 54. --Microlepia herbacea Ching & C.Chr. ex Tardieu & C.Chr. [Notul. Syst. (Paris) 6 (1937) 6. China, Laos, Thailand, Vietnam, Philippines. Photos
   
2. Microlepia platyphylla (D.Don) J.Sm., Lond. J. Bot. 1 (1842) 427b; --Davallia platyphylla D.Don, Prodr. Fl. Nepal. (1825) 10. Bhutan, China, India, Laos, Myanmar, Nepal, Sri Lanka, Taiwan, Thailand, Tibet, Vietnam, Philippines. Photos
3. Microlepia protracta Copel., PJS 81 (1952) 5. Endemic to the Philippines. BALUT. Conservation status: Endangered (DAO 2017-11).
4. Microlepia speluncae (L.) T.Moore, Ind. Fil. (1857) 93; --Polypodium speluncae L., Sp. Pl. 2 (1753) 1093. --Microlepia sablanensis Christ, PJS 2 c (1907) 168. --?Microlepia manilensis C.Presl. --Microlepia trichosticha J.Sm., J. Bot. (Hooker) 3 (1841) 416, ?nom. nud. Africa, Americas, Andaman Isls, Australia, Bangladesh, Bhutan, Bismarck Arch, Borneo, Cambodia, China, India, Indian Ocean, ?Japan, Laos, Lesser Sunda Isls, Moluccas, Myanmar, Nepal, Nicobar Isls, Pacific, Peninsular Malaysia, Ryukyu Isls, Singapore, Solomon Isls, Sri Lanka, Sulawesi, Sumatra, Taiwan, Thailand, Tibet, Vietnam, Philippines. MINDANAO: Bukidnon, PALAWAN. Photos  
5. Microlepia strigosa (Thunb.) C.Presl, Abh. Kön. Böhm. Ges. Wiss. 5(6) (Epim. Bot.) (1851) 95; --Trichomanes strigosum Thunb., Fl. Jap. (1784) 339. Bangladesh, Bhutan, Borneo, China, India, Japan, Java, Lesser Sunda Isls, Myanmar, Nepal, New Caledonia, Pacific, Peninsular Malaysia, Ryukyu Isls, South Korea, Sri Lanka, Sulawesi, Thailand, Vietnam, Philippines. Photos
6. Microlepia todayensis Christ, PJS 3 (1908) 272. China, Myanmar, Peninsular Malaysia, Sumatra, Vietnam, Philippines.

Possibly present:

1. Microlepia firma Mett. ex Kuhn, Linn. 36 (1869) 146. Bhutan, ?Cambodia, China, India, Myanmar, Nepal, Sri Lanka, Thailand, Tibet, Vietnam, ?Philippines.
2. Microlepia rhomboidea (Wall. ex Kunze) Prantl, Arbeiten Königl. Bot. Gart. Breslau 1 (1892) 31; --Davallia rhomboidea Wall. ex Kunze, Bot. Zeit. (Berlin) (1850) 158. Bangladesh, Bhutan, China, India, Myanmar, Nepal, Sri Lanka, Taiwan, Vietnam, ?Philippines.

Monachosorum Kunze

1. Monachosorum henryi Christ, Bull. Herb. Boiss. 6 (1898) 869. Bhutan, China, India, Myanmar, Nepal, Taiwan, Thailand, Tibet, Vietnam, Philippines. LUZON: Mountain Province (Mt Data). Photos  
2. Monachosorum subdigitatum (Blume) Kuhn, Chaetopt. (1882) 345; --Aspidium subdigitatum Blume [Enum. Pl. Jav. (1828) 171. Bhutan, Borneo, ?India, Java, Moluccas, Myanmar, New Guinea, Peninsular Malaysia, Sulawesi, Sumatra, Vietnam, Philippines.

Paesia A.St.-Hil.

1. Paesia elmeri Copel., LPB 3 (1910) 826. Peninsular Malaysia, Sumatra, Philippines. MINDANAO. Photos
2. Paesia radula (Baker) C.Chr., Index Filic. 476 1906; --Pteris radula Baker, J. Bot. (1880) 211. --Paesia luzonica Christ, PJS 3 c (1908) 275. Borneo, Java, Moluccas, New Guinea, Sulawesi, Sumatra, Taiwan, Philippines. MINDANAO: Bukidnon. Photos

Pteridium Gled. ex Scop.

1. Pteridium aquilinum (L.) Kuhn, Bot. Ost-Afrika 3 (1879) 11; --Pteris aquilina L., Sp. Pl. 2 (1753) 1075. Widespread. Photos
   • var. wightianum (Wall. ex J.Agardh) Tryon, Rhodora 43 (1941) 22; --Pteris recurvata var. wightiana Wall. ex J.Agardh., Recens. Spec. Pter. (1839) 50. --Pteridium revolutum (Blume) Nakai, Bot. Mag. (Tokyo) 39 (1925) 109; --Pteris revoluta Blume, Enum. Pl. Javae 2 (1828) 214. China, Indochina, Malesia, Australia, Philippines. MINDANAO: Bukidnon.

Dicksoniaceae

Calochlaena (Maxon) M.D.Turner & R.A.White

1. Calochlaena javanica (Blume) M.D.Turner & R.A.White, Amer. Fern J. 78 (1988) 93; --Culcita javanica (Blume) Maxon, J. Wash. Acad. Sci. 12 (1922) 456; --Dicksonia javanica Blume, Enum. Pl. Javae 2 (1828) 240. --Culcita copelandii (Christ) Maxon, J. Wash. Acad. Sci. 12 (1922) 457. Borneo, Java, Lesser Sunda Isls, Philippines. LUZON, MINDANAO: Bukidnon, NEGROS. Photos
2. Calochlaena straminea (Labill.) M.D.Turner & R.A.White, Amer. Fern J. 78 (1988) 92; --Culcita straminea (Labill.) Maxon, J. Wash. Acad. Sci. 12 (1922) 457; --Dicksonia straminea Labill., Sert. Austro-Caledon. 7 (1824) t.10. Bismarck Arch, New Caledonia, New Guinea, Pacific, Solomon Isls, Philippines. MINDANAO.

Dicksonia L’Her.

1. Dicksonia amorosoana Lehnert & Coritico, Blumea 63 (2018) 269; --Type: F.P. Coritico FPC 016 (PNH, holo; BONN, CMUH iso), Mindanao: North Cotabato, Mt Apo, Kidapawan, Mandarangan, trail to Lake Venado, 2106m, 5-May-2012. Endemic to the Philippines. MINDANAO: North Cotabato.
   
2. Dicksonia mollis Holttum, Kew Bull. 16 (1962) 64; --Coritico & Amoroso, Nature Conservation Research 5 (2020) 81; --Type: Elmer 9874 (K, holo; BM, L, MICH, MO, NY, P, U, US, iso), Negros: Negros Oriental, Dumaguete, Cuernos Mts, Apr-1908. Borneo, Philippines. MINDANAO: Bukidnon, Misamis Occidental, North Cotabato, NEGROS. Conservation status: Endangered (DAO 2017-11). Photos  

Diplaziopsidaceae

Diplaziopsis C.Chr.

1. Diplaziopsis javanica (Blume) C.Chr., Index Filic. (1905) 227; --Diplazium javanicum (Blume) Makino, Phan. Pter. Jap. Ic. I11. 1 (1901) pl. 46; --Asplenium javanicum Blume, Enum. Pl. Jav. (1828) 175. Bangladesh, Bhutan, Bismarck Arch, Borneo, China, India, Japan, Java, Lesser Sunda Isls, Moluccas, Myanmar, Nepal, New Caledonia, New Guinea, Pacific, Solomon Isls, Sri Lanka, Sulawesi, Sumatra, Taiwan, Vietnam, Philippines. MINDANAO: Bukidnon. Photos

Dipteridaceae

Cheiropleuria C.Presl

1. Cheiropleuria bicuspis (Blume) C.Presl, Abh. Königl. Böhm. Ges. Wiss., ser. 5 6 (1851) 549; --Polypodium bicuspe Blume, Enum. Pl. Javae 2 (1828) 125. Borneo, China, Japan, Java, Lesser Sunda Isls, Moluccas, New Guinea, Peninsular Malaysia, Ryukyu Isls, Sulawesi, Sumatra, Thailand, Vietnam, Philippines. MINDANAO: Bukidnon, PALAWAN. Photos  

Dipteris Reinw.

1. Dipteris conjugata Reinw., Syll. pl. 2 (1824) 3. Australia, Borneo, Cambodia, China, Java, Lesser Sunda Isls, Moluccas, New Caledonia, New Guinea, Pacific, Peninsular Malaysia, Peninsular Thailand, Ryukyu Isls, Singapore, Solomon Isls, Sulawesi, Sumatra, Taiwan, Vietnam, Philippines. MINDANAO: Bukidnon, PALAWAN. Photos  
2. Dipteris lobbiana (Hook.) Moore, Ind. Fil. (1857) 80; --Polypodium lobbianum Hook., Hooker's J. Bot. Kew Gard. Misc. 5 (1853) 309. Borneo, Peninsular Malaysia, Sulawesi, Sumatra, Philippines. NEGROS. Conservation status: Critically Endangered (DAO 2017-11). Photos

Dryopteridaceae 

Acrophorus C.Presl = Dryopteris

Acystopteris Nakai = Cystopteris

Arachniodes Blume

1. Arachniodes amabilis (Blume) Tindale, Contr. New South Wales Natl. Herb. 3 (1961) 90; --Rumohra amabilis (Blume) Ching, Sinensia 5 (1934) 41; --Aspidium amabile Blume, Enum. Pl. Javae 2 (1828) 165. China, India, Japan, Java, Lesser Sunda Isls, Myanmar, South Korea, Sri Lanka, Sulawesi, Taiwan, Vietnam, Philippines. MINDANAO: Bukidnon. Photos
2. Arachniodes carvifolia (Kunze) Ching, Acta Bot. Sin. 10 (1962) 256; --Aspidium carvifolium Kunze, Linn. 24 (1851) 292. --Arachniodes aristata auct. non (G.Forst.) Tindale. Australia, Bhutan, Borneo, China, India, Japan, Lesser Sunda Isls, Myanmar, Nepal, New Guinea, Peninsular Malaysia, Ryukyu Isls, South Korea, Sri Lanka, Sulawesi, Vietnam, Philippines.
3. Arachniodes cornucervi (D.Don) Fraser-Jenk., Taxon. Revis. Indian Subcontinental Pteridophytes (2008) 323; --Aspidium cornucervi D.Don, Prodr. Fl. Nepal. (1825) 5. Bhutan, China, India, ?Micronesia, Myanmar, Nepal, South Korea, ?Thailand, Vietnam, Philippines.
4. Arachniodes hasseltii (Blume) Ching, Acta Bot. Sin. 10 (1962) 258; --Rumohra hasseltii (Blume) Ching, Sinensia 5 (1934) 61; --Polypodium hasseltii Blume, Fl. Jav. Fil. 195 (1829) t.92. Australia, Borneo, China, India, Java, ?Lesser Sunda Isls, Moluccas, New Guinea, Pacific, Peninsular Malaysia, Ryukyu Isls, Solomon Isls, Sulawesi, Sumatra, Thailand, Tokara Isl, Vietnam, Yakushima Isl, Philippines.
5. Arachniodes tripinnata (Goldm.) Sledge, Bull. Brit. Mus. (Nat. Hist.) Bot. 5 (1973) 41; --Polystichum tripinnatum Goldm., Nova Acta 19, Suppl. 1 (1843) 463. Borneo, Java, Micronesia, Sulawesi, Philippines. PALAWAN.

Possibly present:

1. Arachniodes coniifolia (Wall. ex T.Moore) Ching, Acta Bot. Sin. 10 (1962) 257; --Lastrea coniifolia T.Moore, Ind. Fil. (1857) 88. --Rumohra wallichii Ching, Sinensia 5 ( 1934) 56. Bhutan, China, India, Myanmar, Nepal, Thailand, Vietnam, ?Philippines.
2. Arachniodes sledgei Fraser-Jenk., Taxon. Revis. Indian Subcontinental Pteridophytes (2008) 323. India, Sri Lanka, ?Philippines.

 Arthrobotrya J.Sm.

Last edited by Julie F. Barcelona, 17 June 2019

No Photos

1. Arthrobotrya articulata (J.Sm. ex Fée) J.Sm., Hist. Fil. (1875) 141; --Teratophyllum articulatum (J.Sm. ex Fée) Mett. ex Kuhn, Ann. Mus. Bot. Lugduno-Batavi 4 (1869) 297; --Polybotrya articulata J.Sm. ex Fée, Acrost. (1845) 74. Bismarck Arch, Moluccas, New Guinea, Solomon Isls, Sulawesi, Philippines. LEYTE, LUZON, MINDANAO, SAMAR.

Bolbitis Schott

1. Bolbitis angustipinna (Hayata) H.Ito, J. Jap. Bot. 14 (1938) 443; --Leptochilus angustipinnus Hayata, Icon. Pl. Formosan. 5 (1915) 297. Bhutan, China, India, Myanmar, Nepal, Sri Lanka, Sulawesi, Taiwan, Thailand, Philippines. LUZON.
2. Bolbitis appendiculata (Willd.) K.Iwats., Acta Phytotax. Geobot. 18 (1959) 48; --Egenolfia appendiculata (Willd.) J.Sm., Ferns Brit. For. (1866) 111; --Acrostichum appendiculatum Willd., Sp. Pl. 5 (1810) 114. Bangladesh, Bhutan, Cambodia, China, India, Japan, Java, Laos, Myanmar, Nicobar Isls, Peninsular Malaysia, Ryukyu Isls, Singapore, Sri Lanka, Sumatra, Taiwan, Thailand, Vietnam, Philippines.
3. Bolbitis heteroclita (C.Presl) Ching, Index Filic., Suppl. 3 (1934) 48; --Acrostichum heteroclitum C.Presl., Reliq. Haenk. 1 (1825) 15. --Bolbitis cuspidata (C.Presl) Ching, Index Filic., Suppl. 3 (1934) 48; --Nephrodium cuspidatum C.Presl., Reliq. Haenk. 1 (1825) 31. --Bolbitis pseudosculpturata (Copel.) Ching, Index Filic., Suppl. Tertium pro Annis 1917-1933 (1934) 49; --Campium pseudoscalpturatum Copel., PJS 37 (1928) 363. --Bolbitis foxworthyi (Copel.) Ching, Index Filic., Suppl. Tertium pro Annis 1917-1933 (1934) 48; --Campium foxworthyi Copel., PJS 37 (1928) 364. --Bolbitis tenuissima (Copel.) Copel., FFP 2 (1960) 259; --Campium tenuissimum Copel., PJS 37 (1928) 364. --Bolbitis membranacea (Copel.) C.Chr., Index Filic., Suppl. Tertium pro Annis 1917-1933 (1934) 198; --Campium membranaceum Copel., PJS 40 (1929) 307. --Bolbitis edanyoi Copel., PJS 81 (1952) 22. --Edanyoa difformis Copel., PJS 81 (1952) 22. Bangladesh, Borneo, China, India, Java, Moluccas, Myanmar, Nepal, New Guinea, Pacific, Peninsular Malaysia, Ryukyu Isls, Singapore, Solomon Isls, Sulawesi, Sulawesi, Sumatra, Sumatra, Taiwan, Thailand, Vietnam, Philippines. MINDANAO: Bukidnon, PALAWAN. Photos
4. Bolbitis quoyana (Gaudich.) Ching in C.Chr., Index Filic., Suppl. Tert. (1934) 49; --Acrostichum quoyanum Gaudich., Voy. Uranie (1827) 306. --Bolbitis enormis (Copel.) C.Chr., Index Filic., Suppl. Tertium pro Annis 1917-1933 (1934) 197; --Campium enorme Copel., PJS 40 (1929) 307. --Stenosemia dimorpha Copel., PJS 84 (1955) 161. Australia, Bismarck Arch, Bonin Isls, Java, Lesser Sunda Isls, Moluccas, New Guinea, Pacific, Solomon Isls, Sulawesi, Philippines.
5. Bolbitis repanda (Blume) Schott, Gen. Fil. ad (1834) t.14; --Acrostichum repandum Blume, Enum. Pl. Jav. (1828) 104. Borneo, Lesser Sunda Isls, ?Nepal, Sulawesi, Philippines. Photos
6. Bolbitis rhizophylla (Kaulf.) Hennipman, Blumea 18 (1970) 148; --Egenolfia rhizophylla (Kaulf.) Fée, Mém. Foug. 5 (1852) 48; --Gymnogramma rhizophylla Kaulf., Enum. Fil. (1824) 78. --Egenolfia intermedia (J.Sm. ex Fée) Fée, Gen. (1852) 308; --Polybotrya intermedia J.Sm. ex Fée, Mém. Foug. (1845) 76. --Egenolfia fluviatilis Copel., PJS 38 (1929) 152. Taiwan, Philippines. BALABAC, LUZON, MINDORO, PALAWAN. Photos
7. Bolbitis scalpturata (Fée) Ching, Index Filic., Suppl. 3 (1934) 50; --Heteroneuron scalpturatum Fée, Mém. Foug. (1845) 95. China, Java, Lesser Sunda Isls, Myanmar, Sulawesi, Sumatra, Taiwan, Thailand, Vietnam, Philippines. LUZON, PALAWAN. Photos
8. Bolbitis sinuata (C.Presl) Hennipman, Blumea 18 (1970) 148; --Polypodium sinuatum C.Presl, Rel. Haenk. 1 (1825) 21. --Bolbitis diversifolia (Blume) Schott, Gen. ad (1834) t.14; --Acrostichum diversifolium Blume, Enum. Pl. Jav. (1828) 103. --Bolbitis subsimplex (Fée) Ching, Index Filic., Suppl. Tertium pro Annis 1917-1933 (1934) 50; --Gymnopteris subsimplex Fée, (Acrost.) 2e mém. 83 (1845) t.40. --Bolbitis hydrophylla (Copel.) Ching, Index Filic., Suppl. Tertium pro Annis 1917-1933 (1934) 48; --Leptochilus hydrophyllus Copel., PJS 1 (Suppl. 2) (1906) 146. --Leptochilus stolonifer Christ, Bull. Herb. Boiss. 2, 6 (1906) 1004; --Type: from Luzon: Rizal prov., Angilog. Borneo, Java, Moluccas, New Guinea, Nicobar Isls, Peninsular Malaysia, Pulau Langkawi, Pulau Tioman, Sulawesi, Sumatra, Thailand, Philippines. LUZON: Rizal, PALAWAN. Photos

Hybrids

1. Bolbitis X arguta (Fée) Ching, Index Filic., Suppl. Tertium pro Annis 1917-1933 (1934) 47. Hennipman doubts whether this fern warrants a separate treatment as a hybrid; it may be close to Edanyoa difformis (=Bolbitis heteroclita).
2. Bolbitis heteroclita X Bolbitis rhizophylla Hennipman, Leid. Bot. Ser. 2 (1977) 291.
3. Bolbitis X sinuosa (Fée) Copel., FFP 2 (1960) 262. Photos

Excluded taxa

1. Bolbitis feeiana (Copel.) Fraser-Jenk. & Gandhi, Indian Fern J. 35(1-2): 114 (2018). --Bolbitis presliana (Fée) Ching, Ind. Fil., Suppl. 3 (1934) 49. Included in Copeland’s treatment in FFP 2 (1960) 260 as Bolbitis presliana. This taxon from India does not occur in the Philippines nor anywhere in Malesia.

Cyrtomium C.Presl

Last edited by Julie F. Barcelona, 20 July 2019

Photos

1. Cyrtomium caryotideum (Wall. ex Hook. & Grev.) C.Presl, Tent. Pterid. (1836) 86; --Polystichum falcatum var. caryotideum (Wall. ex Hook. & Grev.) Matsum., Index Pl. Jap. 1 (1904) 342; --Phanerophlebia caryotidea (Wall. ex Hook. & Grev.) Copel., Gen. Fil. (1947) 111; --Aspidium caryotideum Wall. ex Hook. & Grev., Ic. Fil. (1828) t.69. Bhutan, China, Hawaii, India, Japan, Myanmar, Nepal, Pakistan, South Korea, Taiwan, Tibet, Vietnam, Philippines. Philippine record based on a sterile specimen by Mearns from Mt Sto. Tomas (Copeland, FFP 2 (1960) 248).
   
2. Cyrtomium falcatum (L.f.) C.Presl, Tent. Pterid. (1836) 86; --Polystichum falcatum (L.f.) Diels, Nat. Pflanzenfam. 1 (1899) 194; --Phanerophlebia falcata (L.f.) Copel., Gen. Fil. (1947) 111; --Polypodium falcatum L.f., Suppl. Pl. (1781) 446.
   
   • ssp. falcatum. China, Japan, North Korea, Ryukyu Isls, South Korea, Taiwan, Vietnam, Philippines. Philippine record based on a Loher collection from Mt. Data [Copeland, FFP 2 (1960) 248] at the turn of the 1900s. Photos

Possibly present:

1. Cyrtomium fortunei J.Sm., Ferns Brit. For. (1866) 286; --Cyrtomium fortunei J.Sm. forma latipinna Ching, Ic. Fil. Sin. 3 (1935) t.126. China, Taiwan, Vietnam, North Korea, South Korea, India, Bhutan, Thailand, ?Philippines. Photos

 Diacalpe Blume = Dryopteris

Dryopsis Holttum & P.J.Edwards = Dryopteris

Dryopteris Adans.

1. Dryopteris alpina Rosenst., Repert. Spec. Nov. Regni Veg. 12 (1913) 173. --Stenolepia tristis (Blume) Alderw., Bull. Dept. Agric. Indes Neerl. 27 (1909) 46 t. 7; --Aspidium triste Blume, Enum. Pl. Javae 2 (1828) 169; --Type from Java. Borneo, Java, New Guinea, Sulawesi, Philippines. MINDANAO: Bukidnon, North Cotabato. 2300-2900m. Conservation status: Other Threatened Species (DAO 2017-11 as Stenolepia tristis).
   
2. Dryopteris cacaina Tagawa, Acta Phytotax. Geobot. 6 (1937) 155. Japan, Taiwan, Philippines.
3. Dryopteris chrysocoma C.Chr., Index Filic. 5 (1905) 257. Bhutan, China, India, Myanmar, Nepal, Pakistan, Tibet, Vietnam, Philippines. Conservation status: Endangered (DAO 2017-11).
4. Dryopteris cochleata (Don) C.Chr., Index Filic. 5 (1905) 258; --Nephrodium cochleatum D.Don, Prodr. Fl. Nepal. (1825) 6. Bangladesh, Bhutan, China, India, Java, Lesser Sunda Isls, Myanmar, Nepal, Pakistan, Peninsular Malaysia, Thailand, Vietnam, Philippines.
5. Dryopteris formosana (Christ) C.Chr., Index Filic. 5 (1905) 266; --Aspidium formosanum Christ, Bull. Herb. Boissier, sér. 2, 4 (1904) 615. China, Japan, Ryukyu Isls, South Korea, Taiwan, Philippines. Photos
6. Dryopteris hasseltii (Blume) C.Chr., Index Filic. 5 (1905) 269; --Polypodium hasseltii Blume, Fl. Jav. Fil. (1829) 195. Australia, Borneo, China, India, Java, ?Lesser Sunda Isls, Moluccas, New Guinea, Pacific, Peninsular Malaysia, Ryukyu Isls, Solomon Isls, Sulawesi, Sumatra, Taiwan, Thailand, Vietnam, Philippines.
   
7. Dryopteris hendersonii (Bedd.) C.Chr., Index Filic. 5 (1905) 270; --Chen et al., Phytotaxa 443 (2020) 132; --Nothoperanema hendersonii (Bedd.) Ching, Acta Phytotax. Sin. 11 (1966) 28; --Lastrea hendersonii Bedd., Suppl. Ferns S. Ind. (1876) 17; --Type: from India. --Ctenitis mearnsii Copel., PJS 81 (1952) 24; --Type: Mearns BS 4216 (MICH, lecto), Luzon: Benguet, Haight’s in the Oaks, 1907. Bhutan, Borneo, China, India, Japan, Myanmar, Nepal, Taiwan, Thailand, Vietnam, Philippines. LUZON: Benguet, MINDANAO: Bukidnon. Terrestrial in mossy forest, 2100-2750m. Photos
8. Dryopteris hirtipes (Blume) Kuntze, Revis. Gen. Pl. 2 (1891) 813; --Aspidium hirtipes Blume, Enum. Pl. Javae 2 (1828) 148. Borneo, India, Java, Lesser Sunda Isls, Myanmar, Pacific, Peninsular Malaysia, Sri Lanka, Sulawesi, Sumatra, Thailand, Vietnam, Philippines. Photos
9. Dryopteris manipurensis (Bedd.) C.Chr., Index Filic. (1905) 276; --Dryopsis manipurensis (Bedd.) Holttum & P.J.Edwards, Kew Bull. 41 (1986) 200; --Ctenitis manipurensis (Bedd.) Ching, Bull. Fan Mem. Inst. Biol. Bot. 8 (1938) 297; --Polypodium manipurense Bedd., J. Bot. (1888) 235. --Ctenitis copelandii (Christ) Copel., Gen. Fil. (1947) 124; --Dryopteris copelandii Christ, PJS 2 c (1907) 216. India, Nepal, New Guinea, Philippines.
10. Dryopteris nodosa (C.Presl) Li Bing Zhang, Taxon 61 (2012) 1208; --Acrophorus nodosus C.Presl, Tent. Pterid. (1836) 94. --Acrophorus stipellatus T.Moore, Gard. Chron. (1854) 135. Bismarck Arch, Borneo, Fiji, Japan, Lesser Sunda Isls, Myanmar, Solomon Isls, Sulawesi, Sumatra, Thailand, Vietnam, Philippines. MINDANAO: Bukidnon. Photos
11. Dryopteris peranema Li Bing Zhang, Taxon 61 (2012) 1211. --Dryopteris zhuweimingii Li Bing Zhang, Taxon 61(6) (2012) 1211; --Peranema cyatheoides D.Don var. luzonicum (Copel.) Ching & SH Wu, Acta Phytotax. Sin. 21 (1983) 373; --Peranema luzonica Copel., PJS 4 (1909) 111; --Type: Curran, Merritt & Zschokke 16280 (US, lecto), Luzon: Benguet: Mt Pulag, Jan-1909. Bhutan, China, India, Myanmar, Nepal, Taiwan, Tibet, Vietnam, Philippines. LUZON: Benguet. Conservation status: Critically Endangered (DAO 2017-11), as Dryopteris zhuweimingii). Photos
    
12. Dryopteris permagna M.G.Price, Gard. Bull. Singapore 30 (1977) 249. Endemic to the Philippines. LUZON: Benguet, Kalinga, Mountain Province, MINDANAO: South Cotabato. Conservation status: Endangered (DAO 2017-11).
13. Dryopteris polita Rosenst., Repert. Spec. Nov. Regni Veg. 13 (1914) 218. Borneo, Cambodia, China, Japan, New Guinea, Peninsular Malaysia, Sulawesi, Sumatra, Taiwan, Thailand, Vietnam, Philippines. Conservation status: Vulnerable (DAO 2017-11).
14. Dryopteris pseudocaenopteris (Kunze) Li Bing Zhang, Taxon 61 (2012) 1209; --Diacalpe pseudocaenopteris Kunze, Bot. Zeitung (Berlin) 4 (1846) 457. --Peranema aspidioides (Blume) Mett.; --Diacalpe aspidioides Blume. Bhutan, Borneo, China, India, Java, Laos, Lesser Sunda Isls, Myanmar, Nepal, New Guinea, Peninsular Malaysia, Sri Lanka, Sulawesi, Sumatra, Taiwan, Thailand, Tibet, Vietnam, Philippines. Photos
15. Dryopteris pulvinulifera (Bedd.) Kuntze, Revis. Gen. Pl. 2 (1891) 213; --Lastrea pulvinulifera Bedd., Ferns Brit. India (1870) t.333. Bhutan, China, India, Nepal, Sri Lanka, Philippines.
16. Dryopteris purpurascens (Blume) Christ, PJS 2 c (1907) 213; --Aspidium purpurascens Blume, Enum. Fil. (1828) 169. Borneo, Sulawesi, Philippines. Photos
17. Dryopteris sparsa (D.Don) Kuntze, Revis. Gen. Pl. 2 (1891) 813; --Nephrodium sparsum D.Don, Prodr. Fl. Nepal. (1825) 6. Photos
    •  ssp. rectipinnula Fraser-Jenk., Taxon. Revis. Indian Subcontinental Pteridophytes (2008) 320. Australia, Bhutan, Borneo, ?Cambodia, China, India, Japan, Java, ?Laos, Lesser Sunda Isls, Micronesia, Myanmar, Nepal, New Guinea, Peninsular Malaysia, ?Solomon Isls, Sulawesi, Sumatra, Thailand, Tibet, Vietnam, Philippines. MINDANAO: Bukidnon, PALAWAN.
18. Dryopteris subarborea (Baker) C.Chr., Index Filic. (1905) 295; --Polypodium subarboreum Baker, J. Linn. Soc. 24 (1887) 259. Borneo, Fiji, New Guinea, Peninsular Malaysia, Sulawesi, Sumatra, Philippines.
19. Dryopteris subtriangularis (C.Hope) C.Chr., Index Filic. 5 (1905) 296; --Nephrodium subtriangulare C.Hope, J. Bot. 28 (1890) 327. --Dryopteris uropinna M.G.Price, Gard. Bull. Singapore 30 (1977) 244. --Dryopteris erythrosora sensu Copel. (1960), non (Eaton) Kuntze. China, India, Myanmar, Ryukyu Isls, Taiwan, Thailand, Tibet, Vietnam, Philippines. Photos
20. Dryopteris varia (L.) Kuntze, Revis. Gen. Pl. 2 (1891) 814; --Polypodium varium L., Sp. Pl. 2 (1753) 1090. Bonin Isl, China, India, Japan, Myanmar, Ryukyu Isls, South Korea, Sulawesi, Taiwan, Vietnam, Volcano Isl, Philippines. Photos
21. Dryopteris wallichiana (Spreng.) Hyl., Bot. Not. (1953) 352; --Aspidium wallichianum Spreng., Syst. Veg. 4 (1827) 104. Photos
    • ssp. wallichiana. --Dryopteris paleacea (Lag. ex Sw.) C.Chr., Amer. Fern J. 1 (1911) 94, nom. illeg., auct. non Hand.-Mazz. Americas, Bhutan, Borneo, China, Gough Isl, Hawaii, India, Japan, Java, Myanmar, Nepal, New Guinea, Sri Lanka, Sulawesi, Taiwan, Turkey, Vietnam, Philippines.

 Edanyoa Copel. = Bolbitis heteroclita

 Egenolfia Schott = Bolbitis

Elaphoglossum Schott ex J.Sm.

1. Elaphoglossum angulatum (Blume) T.Moore, Index Fil. 5 (1857) 351; --Acrostichum angulatum Blume, Enum. Pl. Javae 2 (1828) 101. Africa, Borneo, China, India, Indian Ocean, Java, Lesser Sunda Isls, Moluccas, New Guinea, Sri Lanka, Sulawesi, Sumatra, Taiwan, Thailand, Vanuatu, Vietnam, Philippines.
2. Elaphoglossum apoense Holttum, Blumea 14 (1966) 320. New Guinea, Sulawesi, Philippines. MINDANAO.
3. Elaphoglossum blumeanum (Fée) J.Sm., Ferns Brit. For. (1866) 106. --Elaphoglossum copelandii Christ, PJS 2 (1907) 176. --Elaphoglossum blumeanum var. philippinense Christ ex Holttum., Flora Malesiana Series II, 1 (1978) 311. --?Elaphoglossum petiolatum auct. non (Sw.) Urb.; --Coritico et al., PJS 149 (2020) 778. Bismarck Arch, Borneo, Borneo, Java, Moluccas, Peninsular Malaysia, Solomon Isls, Sulawesi, Sumatra, Philippines. LUZON, MINDANAO: Bukidnon. Photos
4. Elaphoglossum calanasanicum Holttum, Kalikasan 3 (1974) 196. Endemic to the Philippines. LUZON. Only known from the type.
5. Elaphoglossum callifolium (Blume) T.Moore, Index Fil. (1857) 7; --Acrostichum callifolium Blume, Enum. Pl. Javae 2 (1828) 100. Australia, Borneo, Java, Lesser Sunda Isls, Moluccas, New Guinea, Pacific, Sulawesi, Sumatra, Taiwan, ?Vietnam, Philippines. MINDANAO: Bukidnon. Photos
6. Elaphoglossum luzonicum Copel., LPB 1 (1907) 235. --Elaphoglossum elmeri Copel., LPB 3 (1910) 849. Borneo, New Guinea, Peninsular Malaysia, Taiwan, Philippines. LUZON, MINDANAO, NEGROS, PANAY. Photos
7. Elaphoglossum melanostictum (Blume) T.Moore, Index Fil. (1862) 361; --Acrostichum melanostictum Blume, Fl. Javae (1829) 26. --Elaphoglossum basilanicum Copel., PJS 11 (1916) 41. Borneo, Java, Peninsular Malaysia, Sumatra, Thailand, Philippines. BASILAN, MINDANAO, NEGROS, SAMAR.
8. Elaphoglossum negrosensis Holttum, Blumea 14 (1966) 323. --Elaphoglossum angulatum sensu Copel. (1960), pro parte, non (Blume) T.Moore. Endemic to the Philippines. NEGROS. Only known from the type.
9. Elaphoglossum ophioglossoides (Goldm.) Holttum, Kalikasan 3 (1974) 197; --Acrostichum ophioglossoides Goldm., Nov. Actorum Acad. Caes. Leop.-Carol. German. Nat. Cur. 19 Suppl. 1 (1843) 451. --Elaphoglossum merrillii Christ, PJS 3 (1908) 275. Endemic to the Philippines. LUZON, MINDORO, SAMAR. Photos

Hemigramma Christ = Tectaria

Lastreopsis Ching

1. Lastreopsis tenera (R.Br.) Tindale, Victoria Naturalist 73 (1957) 181; --Nephrodium tenerum R.Br., Prodr. (1810) 149. --Ctenitis recedens (J.Sm.) Copel., Gen. Fil. (1947) 124; --Lastrea recedens J.Sm. ex Moore, Gard. Chr. (1855) 708. Australia, Fiji, India, Lesser Sunda Isls, New Caledonia, Peninsular Malaysia, Sri Lanka, Sumatra, Taiwan, Philippines. LEYTE, LUZON.

Lomagramma J.Sm.

1. Lomagramma brooksii Copel., PJS 3 (1908) 345. Borneo, Moluccas, Palau Isls, Sulawesi, Philippines. LUZON.
2. Lomagramma copelandii Holttum, Gard. Bull. Straits Settlem. 9 (1937) 201. Moluccas, Philippines. BOHOL, CATANDUANES, LUZON, MINDANAO, NEGROS, SAMAR.
3. Lomagramma merrillii Holttum, Gard. Bull. Straits Settlem. 9 (1937) 208. Sulawesi, Philippines. MINDANAO, NEGROS. Photos
4. Lomagramma pteroides J.Sm., J. Bot. (Hooker) 3 (1841) 402. --Lomagramma subcoriacea Copel., PJS 40 (1929) 308. --Lomagramma pedicellata Copel., PJS 81 (1952) 23. --Lomagramma cordata Copel., PJS 84 (1955) 162. Moluccas, Philippines. LUZON, MINDANAO: Bukidnon, MINDORO, NEGROS, PALAWAN, PANAY. Photos

Nothoperanema (Tagawa) Ching = Dryopteris

Peranema D.Don = Dryopteris

Phanerophlebia C.Presl = Cyrtomium

Polystichum Roth

1. Polystichum biaristatum (Blume) T.Moore, Index Fil. (1858) 86; --Aspidium biaristatum Blume, Enum. Pl. Javae 2 (1828) 164. --Polystichum aquilinum Copel., PJS 81 (1952) 20. Bangladesh, Java, Lesser Sunda Isls, Myanmar, Peninsular Malaysia, Sri Lanka, Sulawesi, Taiwan, Thailand, Vietnam, Philippines.
2. Polystichum copelandii Christ, PJS 2 c (1907) 157. Endemic to the Philippines. LUZON.
3. Polystichum deltodon (Baker) Diels, Nat. Pflanzenfam. 1 (1899) 191; --Aspidium deltodon Baker, Gard. Chr. n. s. 14 (1810) 494. China, Japan, Myanmar, Ryukyu Isls, Taiwan, Vietnam, Philippines. PANAY (photos). Photos
4. Polystichum elmeri Copel., PJS 81 (1952) 21. Endemic to the Philippines. MINDANAO: Bukidnon. Photos
5. Polystichum fuscum Copel., PJS 81 (1952) 20. Endemic to the Philippines. LUZON.
6. Polystichum moluccense T.Moore, Index Fil. (1858) 84. --Polystichum horizontale C.Presl, Epim. Bot. (1849) 57. --Polystichum nudum Copel., PJS 1 (Suppl. 2) (1906) 145. Bismarck Arch, Borneo, ?Fiji, Moluccas, New Guinea, Solomon Isls, Sulawesi, Philippines. MINDANAO: Bukidnon. Conservation status: Endangered (DAO 2017-11, as Polystichum nudum Copel.). Photos
7. Polystichum nepalense (Spreng.) C.Chr., Index Filic. 10 (1905) 84; --Aspidium nepalense Spreng., Syst. 4 (1827) 97. Bhutan, China, India, Myanmar, Nepal, Taiwan, Tibet, Vietnam, Philippines.
8. Polystichum obliquum (D.Don) T.Moore, Index Fil. 87 (1858) 98; --Aspidium obliquum D.Don, Prodr. Fl. Nepal. (1825) 3. Bhutan, China, India, Myanmar, Nepal, Taiwan, Philippines, LUZON.
9. Polystichum obtusum J.Sm. ex C.Presl, Epim. Bot. (1849) 53. --Polystichum edanoi Copel., PJS 83 (1954) 97. Borneo, Philippines. LUZON.
10. Polystichum piceopaleaceum Tagawa, Acta Phytotax. Geobot. 5 (1936) 255. --Polystichum doianum Tagawa, Acta Phytotax. Geobot. 5 (1936) 256. --Polystichum pulogense Copel., 81 PJS (1952) 19. Bhutan, China, India, Japan, Myanmar, Nepal, Pakistan, Sri Lanka, Taiwan, Tibet, Philippines.

Possibly present:

1. Polystichum grandifrons C.Chr., Index Filic., Suppl. 3 (1934) 163. China, India, Japan, Taiwan, Vietnam, ?Philippines.
2. Polystichum manmeiense (Christ) Nakaike, Misc. Publ. Nat. Sci. Mus. Tokyo (1982) 141; --Aspidium manmeiense Christ, Bull. Herb. Boissier 6 (1898) 965. Bhutan, China, India, Myanmar, Nepal, Taiwan, Tibet, Tibet, ?Philippines.

Rumohra sensu Copel. (1960) non Raddi = Arachniodes

Stenolepia Alderw. = Dryopteris

Teratophyllum Mett.

1. Teratophyllum aculeatum (Blume) Mett. ex Kuhn, Ann. Mus. Bot. Lugduno-Batavi 4 (1869) 296; --Lomaria aculeata Blume, Enum. Pl. Javae (1828) 205. Photos
   • var. aculeatum. Borneo, Java, Moluccas, Myanmar, New Guinea, Singapore, Sri Lanka, Sulawesi, Sumatra, Thailand, Philippines. MINDANAO: Bukidnon.
2. Teratophyllum arthropteroides (Christ) Holttum, Gard. Bull. Straits Settlem. 5 (1932) 303; --Stenochlaena arthropteroides Christ, Bull. Herb. Boissier, sér. 2, 6 (1906) 998. Endemic to the Philippines.
3. Teratophyllum koordersii Holttum, Gard. Bull. Straits Settlem. 5 (1932) 301. Sulawesi, Philippines. LUZON. Photos
4. Teratophyllum leptocarpum (Fée) Holttum, Gard. Bull. Straits Settlem. 9 (1937) 143; --Lomariopsis leptocarpa Fée, Mém. Foug. (1845) 69. Micronesia, Sulawesi, Philippines. BASILAN, LUZON, PANAY, SAMAR, SIBUYAN. Photos
5. Teratophyllum luzonicum Holttum, Gard. Bull. Straits Settlem. 5 (1932) 297. Endemic to the Philippines. Photos

Equisetaceae

Equisetum L.

1. Equisetum ramosissimum Desf., Fl. Atlant. 2 (1799) 398.
   
   • ssp. debile (Roxb. ex Vauch.) Hauke, Amer. Fern J. 52 (1962) 33; --Equisetum debile Roxb. ex Vaucher, Mém. Soc. Phys. Genève 1 (1822) 387. Afghanistan, Bangladesh, Borneo, Cambodia, China, Fiji, India, Java, Laos, Lesser Sunda Isls, Moluccas, Myanmar, Nepal, New Caledonia, New Guinea, New Guinea, Peninsular Malaysia, Solomon Isls, Sri Lanka, Sulawesi, Sumatra, Taiwan, Thailand, Tibet, Vanuatu, Vietnam, Philippines. MINDANAO: Bukidnon. Photos.  

Gleicheniaceae

Dicranopteris Bernh.

1. Dicranopteris curranii Copel., PJS 81 (1952) 4; --Copel., FFP 1 (1958) 46. Sumatra, Peninsular Malaysia, Java, Lesser Sunda Isls, Borneo, Sulawesi, Philippines. LUZON: Laguna, NEGROS, MINDANAO. Photos  
2. Dicranopteris linearis (Burm.) Underw., Bull. Torrey Bot. Club 34 (1907) 250; --Copel., FFP 1 (1958) 45. MINDANAO: Bukidnon. Photos
   
   • var. rigida (Blume) Holttum; --Gleichenia hermannii var. rigida Blume. --Dicranopteris crassifolia (C.Presl) Ching, Sunyatsenia 5 (1940) 275; --Copel., FFP 1 (1958) 46; --Mertensia crassifolia C.Presl, Abh. Böhm. Ges. Wiss. 5 (1848) 339. Borneo, Moluccas, Sulawesi, Philippines. LUZON, MINDANAO, NEGROS.
   • var. stipulosa Christ, Bull. Herb. Boiss. 2, 6 (1906) 1010. Endemic to the Philippines.
   • var. subspeciosa Holttum. Borneo, Peninsular Malaysia, Philippines. LUZON, MINDANAO, MINDORO.
3. Dicranopteris splendida (Hand.-Mazz.) Tagawa, Acta Phytotax. Geobot. 8 (1939) 164. --Dicranopteris linearis var. latiloba Holttum. Bangladesh, Bhutan, China, India, Laos, Micronesia, Myanmar, Nepal, Thailand, Tibet, Vietnam, Philippines. LUZON, MINDANAO, NEGROS.
   
4. Dicranopteris taiwanensis Ching & Chiu in Chien & Chun, Fl. Reipubl. Popularis Sin. 2 (1959) 346. --Dicranopteris linearis var. altissima Holttum. ?Andaman Isls, Australia, Bhutan, Borneo, Borneo, Cambodia, China, India, Java, Laos, Moluccas, Myanmar, Nepal, New Guinea, Peninsular Malaysia, Solomon Isl., Sumatra, Taiwan, Thailand, Philippines. PALAWAN.
   

Diplopterygium (Diels) Nakai

Photos

1. Diplopterygium elmeri (Copel.) Nakai, Bull. Natl. Sci. Mus. 29 (1950) 49; --Gleichenia elmeri Copel. in Elmer, LPB 3 (1910) 799; --Hicriopteris elmeri (Copel.) Copel., FFP 1 (1958) 43. Endemic to the Philippines. MINDANAO. Photos
2. Diplopterygium laevissimum (Christ) Nakai, Bull. Natl. Sci. Mus. 29 (1950) 52; --Gleichenia laevissima Christ, Bull. Ac. Inst. Geogr. Bot. 3, 11 (1902) 268; --Hicriopteris laevissima (Christ) Ching, Sunyatesenia 5 (1940) 280; --Copel., FFP 1 (1958) 44. China, Japan, Peninsular Malaysia, South Korea, Taiwan, Vietnam, Philippines. LUZON.
3. Diplopterygium longissimum (Blume) Nakai, Bull. Natl. Sci. Mus. 29 (1950) 53; --Gleichenia longissima Blume, Enum. Pl. Jav. (1828) 250. --Hicriopteris glauca (Thunb. ex Houtt.) Ching, Sunyatsenia 5 (1940) 279; --Copel., FFP 1 (1958) 43; --Polypodium glaucum Thunb. ex Houtt., Nat. Hist. 2 (1783) 177. S China to Indochina through Malesia to Melanesia (Fiji, Tahiti). In Malesia not collected in the Lesser Sunda Isls E of Flores, Sulawesi, the Moluccas and New Guinea. MINDANAO: Bukidnon. Throughout the Philippines, usually at 800-1500m in exposed places. Photos
   
4. Diplopterygium sordidum (Copel.) Parris, Malayan Nat. J. 50 (1997) 252; --Gleichenia sordida Copel. in Elmer, LPB 3 (1910) 798; --Hicriopteris sordida (Copel.) Copel., FFP 1 (1958) 44. Moluccas, New Guinea, Peninsular Malaysia, Solomon Isls, Sulawesi, Sumatra, Philippines.

Gleichenia Sm.

1. Gleichenia dicarpa R.Br., Prodr. Fl. Nov. Holl. (1810) 161; --Copel., FFP 1 (1958) 38. Australia, Borneo, Chatham Isls, New Caledonia, New Guinea, New Zealand, Peninsular Malaysia, Sulawesi, Western Samoa, Philippines. MINDANAO: Agusan del Norte, Bukidnon, Davao. Photos  
2. Gleichenia microphylla R.Br., Prodromus Florae Novae Hollandiae (1810) 161. Australia, Borneo, Cambodia, Moluccas, New Zealand, Peninsular Malaysia, Sulawesi, Sumatra, Thailand, Vietnam, Philippines. Photos
3. Gleichenia paleacea (Copel.) Holttum, Reinwardtia 4 (1957) 265; --Hicriopteris paleacea Copel., PJS 81 (1952) 3; FFP 1 (1958) 44. Endemic to the Philippines. NEGROS.
4. Gleichenia peltophora Copel., PJS 40 (1929) 292, t. 1; --Copel., FFP 1 (1958) 39. Borneo, New Guinea, Sulawesi, Philippines. MINDANAO: South Cotabato (Mt Matutum), 1600m.
5. Gleichenia vulcanica Blume, Enum. Pl. Jav. (1828) 251. Borneo, Java, New Guinea, Peninsular Malaysia, Solomon Isls, Sulawesi, Sumatra, Philippines. MINDORO: Mindoro Oriental (Mt Halcon), PALAWAN (Mt Pulgar and Mt Mantalingahan, Co 1773), MINDANAO: Bukdnon. Photos

Sticherus C.Presl

Last edited by Julie F. Barcelona, 7 September 2020

Photos

1. Sticherus hirtus (Blume) Ching, Sunyatsenia 5 (1940) 203 (err. ‘hirsutus’); --Gleichenia hirta Blume, Enum. Pl. Jav. (1828) 250; --Copel., FFP 1 (1958) 40. Moluccas, Sulawesi, Thailand, Philippines. LUZON, MINDORO, MINDANAO: Bukidnon. Photos 
   
2. Sticherus hispidus (Mett. ex Kuhn) Copel., Gen. Fil. (1947) 27; FFP 1 (1958) 41; --Gleichenia hispida Mett. ex Kuhn, Verh. K.K. Zool.-Bot. Ges. Wien 25 (1875) 600. --Sticherus koordersii (Christ) Copel., FFP 1 (1958) 40; --Gleichenia koordersii Christ, Ann. Jard. Bot. Buitenzorg 15 (1897) 7. --Sticherus pinnatus Copel. Java, Lesser Sunda Isls, Moluccas, Sulawesi, Sumatra, Philippines. LUZON: Benguet, Albay (Mt Mayon), NEGROS.
3. Sticherus loheri (Christ) Copel., FFP 1 (1958) 42; --Gleichenia loheri Christ, Bull. Herb. Boiss. 2, 6 (1906) 1009. --Sticherus perpaleaceus Copel., PJS 81 (1952) 3; FFP 1 (1958) 42. Borneo, Moluccas, Sulawesi, Philippines. MINDANAO: Bukidnon. Photos
   • var. loheri. Endemic to the Philippines. LUZON: Benguet (Mt Pulog), Rizal, Quezon (Mt Banahaw).
   • var. major Holttum, Reinwardtia 4 (1959) 272. Borneo, Moluccas, Sulawesi, Philippines. NEGROS: Negros Occidental (Mt Kanlaon). 
4. Sticherus milnei (Baker) Ching, Sunyatsenia 5 (1940) 284; --Gleichenia milnei Baker, Syn. Fil. (1874) 449. Australia, Bismarck Arch, Moluccas, New Guinea, New Hebrides, Pacific, Solomon Isls, Sulawesi, Philippines. LUZON. Photos
5. Sticherus truncatus (Willd.) Nakai, Bull. Natl. Sci. Mus. 29 (1950) 20; --Gleichenia truncata (Willd.) Sprengel, Syst. Veg. ed. 16, 4 (1827) 25; --Mertensia truncata Willd., Kongl. Vetensk. Acad. Nya Handl. 25 (1804) 169. Borneo, Cambodia, China, Java, Lesser Sunda Isls, Moluccas, Singapore. Sulawesi, Sumatra, Thailand, Viernam, Philippines. PALAWAN. Photos
• var. truncatus. --Sticherus laevigatus (Willd.) C.Presl, Tent. (1836) 52; --Copel., FFP 1 (1958) 41. Throughout Malesia, apparently most common in the west. 

Hymenophyllaceae

Abrodictyum C.Presl

Callistopteris C.Presl

Cephalomanes C.Presl

1. Cephalomanes atrovirens C.Presl, Hymen. (1843) 17; --Copel., FFP 1 (1958) 77; --Trichomanes atrovirens (C.Presl) Kunze, Bot. Zeit. 6 (1847) 371; --Copel., PJS 51 (1933) 251.
   
   • ssp. atrovirens. Australia, New Guinea, Pacific, Palawan, Ryukyu Isls, Solomon Isls, Sulawesi, Philippines. MINDANAO: Bukidnon. Photos.  
2. Cephalomanes crassum (Copel.) M.G.Price, Contr. Univ. Michigan Herb. 17 (1990) 268; --Trichomanes crassum Copel., PJS 51 (1933) 256. Endemic to the Philippines. LEYTE, SAMAR.
3. Cephalomanes javanicum (Blume) Bosch, Hymenophyll. Javan. (1861) 30–31; --Trichomanes javanicum Blume, Enum. Pl. Javae 2 (1828) 224.
   
   • var. asplenioides (C.Presl) K.Iwats., J. Fac. Sci. Univ. Tokyo, Sect. 3, Bot. 13 (1985) 549; --Trichomanes asplenioides C.Presl., Abhandl. Kön. Böhm. Ges. Wiss. 5 (1843) 15. --Cephalomanes oblongifolium C.Presl, Epim. Bot. (1849) 19; --Copel., FFP 1 (1923) 78. --Trichomanes laciniatum Roxb., Calcutta J. Nat. Hist. 4 (1844) 518. Borneo, Peninsular Malaysia, Ryukyus Isls, Sulawesi, Taiwan, Philippines. MINDANAO: Bukidnon, PALAWAN. Photos

Crepidomanes (C.Presl) C.Presl

1. Crepidomanes bipunctatum (Poir.) Copel., PJS 67 (1938) 59; FFP 1 (1958) 72; --Trichomanes bipunctatum Poir. in Lam., Encycl. 8 (1808) 69; --Copel., PJS 51 (1933) 177. Australia, Bangladesh, Bismarck Arch, Borneo, Cambodia, China, Comoros, India, La Reunion, Laos, Lesser Sunda Isls, Madagascar, Mauritius, Micronesia, Myanmar, New Caledonia, New Guinea, Pacific, Palau Isls, Peninsular Malaysia, Ryukyu Isls, Solomon Isls, Sulawesi, Sumatra, Taiwan, Thailand, Thailand, Vietnam, Vietnam, Philippines. Photos
2. Crepidomanes brevipes (C.Presl) Copel., PJS 67 (1938) 60; FFP 1 (1958) 71; --Trichomanes brevipes (C.Presl) Baker, Syn. Fil. (1867) 84; --Copel., PJS 51 (1933) 182, pl. 20; --Didymoglossum brevipes C.Presl. Borneo, Micronesia, New Guinea, Southern Marianas, Sulawesi, Thailand, Philippines. Photos
3. Crepidomanes campanulatum (Roxb.) Panigrahi & Sarn. Singh, Ferns Fern-Allies Arunachal Pradesh 1 (2005) 333; --Hymenophyllum campanulatum (Roxb.) Christ, PJS 2 c (1907) Bot.155; --Copel., PJS 64 (1937) 32; --Meringium campanulatum (Roxb.) Copel., PJS 67 (1938) 40; FFP 1 (1958) 60; --Trichomanes campanulatum Roxb. Bhutan, China, India, Myanmar, Nepal, Nicobar Isls, Peninsular Malaysia, Sri Lanka, Tibet, Vietnam, Philippines. NEGROS, PANAY.
   
4. Crepidomanes christii (Copel.) Copel., PJS 67 (1938) 60; FFP 1 (1958) 73; --Trichomanes christii Copel., PJS 1 (1906) Suppl. 251; PJS 51 (1933) 185. Borneo, India, New Guinea, Peninsular Malaysia, Singapore, Sumatra, Thailand, Philippines. LUZON.
5. Crepidomanes grande (Copel.) Ebihara & K.Iwats., Blumea 51 (2006) 239; --Cephalomanes grande (Copel.) K.Iwats.; --Trichomanes grande Copel., PJS 6 c (1911) 70; PJS 51 (1933) 224; --Nesopteris grandis (Copel.) Copel., PJS 67 (1938) 66; FFP 1 (1958) 76. Borneo, Lesser Sunda Isls, Micronesia, ?Nicobar Isls, Solomon Isls, Sulawesi, Taiwan, Philippines. Photos
6. Crepidomanes humile (G.Forst.) Bosch, Hym. Jav. (1861) 16; --Trichomanes humile G.Forst., Prodr. (1786) 12; --Copel., PJS 51 (1933) 164; --Crepidopteris humilis (G.Forst.) Copel., PJS 67 (1938) 58; FFP 1 (1958) 70. --Trichomanes gracillimum Copel., PJS 51 (1933) 168; --Crepidopteris gracillima (Copel.) Copel., PJS 67 (1938) 58; FFP 1 (1958) 70. Andaman Isls, Australia, Bismarck Arch, Borneo, Java, Kermadec Isls, Lesser Sunda Isls, Micronesia, Moluccas, New Caledonia, New Guinea, Nicobar Isls, Pacific, Palau Isls, Peninsular Malaysia, Ryukyu Isls, Singapore, Solomon Isls, Sulawesi, Sumatra, Taiwan, Thailand, Philippines. LUZON, PALAWAN. Photos
7. Crepidomanes minutum (Blume) K.Iwats., J. Fac. Sci. Univ. Tokyo, Sect. 3, Bot. 13 (1985) 524; --Trichomanes minutum Blume, Enum. Pl. Jav. (1828) 223. --Gonocormus minutus (Blume) Bosch, Hymen. Javan. (1861) 7; --Copel., FFP 1 (1958) 68. --Trichomanes minutum Blume forma alagensis (Christ) Copel; --Trichomanes alagensis Christ, PJS 3 c (1908) Bot. 270; --Copel., PJS 51 (1933) 152; --Gonocormus alagensis (Christ) Copel., PJS 67 (1938) 57. Andaman Isls, Australia, Bonin Isl, Borneo, Cambodia, China, Comoros, India, Japan, Java, Korea Peninsula, Lesser Sunda Isls, Mauritius, Micronesia, Moluccas, New Caledonia, New Guinea, Norfolk Isl, Pacific, Palau Isls, Peninsular Malaysia, Russia, Ryukyu Isls, Singapore, Solomon Isls, Sri Lanka, Sulawesi, Sumatra, Sumatra, Taiwan, Thailand, Vietnam, Philippines. MINDANAO: Bukidnon, PALAWAN. Photos
8. Crepidomanes rupicola (Racib.) Copel., PJS 67 (1938) 59; FFP 1 (1958) 73; --Trichomanes rupicolum Racib., Pterid. Buitenz. (1898) 24. Java, Lesser Sunda Isls, Solomon Isls, Taiwan, Philippines.
9. Crepidomanes thysanostomum (Makino) Ebihara & K.Iwats., Blumea 51 (2006) 240; --Cephalomanes thysanostomum (Makino) K.Iwats.; --Trichomanes thysanostomum Makino, Bot. mag. Tokyo 12 (1898) 193; --Nesopteris thysanostoma (Makino) Copel., PJS 67 (1938) 66. Japan, ?Nicobar Isls, Ryukyu Isls, Sulawesi, Taiwan, Philippines.

Possibly present:

1. Crepidomanes kurzii (Bedd.) Tagawa & K.Iwats., Acta Phytotax. Geobot. 26 (1975) 169; --Trichomanes kurzii Bedd., Handb. Ferns Brit. India (1868) t.286. Andaman Isls, Australia, Borneo, Cambodia, Hong Kong, India, Laos, Myanmar, New Guinea, Pacific, Peninsular Malaysia, Ryukyu Isls, Solomon Isls, Sri Lanka, Sumatra, Taiwan, Thailand, Vietnam, ?Philippines.
2. Crepidomanes schmidtianum (Zenker ex Taschner) K.Iwats., J. Fac. Sci. Univ. Tokyo, Sect. 3, Bot. 13(5) (1985) 526; --Trichomanes schmidtianum Zenker ex Taschner, Dissertation (1843) 34. Bhutan, China, India, Japan, Myanmar, Nepal, Sulawesi, Sumatra, Tibet, Philippines.
   
   • var. latifrons (Bosch) K.Iwats., J. Fac. Sci. Univ. Tokyo, Sect. 3, Bot. 13 (1985) 527; --Trichomanes latifrons Bosch, Ned. Kruidk. Arch. 5 (1863) 209; --Vandenboschia latifrons (Bosch) Copel., PJS 67 (1938) 104. China, Himalaya, Taiwan, ?Philippines.

Crepidopteris Copel. = Crepidomanes

Didymoglossum Desv.

1. Didymoglossum beccarianum (Ces.) Senterre & Rouhan, Phytotaxa 292 (2017) 210; --Trichomanes beccarianum Ces., Atti Accad. Napoli, Part 8 7 (1876) 8; --Copel., PJS 51 (1933) 200; --Microgonium beccarianum (Cesati) Copel., PJS 67 (1938) 63; FFP 1 (1958) 74. --Didymoglossum motleyi auct. non (Bosch) Ebihara & K.Iwats.; --Trichomanes motleyi auct. non (Bosch) Bosch; --Copel., PJS 51 (1933) 201; --Microgonium motleyi auct. non Bosch; --Copel., FFP 1 (1958) 75. Andaman Isls, Australia, Bismarck Arch, Borneo, Java, Micronesia, Moluccas, Myanmar, New Guinea, Palau Isls, Peninsular Malaysia, Ryukyu Isls, Singapore, Solomon Isls, Sri Lanka, Sulawesi, Taiwan, Thailand, Vietnam, Philippines. PALAWAN. Photos
   
2. Didymoglossum mindorense (Christ) K.Iwats., Blumea 51 (2006) 236; -- Trichomanes mindorense Christ, PJS 3 c (1908) Bot. 270; --Copel., PJS 51 (1933) 209; --Microgonium mindorense (Christ) Copel., PJS 67 (1938) 61; PJS 51 (1933) 164. --Trichomanes craspedoneurum Copel., PJS 7 c (1912) Bot. 58; PJS 51 (1933) 208; --Microgonium craspedoneurum (Copel.) Copel., PJS 67 (1938) 61; FFP 1 (1958) 74. Australia, Bismarck Arch, India, Micronesia, New Guinea, Peninsular Malaysia, Singapore, Solomon Isls, Philippines. LUZON.

Gonocormus Bosch = Crepidomanes

Hymenophyllum Sm.

1. Hymenophyllum acanthoides (Bosch) Rosenst., Bull. Jard. Bot. Buitenz. 2, 2 (1911) 25; --Copel., PJS 64 (1937) 45, pl. 17; --Meringium acanthoides (Bosch) Copel., PJS 67 (1938) 42; FFP 1 (1958) 63; --Didymoglossum acanthoides Bosch, Pl. Jungh. 1 (1856) 560. Borneo, India, Java, New Guinea, Peninsular Malaysia, Singapore, Sulawesi, Sumatra, Thailand, Philippines. MINDANAO: Bukidnon. Photos
2. Hymenophyllum acutum (C.Presl) Ebihara & K.Iwats., Blumea 51 (2006) 232; --Crepidomanes acutum (C.Presl) K.Iwats.; --Trichomanes acutum C.Presl, Epim. Bot. (1849) 258. Micronesia, New Guinea, Solomon Isls, Taiwan, Philippines.
3. Hymenophyllum angulosum Christ, PJS 3 c (1908) Bot. 269; --Copel., PJS 64 (1937) 33; --Mecodium angulosum (Christ) Copel., PJS 64 (1937) 109, pl. 50; FFP 1 (1958) 49. New Guinea, Solomon Isls, Vanuatu, Philippines. Photos
4. Hymenophyllum badium Hook. & Grev., Icon. Fil. (1828) pl. 76; --Copel., PJS 64 (1937) 144; --Mecodium badium (Hook. & Grev.) Copel., PJS 67 (1938) 23; FFP 1 (1958) 53. --Hymenophyllum crispatum Wall. ex Hook. & Grev., Icon. Fil. (1828) pl. 72; --Copel., PJS 64 (1937) 148; --Mecodium crispatum (Wall.) Copel., PJS 67 (1938) 23. Bhutan, Borneo, Cambodia, China, India, Japan, Laos, Myanmar, Nepal, Peninsular Malaysia, Ryukyu Isls, Solomon Isls, Sulawesi, Sumatra, Taiwan, Thailand, Tibet, Vietnam, Philippines. Photos
5. Hymenophyllum bartlettii (Copel.) C.V.Morton, Contr. U.S. Natl. Herb. 38 (1968) 167; --Meringium bartlettii Copel., PJS 73 (1940) 464; FFP 1 (1958) 62. Endemic to the Philippines. MINDANAO.
6. Hymenophyllum bicolanum Copel., PJS 64 (1937) 31; --Meringium bicolanum (Copel.) Copel., PJS 67 (1938) 40; FFP 1 (1958) 60. Endemic to the Philippines. LUZON.
7. Hymenophyllum blandum Racib., Pterid. Buitenz. (1898) 20; --Copel., PJS 64 (1937) 50; --Meringium blandum (Racib.) Copel., PJS 67 (1938) 43; FFP 1 (1958) 63.Borneo, Java, Lesser Sunda Isls, New Guinea, Peninsular Malaysia, Sulawesi, Sumatra, Taiwan, Thailand, Philippines.
8. Hymenophyllum bontocense Copel., PJS 64 (1937) 33; --Meringium bontocense (Copel.) Copel., PJS 67 (1938) 41; FFP 1 (1958) 60. Thailand, Philippines.
9. Hymenophyllum denticulatum Sw., Schrad. J. (1801) 100; --Copel., PJS 64 (1937) 41; --Meringium denticulatum (Sw.) Copel., PJS 67 (1938) 42; FFP 1 (1958) 62. Bangladesh, Borneo, China, Fiji, India, Java, Lesser Sunda Isls, Myanmar, Peninsular Malaysia, Ryukyu Isls, Singapore, Solomon Isls, Sri Lanka, Sulawesi, Sumatra, Taiwan, Thailand, Vietnam, Philippines. MINDANAO: Bukidnon, PALAWAN. Photos
10. Hymenophyllum digitatum (Sw.) Fosberg, Smithsonian Contr. Bot. 45 (1980) 1; --Crepidomanes digitatum (Sw.) K.Iwats.; --Trichomanes digitatum Sw., Syn. Fil. (1806) 370; --Copel., PJS 51 (1933) 159; --Microtrichomanes digitatum (Sw.) Copel., PJS 67 (1938) 36; FFP 1 (1958) 55. Africa, Australia, Bismarck Arch, Borneo, Java, La Reunion, Lesser Sunda Isls, Madagascar, Mauritius, Micronesia, Pacific, Peninsular Malaysia, Philippines, Solomon Isls, Sulawesi, Taiwan, Tanzania, Thailand, Vietnam, Philippines.
11. Hymenophyllum edanoi (Copel.) C.V.Morton, Contr. U.S. Natl. Herb. 38 (1968) 173; --Mecodium edanoi Copel., PJS 81 (1952) 2; FFP 1 (1958) 50. Endemic to the Philippines. LUZON, MINDORO.
12. Hymenophyllum edentulum (Bosch) C.Chr., Index Fil. (1905) 360; --Meringium edentulum (Bosch) Copel., PJS 67 (1938) 41; FFP 1 (1958) 58. --Hymenophyllum bakeri Copel., Sarawak Mus. J. 2 (1917) 309; --Copel., PJS 64 (1937) 29; --Meringium bakeri (Copel.) Copel., PJS 67 (1938) 40; FFP 1 (1958) 59. Micronesia, Philippines. PALAWAN.
    
13. Hymenophyllum emarginatum Sw., Schrad. J. 2 of 1800 (1801) 101; --Copel., PJS 64 (1937)  118, pl. 58; --Mecodium emarginatum (Sw.) Copel., PJS 67 (1938) 21; FFP 1 (1958) 52. --Hymenophyllum demissum auct. non Sw.; --Hook., Sp. Fil. 1 (1846) 109; --Baker, Syn. Fil. ed. 2 (1874) 61. Borneo, Lesser Sunda Isls, Peninsular Malaysia, Society Isls, Sulawesi, Sumatra, Philippines. MINDANAO: Bukidnon. Photos  
14. Hymenophyllum fimbriatum J.Sm., J. Bot. 3 (1841) 418, nomen; --Hook., Sp. Fil. (1844) 102; --Copel., PJS 64 (1937) 122; --Mecodium fimbriatum (J.Sm.) Copel., PJS 67 (1938) 21; FFP 1 (1958) 52. Borneo, Moluccas, New Guinea, Taiwan, Vietnam, Philippines. Photos
15. Hymenophyllum holochilum (Bosch) C.Chr., Index Fil. (1905) 226, 362; --Copel., PJS 64  (1937) 34; --Meringium holochilum (Bosch) Copel., PJS 67 (1938) 41; FFP 1 (1958) 62; --Didymoglossum holochilum Bosch, Pl. Jungh. 5 (1857) 561. --Hymenophyllum merrillii Christ, PJS 2 c (1907) Bot. 154; --Copel., PJS 64 (1937) 33; --Meringium merrillii (Christ) Copel., PJS 67 (1938) 41; FFP 1 (1958) 61. Australia, Bismarck Arch, Borneo, Java, Lesser Sunda Isls, Micronesia, New Caledonia, Pacific, Palawan, Peninsular Malaysia, Singapore, Solomon Isls, Sulawesi, Sumatra, Taiwan, Thailand, Philippines. Photos
16. Hymenophyllum imbricatum Blume, Enum. Pl. Jav (1828) 220; --Copel., PJS 64 (1937) 137; --Mecodium imbricatum (Blume) Copel., PJS 67 (1938) 22; FFP 1 (1958) 53. Bismarck Arch, Java, New Caledonia, New Guinea, Pacific, Peninsular Malaysia, Solomon Isls, Sulawesi, Sumatra, Philippines. Photos
17. Hymenophyllum inaequale (Poir.) Desv., Mem. Soc. Linn. Paris 6, 2 (1827) 335; --Trichomanes inaequale Poir., Encycl. 8 (1808) 74. --Hymenophyllum polyanthos (Sw.) Sw., Schrad. J. ii of 1800 (1801) 102; --Copel., PJS 64 (1937) 97; --Mecodium polyanthos (Sw.) Copel., PJS 67 (1938) 19; FFP 1 (1958) 49; --Trichomanes polyanthos Sw., Prodr. (1788) 137. Africa, Australia, Bhutan, Bismarck Arch, Borneo, Cambodia, China, India, Indian Ocean, Japan, Java, Laos, Lesser Sunda Isls, Myanmar, Nepal, New Guinea, Pacific, Palau Isls, Peninsular Malaysia, Ryukyu Isls, Singapore, Solomon Isls, South Korea, Sri Lanka, Sulawesi, Sumatra, Taiwan, Thailand, Vietnam, Philippines. MINDANAO: Bukidnon, PALAWAN.
    
18. Hymenophyllum javanicum Spreng., Syst. Veg. (ed. 16) 4 (1827) 132. ?Andaman Isls, Australia, Borneo, Cambodia, India, Laos, Lesser Sunda Isls, Moluccas, Myanmar, New Caledonia, New Guinea, Pacific, Peninsular Malaysia, Solomon Isls, Sri Lanka, Sulawesi, Sumatra, Taiwan, Thailand, Vietnam, Philippines.
19. Hymenophyllum johorense Holttum, Gard. Bull. Sing. 4 (1929) 408; --Copel., PJS 64  (1937) 51; --Meringium johorense (Holttum) Copel., FFP 1 (1958) 64. --Hymenophyllum reductum Copel., PJS 64 (1937) 53, pl. 20; --Meringium reductum (Copel.) Copel., PJS 67 (1938) 43; FFP 1 (1958) 64. Borneo, Peninsular Malaysia, Philippines. Photos
20. Hymenophyllum klabatense Christ, Verh. Nat. Ges. Basel 11 (1894) 4; --Copel., PJS 64 (1937) 30; --Meringium klabatense (Christ) Copel., PJS 67 (1938) 40; FFP 1 (1958) 59. Sulawesi, Philippines.
21. Hymenophyllum nitiduloides Copel., PJS 64 (1937) 112; --Mecodium nitiduloides (Copel.) Copel., PJS 64 (1937) 112, pl. 52; FFP 1 (1958) 50. Endemic to the Philippines. LUZON.
22. Hymenophyllum pachydermicum Cesati, Atti Accad. Napoli 8 (1876) 8; --Meringium pachydermicum (Cesati) Copel., PJS 67 (1938) 41; FFP 1 (1958) 57. Borneo, Sulawesi, Philippines. PALAWAN. Photos
23. Hymenophyllum pallidum (Blume) Ebihara & K.Iwats., Blumea 51 (2006) 232; --Crepidomanes pallidum (Blume) K.Iwats.; --Pleuromanes pallidum (Blume) C.Presl, Epim. Bot. (1849) 258; --Copel., FFP 1 (1958) 67; --Trichomanes pallidum Blume, Enum. Pl. Jav. (1828) 226; --Copel., PJS 51 (1933) 141. Australia, Bismarck Arch, Borneo, China, Laos, Lesser Sunda Isls, Micronesia, New Caledonia, New Guinea, Pacific, Peninsular Malaysia, Singapore, Society Isls, Solomon Isls, Sri Lanka, Sulawesi, Sumatra, Sumatra, Taiwan, Thailand, Vietnam, Philippines. MINDANAO: Bukidnon, PALAWAN. Photos
24. Hymenophyllum paniculiflorum C.Presl, Hymen. (1843) 147; --Copel., PJS 64 (1937) 110; --Chen et al., Phytotaxa 443 (2020) 132; --Mecodium paniculiflorum (C.Presl) Copel., PJS 67 (1938) 19; FFP 1 (1958) 50; --Type: Cuming 214 (B, BM, E, GH, K, MW, P, PNH, PRC), Luzon: 1841. --Hymenophyllum discosum Christ, Bull. Herb. Boissier 6 (1898) 140; --Hsu et al., Syst. Bot. 44 (2019) 762; --Type: Loher 925 (K, holo), Luzon: Benguet Prov.: Mt. Santo Tomas ('Monte Tonglon'), 2250m, Feb-1894. ?Borneo, ?Japan, ?Lesser Sunda Isls, ?Sulawesi, Taiwan, Vietnam, Philippines. LUZON: Mountain Province, MINDANAO: Bukidnon, NEGROS, PALAWAN. Conservation status: Least Concern (Hsu et al., 2019). Photos
25. Hymenophyllum pilosissimum C.Chr., Gard. Bull. 7 (1934) 213; --Sphaerocionium pilosissimum (C.Chr.) Copel., PJS 67 (1938) 33; FFP 1 (1958) 54. Borneo, New Guinea, Taiwan, Philippines. Photos
26. Hymenophyllum productum Kunze, Bot. Zeit. 6 (1848) 305; --Copel., PJS 64 (1937) 118, pl.  54; --Mecodium productum (Kunze) Copel., PJS 67 (1938) 20; FFP 1 (1958) 51. Borneo, ?Cambodia, China, Java, Lesser Sunda Isls, Peninsular Malaysia, Sulawesi, Sumatra, Taiwan, Thailand, Vietnam, Philippines.
27. Hymenophyllum ramosii Copel., PJS 64 (1937) 34, pl. 9, Fig. 4-6; --Meringium ramosii (Copel.) Copel., PJS 67 (1938) 31; FFP 1 (1958) 61. Endemic to the Philippines. MINDANAO. Photos
28. Hymenophyllum reinwardtii Bosch., Pl. Jungh. 1 (1856) 567; --Copel., PJS 64 (1937) 115; --Mecodium reinwardtii (Bosch.) Copel., PJS 67 (1938) 20; FFP 1 (1958) 51. Bismarck Arch, Borneo, Java, Micronesia, Moluccas, New Guinea, Palau Isls, Sulawesi, Sumatra, Philippines. MINDANAO: Bukidnon. Photos
29. Hymenophyllum serrulatum (C.Presl) C.Chr., Index Filic. (1905) 367. --Meringium meyenianum C.Presl, Hymen. (1843) 116, pl. 8 B; --Copel., FFP 1 (1958) 58; --Hymenophyllum meyenianum (C.Presl) Copel., PJS 64 (1937) 25; FFP 1 (1958) 58. --Hymenophyllum pulchrum Copel., PJS 64 (1937) 22; --Meringium pulchrum (Copel.) Copel., PJS 67 (1938) 42; FFP 1 (1958) 57. Bismarck Arch, Borneo, Java, ?Lesser Sunda Isls, Micronesia, ?Moluccas, New Guinea, Pacific, Palau Isls, Peninsular Malaysia, Singapore, Solomon Isls, Sulawesi, Sumatra, Thailand, Philippines. MINDANAO: Bukidnon. Photos
30. Hymenophyllum thuidium Harrington, J. Linn. Soc. Bot. 16 (1877) 25; --Copel., PJS 64 (1937) 116; --Mecodium thuidium (Harrington) Copel., PJS 67 (1938) 20; FFP 1 (1958) 52. Borneo, New Guinea, Philippines. MINDANAO: Bukidnon.
31. Hymenophyllum vittatum Copel., PJS 64 (1937) 31; FFP 1 (1958) 59; --Meringium vittatum (Copel.) Copel., PJS 67 (1938) 40; FFP 1 (1958) 59. Endemic to the Philippines. LUZON.
32. Hymenophyllum zamboanganum (Copel.) ined.; --Microtrichomanes zamboanganum Copel., PJS 81 (1952) 2; FFP 1 (1958) 56; --Trichomanes zamboanganum (Copel.) C.V.Morton. Endemic to the Philippines. MINDANAO: Zamboanga.

Possibly present:

1. Hymenophyllum nitidulum (Bosch) Ebihara & K.Iwats., Taxon 53(4) (2004) 941; --Trichomanes nitidulum Bosch, Pl. Jungh. 1 (1856) 547; --Gonocormus nitidulus (Bosch) Prantl, Unters. Morph. Gefasskrypt. (1875) 51; --Microtrichomanes nitidulum (Bosch) Copel., PJS 67 (1938) 37; --Sphaerocionium nitidulum (Bosch) K.Iwats., J. Fac. Sci. Univ. Tokyo, Bot. 13 (1982) 211. Borneo, India, Java, Lesser Sunda Isls, Peninsular Malaysia, Sri Lanka, Sumatra, Taiwan, Vietnam, ?Philippines. Photos

Sphaerocionium C.Presl = Hymenophyllum

Vandenboschia Copel.

Photos

1. Vandenboschia auriculata (Blume) Copel., PJS 67 (1938) 55; --Crepidomanes auriculatum (Blume) K.Iwats., J. Fac. Sci. Univ. Tokyo, Bot. 13 (1985) 528; --Trichomanes auriculatum Blume, Enum. Pl. Javae 2 (1828) 225. Bhutan, Borneo, Cambodia, China, India, Japan, Java, Laos, Lesser Sunda Isls, Moluccas, Myanmar, Nepal, New Guinea , Peninsular Malaysia, Ryukyu Isls, Sulawesi, Sumatra, Taiwan, Thailand, Tibet, Vietnam, Philippines. PALAWAN. Photos
2. Vandenboschia johnstonensis (Bailey) Copel., PJS 67 (1938) 54; --Crepidomanes johnstonensis (Bailey) K.Iwats., J. Fac. Sci. Univ. Tokyo, Sect. 3, Bot. 13 ((1985)) 532; --Trichomanes johnstonensis Bailey, Proc. Roy. Soc. Queensl. 1 (1884) 14. Australia, Philippines.
3. Vandenboschia maxima (Blume) Copel., 67 (1938) 54; --Crepidomanes maximum (Blume) K.Iwats., J. Fac. Sci. Univ. Tokyo, Bot. 13 (1985) 531; --Trichomanes maximum Blume, Enum. Pl. Javae 2 (1828) 228. Bismarck Arch, Borneo, Japan, Java, Lesser Sunda Isls, New Caledonia, Nicobar Isls, Pacific, Peninsular Malaysia, Ryukyu Isls, Solomon Isls, Sulawesi, Sumatra, Taiwan, Thailand, Vietnam, Philippines. Photos

Possibly present:

1. Vandenboschia kalamocarpa (Hayata) Ebihara, Acta Phytotax. Geobot. 60 (2009) 32; --Trichomanes kalamocarpum Hayata, Ic. Pl. Formos. 6  (1916) 260. China, Japan, Taiwan, ?Philippines.

Hypodematiaceae 

Didymochlaena Desv.

1. Didymochlaena philippensis Li Bing Zhang & H.Shang, Phytotaxa 479 (2021) 77. --Didymochlaena truncatula auct. non (Sw.) J.Sm. Endemic to the Philippines. LUZON: Isabela (photos), Sorsogon, MINDANAO: Agusan, Davao, Sarangani, NEGROS, PALAWAN, PANAY. Photos

Hypodematium Kunze

1. Hypodematium crenatum (Forssk.) Kuhn, Reisen Ost Afr. [Decken] 3(3) Bot. (1879) 37; --Polypodium crenatum Forssk., Fl. Aegypt.-Arab. (1775) 185. Afghanistan, Africa, Bhutan, China, India, Japan, La Reunion, Madagascar, Mauritius, Middle East, Myanmar, Nepal, New Guinea, Pakistan, Socotra, Sri Lanka, Sulawesi, Sumatra, Taiwan, Thailand, Vietnam, Philippines. Photos

Leucostegia C.Presl

1. Leucostegia pallida (Mett. ex Kuhn) Copel., PJS 34 (1927) 252; --Davallia pallida Mett. ex Kuhn, Index. 1869. Bismarck Arch, Borneo, Micronesia, Moluccas, Myanmar, New Guinea, Pacific, Peninsular Malaysia, Solomon Isls, Sulawesi, Sumatra, Philippines. Photos
2. Leucostegia truncata (D.Don) Fraser-Jenk., Taxon. Revis. Indian Subcontinental Pteridophytes (2008) 348; --Davallia truncata D.Don, Prodr. Fl. Nepal. (1825) 10. --Leucostegia immersa (Wall. ex Hook.) C.Presl, Tent. (1836) 95; --Davallia immersa Wall. ex Hook., Sp. Fil. 1 (1845) 156. Bhutan, Borneo, Cambodia, China, India, Java, Lesser Sunda Isls, Moluccas, Myanmar, Nepal, New Guinea, Peninsular Malaysia, Sri Lanka, Sulawesi, Sumatra, Taiwan, Thailand, Tibet, Vietnam, Philippines. MINDANAO: Bukidnon. Photos

Lindsaeaceae

Isoloma J.Sm. = Lindsaea

Lindsaea Dryand. ex Sm.

1. Lindsaea adiantoides J.Sm., J. Bot. 3 (1841) 415. --Lindsaea tropidorachis Alderw., Bull. Jard. Bot. Buitenzorg, sér. 3, 5 (1922) 211; --Type: Cuming 176 (K, holo; MO, P, iso), Luzon. Endemic to the Philippines. LUZON. Photos
2. Lindsaea apoensis Copel., FFP (1904) 182. --Lindsaea havicei Copel., PJS 1 (Suppl. 2) (1906) 149. Endemic to the Philippines. LEYTE, LUZON, MINDANAO: Bukidnon, NEGROS. Photos
3. Lindsaea bonii Christ, Notul. Syst. (Paris) 1 (1910) 187. --Lindsaea orbiculata var. commixta (Tagawa) K.U.Kramer, Fl. Males. Ser. 2, 1 (1971) 207; --Lindsaea commixta Tagawa, Acta phytotax. Geobot. 6 (1937) 37. Borneo, Cambodia, China, Hongkong, India, Japan, ?Laos, Lesser Sunda Isls, Luzon, Myanmar, Nepal, Peninsular Malaysia, Ryukyu Isls, Sri Lanka, Sulawesi, Sumatra, Taiwan, Thailand, Vietnam, Philippines.
   
4. Lindsaea bouillodii Christ, Not. Syst. 1 (1909) 59. Borneo, Cambodia, Java, Peninsular Malaysia, Sulawesi, Sumatra, Thailand, Vietnam, Philippines. PALAWAN: Co 1793 (A, PNH, PUH), Mt Mantalingahan, 1200m.
5. Lindsaea capillacea Christ, Bull. Herb. Boissier 6 (1898) 144. Endemic to the Philippines. LUZON.
6. Lindsaea cultrata (Willd.) Sw., Syn. Fil. (1806) 119; --Adiantum cultratum Willd., Phytographia 14 (1794) t.10. --Lindsaea decomposita sensu Copel. (1960), auct. non Willd., pro parte. Borneo, ?India, Japan, Java, Lesser Sunda Isls, Moluccas, Nicobar Isls, Pacific, Palau Isls, Peninsular Malaysia, Singapore, Solomon Isls, Sri Lanka, Sulawesi, Sumatra, Taiwan, Thailand, Philippines. PALAWAN, SIBUYAN.
7. Lindsaea divergens Wall. ex Hook. & Grev., Icon. Filic. (1831) 226; --Isoloma divergens (Wall. ex Hook. & Grev.) J.Sm., J. Bot. 3 (1841) 414. Borneo, Peninsular Malaysia, Singapore, Sumatra, Thailand, Philippines. PALAWAN.
8. Lindsaea doryphora K.U.Kramer, Blumea 15 (1968) 566. --Lindsaea parasitica sensu Copel. (1960), auct. non Wall. Bangladesh, Borneo, Java, Myanmar, Peninsular Malaysia, Singapore, Sumatra, Thailand, Philippines. LEYTE, MINDANAO, NEGROS. Photos  
9. Lindsaea ensifolia Sw., J. Bot. 1800 (1801) 77. Africa, Australia, Bangladesh, Bismarck Arch, Cambodia, China, India, Indian Ocean, Japan, Laos, Lesser Sunda Isls, Moluccas, Myanmar, Nepal, New Guinea, Nicobar Isls, Peninsular Malaysia, Ryukyu Isls, Solomon Isls, Sri Lanka, Sulawesi, Sumatra, Taiwan, Thailand, Vietnam, Philippines. PALAWAN. Photos
10. Lindsaea fissa Copel., PJS 38 (1929) 143. --Lindsaea hymenophylloides sensu Copel. (1960), auct. non Blume. Endemic to the Philippines. BILIRAN, LUZON, MINDANAO: Bukidnon, MINDORO, NEGROS, PALAWAN, PANAY. Photos  
11. Lindsaea gueriniana (Gaudich.) Desv., Mem. Soc. Linn. Paris 6 (1827) 312; --Isoloma guerinianum (Gaudich.) Fée, Mem. Foug., Gen. Filic. (1852) 108; --Schizoloma guerinianum Gaudich., Ann. Sci. Nat. (Paris) 3 (1824) 508. --Isoloma fuligineum (Copel.) Copel., PJS 78 (1950) 24; --Schizoloma fuligineum Copel., PJS 1 (Suppl. 4) (1906) 254. Andaman Isls, Borneo, Fiji, Moluccas, New Guinea, Nicobar Isls, Solomon Isls, Sulawesi, Philippines. MINDANAO, SIBUYAN. Photos
12. Lindsaea hamiguitanensis D.N.Karger & V.B.Amoroso, Phytotaxa 56 (2012) 18; --Type: Karger 444 (holo PNH, UC, Z, iso). Endemic to the Philippines. MINDANAO: Bukidnon, Davao Oriental. Conservation status: Vulnerable (DAO 2017-11). Photos
13. Lindsaea heterophylla Dryand., Trans. Linn. Soc. London 3 (1797) 41. Borneo, Cambodia, China, Hongkong, India, Indian Ocean, Java, Laos, Moluccas, Nicobar Isls, Peninsular Malaysia, Ryukyu Isls, Singapore, Sri Lanka, Sumatra, Taiwan, Thailand, Vietnam, Philippines. LUZON, MINDANAO.
14. Lindsaea javanensis Blume, Enum. Pl. Javae 2 (1828) 219. Borneo, Cambodia, China, India, Japan, Java, Laos, Myanmar, Peninsular Malaysia, Ryukyu Isls, Sulawesi, Sumatra, Taiwan, Thailand, Vietnam, Philippines. MINDORO, SIBUYAN.
15. Lindsaea lobata Poir. in Lam., Encycl. Suppl. 3 (1813) 448. Bismarck Arch, Borneo, Carolina Isl, Java, Lesser Sunda Isls, Micronesia, Moluccas, New Guinea, Palau Isls, ?Peninsular Malaysia, Sulawesi, Sumatra, Vietnam, Philippines. PALAWAN.
16. Lindsaea longifolia Copel., PJS 38 (1929) 145. Endemic to the Philippines. BASILAN, BILIRAN, CATANDUANES, LEYTE, LUZON, MINDANAO, PANAY, POLILLO, SAMAR.
17. Lindsaea lucida Blume, Enum. Pl. Jav. (1828) 216. --Lindsaea securifolia C.Presl ex Goldm., Nova Acta Suppl. 1 ad 16 (1843) 464. --Lindsaea gracilis Blume, Enum. Fil. (1828) 217. Andaman Isls, Bangladesh, Borneo, ?Cambodia, China, India, Japan, Java, ?Laos, Moluccas, Myanmar, New Guinea, Peninsular Malaysia, Ryukyu Isls, Singapore, Sulawesi, Sumatra, Taiwan, Thailand, Vietnam, Philippines. MINDANAO: Bukidnon, PALAWAN. Photos  
18. Lindsaea merrillii Copel., Perkin's Fragm. (1905) 181. Endemic to the Philippines. BATANES, CAMIGUIN, LEYTE, LUZON, MINDANAO, MINDORO, PALAWAN, POLILLO, SAMAR.
19. Lindsaea oblanceolata Alderw., Bull. Jard. Bot. Buitenz. 2, 23 (1916) 15. Borneo, Borneo, Java, Moluccas, Nicobar Isls, Peninsular Malaysia, Sumatra, Thailand, Vietnam, Philippines. LEYTE, LUZON, MINDANAO. Photos
20. Lindsaea obtusa J.Sm. ex Hook., Sp. Fil. 1 (1846) 224. --Lindsaea decomposita sensu Copel. (1960), auct. non Willd., excl. syn. --Lindsaea davallioides sensu Copel. (1960), auct. non Blume. Andaman Isls, Australia, Bismarck Arch, Borneo, Moluccas, New Guinea, Pacific, Palau Isls, Solomon Isls, Sulawesi, Taiwan, Philippines. MINDANAO: Bukidnon, PALAWAN. Photos  
21. Lindsaea orbiculata (Lam.) Mett. ex Kuhn, Ann. Mus. Bot. Lugduno-Batavi 4 (1869) 279; --Adiantum orbiculatum Lam., Encycl. 1 (1783) 41. Australia, Borneo, Cambodia, China, Japan, Java, Laos, Lesser Sunda Isls, Moluccas, Moluccas, Myanmar, Peninsular Malaysia, Ryukyu Isls, Singapore, Sulawesi, Sumatra, Taiwan, Thailand, Vietnam, Philippines. LUZON. Photos
22. Lindsaea ovata J.Sm., Sp. Fil. (1846) 204; --Isoloma ovatum (J.Sm.) C.Presl, Abh. Königl. Böhm. Ges. Wiss., ser. 5 6 (1851) 461. Borneo, Peninsular Malaysia, Singapore, Sumatra, Philippines. LUZON, MINDANAO.
23. Lindsaea pectinata Blume, Enum. Pl. Javae (1828) 217; --Lindsaea repens var. pectinata (Blume) Mett. ex Kuhn in Miq., Ann. Mus. Bot. Lugd.-Bat. 4 (1869) 277.  --Lindsaea roxasii Copel., PJS 81 (1952) 6. --Lindsaea angusta Copel., PJS 81 (1952) 5. Borneo, Cambodia, India, Java, Lesser Sunda Isls, Moluccas, ?Palau Isls, Peninsular Malaysia, Singapore, Sri Lanka, Sulawesi, Sumatra, Thailand, Vietnam, Philippines. PALAWAN.
    
24. Lindsaea pellaeiformis Christ, Verh. Naturf. Ges. Basel 11 (1897) 430. Sulawesi, Philippines. LUZON: Isabela.
25. Lindsaea philippinensis K.U.Kramer, Fl. Malesiana, Ser. 2, Pterid. 1 (1971) 232. --Isoloma angustum (Copel.) Tard., Notul. Syst. (Paris) 14 (1952) 332; --Schizoloma angustum Copel., PJS 1 (Suppl. 4) (1906) 252. Endemic to the Philippines. LUZON, PALAWAN, SIBUYAN. Photos  
26. Lindsaea pulchella (J.Sm.) Mett. ex Kuhn, Linnaea 36 (1869) 81; --Odontoloma pulchellum J.Sm., Sert. Pl. (1844) pl.51. MINDANAO: Bukidnon. Photos
    
    • var. pulchella. --Lindsaea adiantifolia (Hook.) Copel., FFP 1 (1958) 108. --Lindsaea adiantoides sensu Copel. (1960), pro parte. Lesser Sunda Isls, Moluccas, New Guinea, Sulawesi, Philippines. DINAGAT, LEYTE, LUZON, MINDANAO, MINDORO, NEGROS, PANAY, SAMAR.
    • var. blanda (Mett. ex Kuhn) K.U.Kramer, Blumea 15 (1968) 571; --Lindsaea blanda Mett. ex Kuhn, Linn. 36 (1869) 80. --Lindsaea cyathicola Copel., PJS 1 (Suppl. 2) (1906) 149. Australia, Java, Moluccas, New Guinea, Solomon Isls, Sulawesi, Sumatra, Philippines. LUZON, MINDANAO.
27. Lindsaea ramosii Copel., PJS 38 (1929) 144. Very doubtfully distinct from Lindsaea longifolia Copel. Endemic to the Philippines. LUZON, MINDANAO: Bukidnon.
28. Lindsaea repens (Bory) Bedd., Enum. Pl. Zeyl. (1864) 388. Photos
    • var. sessilis (Copel.) K.U.Kramer, Blumea 15 (1968) 568. --Lindsaea longa Copel., PJS 46 (1931) 216. --Lindsaea cultripinna Copel., PJS 81 (1952) 6. Australia, Bismarck Arch, Borneo, Moluccas, New Guinea, Pacific, Solomon Isls, Sulawesi, Philippines. MINDANAO: Bukidnon, PALAWAN.
29. Lindsaea rigida J.Sm. ex Hook., Sp. Fil. 1 (1846) 217. --Lindsaea monosora Copel., LPB 2 (1908) 398. Bismarck Arch, Borneo, Micronesia, Moluccas, New Guinea, Pacific, Peninsular Malaysia, Solomon Isls, Sulawesi, Sumatra, Philippines. LUZON, MINDANAO: Bukidnon, NEGROS, PALAWAN.
30. Lindsaea tenuifolia Blume, Enum. Pl. Javae (1828) 219. --Lindsaea polyctena K.U.Kramer, Blumea 15 (1968) 565. Java, Moluccas, New Guinea, Sulawesi, Sumatra, Western Samoa, Philippines. LEYTE, MINDANAO, SAMAR. Photos
    
31. Lindsaea tetragona K.U.Kramer, Blumea 15 (1967) 564.
    
    • var. tetragona. --Lindsaea tenuifolia sensu Copel. (1960), auct. non Blume, excl. syn. American Samoa, Fiji, Moluccas, New Caledonia, New Guinea, Nicobar Isls, Solomon Isls, Sulawesi, Philippines. MINDANAO.

Doubtful species

1. Lindsaea decrescens Copel., PJS 81 (1952) 6. “As stated by Copeland, this is intermediate between L. repens (‘L. macraena, L. longa’) and L. fissa. The isotype in MICH from Copeland’s herbarium is rather scrappy. I doubt whether it represents a distinct species” –Kramer, Fl. Males. Ser. ii 1 (1971) 253.

Sphenomeris Maxon: see Odontosoria for the Philippine species

Odontosoria (Pr.) Fée

Last edited by Julie F. Barcelona, 7 September 2020

Photos

1. Odontosoria biflora (Kaulf.) C.Chr., Index Filic. (1905) 207; --Sphenomeris biflora (Kaulf.) Tagawa, J. Jap. Bot. 33 (1958) 203; --Davallia biflora Kaulf., Enum. Fil. (1824) 221. --Sphenomeris chusana sensu Copel. (1960), pro parte. Bonin Isls, China, Hongkong, Izu Isl, Japan, Pacific, Ryukyu Isls, South Korea, Taiwan, Volcano Isls, Philippines. BATANES, LUZON.
2. Odontosoria chinensis (L.) J.Sm., Bot. Voy. Herald 10 (1857) 430; --Sphenomeris chinensis (L.) Maxon, J. Wash. Acad. Sci. 3 (1913) 144; --Trichomanes chinense L., Sp. Pl. 2 (1753) 1099. Photos
   • ssp. chinensis. --Sphenomeris chusana (L.) Copel., Bernice P. Bishop Mus. Bull. 59 (1929) 69; --Adiantum chusanum L., Sp. 2 (1753) 1096. Bangladesh, Bhutan, Borneo, China, India, Laos, Lesser Sunda Isls, Moluccas, Myanmar, Nepal, New Caledonia, Pacific, Palau Isls, Peninsular Malaysia, Solomon Isls, South Korea, Sri Lanka, Sulawesi, Sumatra, Taiwan, Thailand, Tibet, Vietnam, Philippines. MINDANAO: Bukidnon, PALAWAN. Terrestrial or epilithic in exposed sites, from low elevation to 2100m. 
3. Odontosoria krameri Fraser-Jenk., Taxon. Revis. Indian Subcontinental Pteridophytes (2008) 152. --Odontosoria chinensis var. divaricata (Christ) K.U.Kramer, Blumea 15 (1967) 572; --Odontosoria chinensis var. divaricata Christ, Journ. de Bot., ser. 2, 2 (1909) 23. Borneo, ?Cambodia, Indian Ocean, Java, Laos, Myanmar, New Guinea, New Guinea, Sulawesi, Sumatra, Thailand, Vietnam, Philippines. LUZON, MINDANAO.
   
4. Odontosoria retusa (Cav.) J.Sm., Bot. Voy. Herald 10 (1857) 430; --Sphenomeris retusa (Cav.) Maxon, J. Wash. Acad. Sci. 3 (1913) 144; --Davallia retusa Cav., Descr. Pl. (1802) 278. Bismarck Arch, Borneo, Moluccas, New Guinea, Solomon Isls, Sulawesi, Vanuatu, Philippines. MINDANAO: Bukidnon, PALAWAN. Photos  

Osmolindsaea (K.U.Kramer) Lehtonen & Christenh.

Last edited by Julie F. Barcelona, 4 July 2019

Photos

1. Osmolindsaea minor (Hook.) Lehtonen & Christenh., Syst. Bot. 38 (2014) 896; --Lindsaea cultrata var. minor Hook., Sp. Fil. 1 (1846) 204. Borneo, Moluccas, New Guinea, Solomon Isls, Philippines. LUZON.
   
2. Osmolindsaea odorata (Roxb.) Lehtonen & Christenh., Bot. J. Linn. Soc. 163 (2010) 335; --Lindsaea odorata Roxb., Calc. J. Nat. Hist. 4 (1844) 511. --Lindsaea loheriana Christ., Bull. Herb. Boissier 6 (1898) 44. --Lindsaea cultrata sensu Copel. (1960), pro parte. Bangladesh, Bhutan, Borneo, ?Cambodia, China, India, Japan, Java, ?Laos, Lesser Sunda Isls, Moluccas, Myanmar, Nepal, Peninsular Malaysia, Ryukyu Isls, Sri Lanka, Sumatra, Taiwan, Thailand, Tibet, Vietnam, Philippines. LUZON, MINDANAO, MINDORO, NEGROS, PANAY. Photos
3. Osmolindsaea plumula (Ridl.) Lehtonen & Tuomisto, Syst. Bot. 38 (2014) 898; --Lindsaea plumula Ridl., J. Mal. Br. R. As. Soc. 4 (1926) 22. --Lindsaea cultrata var. varia Copel., Fragm. Fl. Philip. (1905) 181. Borneo, Peninsular Malaysia, Philippines. NEGROS.

Tapeinidium (C.Presl) C.Chr.

1. Tapeinidium acuminatum K.U.Kramer, Blumea 15 (1968) 554. Borneo, Philippines. LUZON. Photos  
2. Tapeinidium biserratum (Blume) Alderw., Mal. Ferns Suppl. (1917) 509; --Davallia biserrata Blume, Enum. Pl. Javae 2 (1828) 232. Borneo, Lesser Sunda Isls, New Guinea, Sulawesi, Taiwan, Philippines.
   Probably a natural hybrid. According to Kramer (Fl. Males. Ser. ii 1 (1971): “The type of this ‘species’ is intermediate between T. luzonicum and T. pinnatum; there are 25-30 intermediates… Their status is not clear. They occur throughout the common range of the two species”.
   
3. Tapeinidium calomelanos K.U.Kramer, Blumea 15 (1968) 551. Borneo, ?Sulawesi, ?Sumatra, Philippines. LUZON.
4. Tapeinidium gracile (Blume) Alderw., Malayan Ferns (1909) 315; --Davallia gracilis Blume, Enum. Pl. Javae 2 (1828) 233. Borneo, Java, Lesser Sunda Isls, Moluccas, Sulawesi, Vietnam, Philippines. MINDANAO: Bukidnon, PALAWAN.
5. Tapeinidium luzonicum (Hook.) K.U.Kramer, Blumea 15 (1968) 552. MINDANAO: Bukidnon. Photos
   • var. luzonicum. Borneo, Java, Moluccas, Peninsular Malaysia, Sulawesi, Sumatra, Thailand, Philippines. LUZON, MINDANAO, POLILLO.
     
   • var. leptophyllum K.U.Kramer. Endemic to the Philippines. LEYTE, LUZON, MINDANAO, NEGROS, PANAY.
6. Tapeinidium pinnatum (Cav.) C.Chr., Index Filic. (1905) 213; --Davallia pinnata Cav., Descr. Pl. (Cavanilles) (1802) 277. Borneo, Java, New Guinea, Peninsular Malaysia, Ryukyu Isls, Singapore, Sulawesi, Sumatra, Taiwan, Thailand, ?India, Philippines. MINDANAO: Bukidnon, PALAWAN. Photos  

Lomariopsidaceae 

Cyclopeltis J.Sm.

1. Cyclopeltis crenata (Fée) C.Chr., Index Filic., Suppl. Tert. (1934) 64; --Hemicardion crenatum Fée, Mém. Foug., 5. Gen. Filic. (1850) 283. --Cyclopeltis zamboangana Copel., PJS 81 (1952) 21. Borneo, China, Java, Laos, Myanmar, Peninsular Malaysia, Sulawesi, Sumatra, Thailand, Vietnam, Philippines. Photos
2. Cyclopeltis presliana (J.Sm.) Berkeley, Introd. Crypt. Bot. (1857) 517; --Lastrea presliana J.Sm., J. Bot. (Hooker) 3 (1841) 412. --Cyclopeltis cumingiana (Fée) C.V.Morton, Contr. U.S. Natl. Herb. 38 (1967) 46; --Hemicardion cumingianum Fée, Mém. Foug., 5. Gen. Filic. (1850) 283. Borneo, Moluccas, ?Myanmar, New Guinea, Palau Isls, Sulawesi, Philippines.

Lomariopsis Fée

1. Lomariopsis kingii (Copel.) Holttum, Bot. Mag. (Tokyo) 47 (1933) 273; --Stenochlaena kingii Copel., PJS 6 c (1911) 80. Australia, Moluccas, New Guinea, Sulawesi, Philippines.
2. Lomariopsis lineata (C.Presl) Holttum, Novit. Bot. 1968 (1969) 9; --Olfersia lineata C.Presl, Tent. Pterid. (1836) 235. --Lomariopsis smithii Fée, Mém. Foug., 2. Hist. Acrostich. (18450 71. --Lomariopsis papyracea Copel., PJS 56 (1935) 104. Borneo, Cambodia, China, Java, ?Lesser Sunda Isls, Myanmar, Moluccas, Peninsular Malaysia, Singapore, Sulawesi, Sumatra, Thailand, Vietnam, Philippines. MINDANAO: Bukidnon. Photos
3. Lomariopsis subtrifoliata (Copel.) Holttum, Bot. Mag. (Tokyo) 47 (1933) 274; --Stenochlaena subtrifoliata Copel., PJS 1 (Suppl. 2) (1906) 152. Endemic to the Philippines. LEYTE, LUZON.

Doubtfully occurring in the Philippines

1. Lomariopsis spectabilis (Kunze) Mett., Fil. Hort. Bot. Lips. (1856) 22; --Lomaria spectabilis Kunze, Bot. Zeitung (Berlin) 6 (1848) 144. Borneo, China, Java, Lesser Sunda Isls, Moluccas, Ryukyu Isls, Sulawesi, Sumatra, Taiwan, Vietnam, ?Philippines. Philippine specimens formerly so-named to this species all appear to have narrower fertile pinnae and hence believed by Holttum to be L. kingii. (Fl. Males. ser. 2, 1 (1978) 262. Photos

Lygodiaceae

Lygodium Sw.

1. Lygodium auriculatum (Willd.) Alston, Reinwardtia 5 (1959) 16; --Hydroglossum auriculatum Willd., Sp. 5 (1810) 84. --Lygodium semihastatum (Cav.) Desv., Prodr. (1827) 203, nom. illeg.; --Copel., FFP 1 (1958) 34; --Ugena semihastata Cav. [Ic. Descr. Pl. 6 (1801) 74, nom. illeg. Borneo, Guam, Peninsular Malaysia, Philippines. LUZON, PALAWAN, POLILLO, MINDORO, SAMAR, MINDANAO. Terrestrial, usually in thickets and disturbed forests. Photos  
2. Lygodium circinnatum (Burm.f.) Sw., Syn. Fil. (1806) 153; --Copel., FFP 1 (1958) 33; --Ophioglossum circinnatum Burm.f., Fl. Ind. (1768) 2. --Lygodium basilanicum Christ, PJS 2 c (1907) Bot. 179; --Copel., FFP 1 (1958) 34. Andaman Isls, Bangladesh, Bismarck Arch, Borneo, China, Hong Kong, ?India, Java, Laos, Lesser Sunda Isls, Moluccas, New Guinea, Nicobar Isls, Pacific, Palau Isls, Peninsular Malaysia, Singapore, Solomon Isls, Sri Lanka, Sulawesi, Sumatra, Thailand, Vietnam, Philippines. Throughout the Philippines. Terrestrial, common in brush and open sites and disturbed lowland forests. Photos  
3. Lygodium flexuosum (L.) Sw., J. Bot. (Schrader) 1800 (1801) 106; --Copel., FFP 1 (1958) 35; --Ophioglossum flexuosum L., Sp. Pl. 2 (1753) 1063. --Lygodium scandens (L.) Sw., J. Bot. (Schrader) 1800 (1801) 106; --Copel., FFP 1 (1958) 36; --Ophioglossum scandens L., Sp. Pl. 2 (1753) 1106. Andaman Isls, Australia, Bangladesh, Bhutan, Cambodia, China, India, Laos, Lesser Sunda Isls, Moluccas, Myanmar, Nepal, New Guinea, Peninsular Malaysia, Ryukyu Isls, Singapore, Sri Lanka, Sulawesi, Thailand, Vietnam, Philippines. PALAWAN. Terrestrial, in brush and open forests at low elevation. Photos
4. Lygodium japonicum (Thunb.) Sw., J. Bot. (Schrader) 1800 (1801) 106; --Copel., FFP 1 (1958) 36; --Batuyong et al., Biodiversitas 21 (2020) 3975; --Ophioglossum japonicum Thunb., Syst. Veg. ed. 14 (1784) 926. --Lygodium mearnsii Copel., PJS 3 c (1908) 37; --Copel., FFP 1 (1958) 35. Bangladesh, Bhutan, Cambodia, China, India, Japan, Java, Laos, Laos, Lesser Sunda Isls, Moluccas, Nepal, New Guinea, Pakistan, Ryukyu Isls, South Korea, Sulawesi, Sumatra, Taiwan, Thailand, Tibet, Vietnam, Philippines. LUZON: Ilocos Norte. Terrestrial in open forests and throroughly cleared sites at low to medium elevation. Photos  
5. Lygodium longifolium (Willd.) Sw., J. Bot. (Schrader) 2 (1803) 305; --Hydroglossum longifolium Willd., Abh. Kurfüstl.-Mainz. Akad. Nützl. Wiss. Erfurt 2 (1802) 22. Bangladesh, Borneo, China, India, Myanmar, Peninsular Malaysia, Singapore, Sulawesi, Sumatra, Vietnam, Philippines. ?LUZON, PALAWAN.
6. Lygodium merrillii Copel., PJS 2 c (1907) 146; FFP 1 (1958) 37. --Lygodium matthewii Copel., PJS 3 c (1908) 36; --Copel., FFP 1 (1958) 37. Borneo, China, Sumatra, Vietnam, Philippines. LUZON: Isabela, MINDORO, LEYTE, PALAWAN. Photos
7. Lygodium microphyllum (Cav.) R.Br., Prodr. Fl. Nov. Holl. (1810) 162; --Ugena microphylla Cav., Icon. 6 (1801) 76. Africa, Andaman Isls, Australia, Bangladesh, Bismarck Arch, Borneo, Cambodia, China, India, Indian Ocean, Java, Laos, Lesser Sunda Isls, Moluccas, Myanmar, Nepal, New Caledonia, New Guinea, Pacific, Palau Isls, Peninsular Malaysia, Ryukyu Isls, Singapore, Solomon Isls, Sri Lanka, Sulawesi, Sumatra, Taiwan, Thailand, Vietnam, Philippines. PALAWAN. LUZON to MINDANAO (along the Pacific side), MINDORO, CULION, BASILAN. Terrestrial in more or less open sites. Photos
8. Lygodium salicifolium C.Presl., Suppl. Tent. Pterid. (1845) 102. Andaman Isls, Bangladesh, Bhutan, Borneo, Cambodia, China, India, Java, Laos, Micronesia, Myanmar, Nepal, New Guinea, Palau Isls, Peninsular Malaysia, Singapore, Sumatra, Thailand, Vietnam, Philippines.
9. Lygodium versteegii Christ, Nova Guinea 8 (1909) 161; --Copel., FFP 1 (1958) 37. New Guinea, Solomon Isls, Sulawesi, Philippines. LUZON: Laguna (Mt Makiling), Quezon (Guinayangan), Camarines (Bicol National Park), MINDANAO: Agusan del Norte (Mt Hilong-hilong). Photos  

Marattiaceae

Angiopteris Hoffm.

1. Angiopteris angustifolia C.Presl, Suppl. Tent. Pterid. (1845) 21. Hong Kong, Japan, Moluccas, ?Pacific, Ryukyu Isls, Sulawesi, Sumatra, Taiwan, Thailand, Vietnam, Philippines.
   
2. Angiopteris cartilagidens Christ, Bull. Herb. Boissier 6 (1898) 207. China, Philippines.
   
3. Angiopteris evecta (G.Forst.) Hoffm., Commentat. Soc. Regiae Sci. Gott. 12 (1794) 29; --Sw., Syn. Fil. (1806) 395; --Copel., FFP 1 (1958) 22, in obs., pro syn. A. palmiformis; --Coritico & Amoroso, Nature Conservation Research 5 (2020) 81; --Polypodium evectum G.Forst., Fl. Ins. Austr. (1786) 81. --Angiopteris palmiformis (Cav.) C.Chr., Dansk Bot. Arki 3, 9 (1937) 30; --Copel., FFP 1 (1958) 22; --Clementea palmiformis Cav., Descr. Pl. (1803) 554. --Angiopteris uncinata Christ, PJS 2 c (1907) Bot. 186, non Vriese (1853); --Copel., FFP 1 (1958) 24. --Angiopteris caudata Vriese, Monog. (1853) 20; --Copel., FFP 1 (1958) 24. --Angiopteris cartilagidens Christ, Bull. Boiss. 6 (1898) 207; --Copel., FFP 1 (1958) 24. --Angiopteris pruinosa Kunze, Bot. Zeit. 4 (1846) 417; --Copel., FFP 1 (1958) 25. --?Angiopteris angustifolia C.Presl var. prunifera Hieron., Hedwigia 61 (1919) 257; --Copel., FFP 1 (1958) 25. MINDANAO: Bukidnon, Misamis Occidental, North Cotabato. Conservation status: Other Threatened Species (DAO 2017-11 as Angiopteris palmiformis (Cav.) C.Chr.). Photos
4. Angiopteris smithii Racib., Bull. Int. Acad. Sci. Cracovie 1902 (1902) 54. --Macroglossum alidae Copel., PJS 3 (1909) 342. Borneo, Sulawesi, Philippines. PALAWAN.
5. Angiopteris subcuspidata Rosenst., Meded. Rijks Herb. no. 31 (1917) 3. --Angiopteris elmeriana Copel. in Elmer, LPB 5 (1913) 1679; --Copel., FFP 1 (1958) 23. Endemic to the Philippines.

Christensenia Maxon

1. Christensenia aesculifolia (Blume) Maxon, Proc. Biol. Soc. Wash. 18 (1905) 240; --Copel., FFP 1 (1958) 27; --Coritico & Amoroso, Nature Conservation Research 5 (2020) 82; --Kaulfussia aesculifolia (Blume) Blume, Enum. Pl. Javae 2 (1898) 260; --Aspidium aesculifolium Blume, Enum. Pl. Javae 2 (1898) 143; --Type: Blume? (L51148). Bangladesh, Bismarck Arch, Borneo, China, India, Java, Moluccas, Peninsular Malaysia, Solomon Isls, Sulawesi, Sumatra, Thailand, Vietnam, Philippines. MINDANAO: Bukidnon, Misamis Occidental, PALAWAN. Conservation status: Endangered (DAO 2017-11). Photos
   • ssp. aesculifolia Java. India (Assam) to Malesia.  
2. Christensenia lobbiana (de Vriese) Rolleri, Amer. Fern Jour. 83(1) (1993) 17; --Kaulfussia lobbiana de Vriese & Harting, Monogr. Marattiac. (1853) 14; --Type: from Java. --Christensenia cumingiana Christ, PJS 2 (1907) 186; --Type: Copeland, s.n. (?MICH, holo; ?PNH, iso). Philippines, Borneo, Thailand, Sumatra and Java. Photos. 

Macroglossum Copel. = Angiopteris

Marattia Sw. p.p. = Ptisana

Ptisana Murdock

Last edited by Julie F. Barcelona, 7 September 2020

Photos 

1. Ptisana pellucida (C.Presl) Murdock, Taxon 57 (2008) 747; --Marattia pellucida C.Presl, Suppl. Tent. (1845) 10; --Copel., FFP 1 (1958) 26; --Type: Cuming 177 (L, ?holo; BM, P, US), Luzon: Camarines. --Marattia haenkeana C.Presl, Suppl. Tent. Pterid. (1845) 11; --Type: Haenke s.n. (PR or PRC, holo), Luzon. --Marattia vestita Christ, PJS 2c (1907) 185; --Type: Copeland 1179 (MICH, holo; US, iso). Mindanao; Davao District, Mt Apo, Apr-1904. Sulawesi, Taiwan, Vietnam, Philippines. C LUZON southwards to MINDANAO: Mt. Matutum. Photos  
2. Ptisana sylvatica (Blume) Murdock, Taxon 57 (2008) 747; --Marattia sylvatica Blume, Enum. (1828) 256; --Copel., FFP 1 (1958) 27. Bismarck Arch, Borneo, Caroline Isl, Java, Peninsular Malaysia, Sulawesi, Sumatra, Moluccas, New Guinea, Vanuatu, Philippines. LUZON: Cordillera Central, MINDANAO: Bukidnon, PALAWAN. Common in mossy montane forests to c. 2400m. Photos

Possibly present:

1. Ptisana ternatea (de Vriese) Murdock, Taxon 57 (2008) 747; --Marattia ternatea Christ, PJS 2 c (1907) Bot. 185; --Copel., FFP 1 (1958) 26; --Type: from Moluccas. Bismarck Arch, Moluccas, New Guinea, Santa Cruz Isls, Solomon Isls, Sulawesi, ?Philippines. ?LUZON.

Marsileaceae

Marsilea L.

1. Marsilea crenata C.Presl., Reliq. Haenk. 1 (1825) 84. Australia, ?China, ?Indonesia, New Guinea, ?Peninsular Malaysia, Vietnam, Philippines. Photos
2. Marsilea minuta L., Mant. Pl. (1771) 308. Afghanistan, Africa, Andaman Isls, Australia, Bangladesh, Bhutan, Cambodia, China, India, Indian Ocean, Japan, Java, Laos, Lesser Sunda Isls, Mediterranean, Middle East, Myanmar, Nepal, New Guinea, Pakistan, Peninsular Malaysia, Ryukyu Isls, Sri Lanka, Sulawesi, Taiwan, Thailand, Vietnam, Philippines. Conservation status: Vulnerable (DAO 2017-11).

Possibly present:

1. Marsilea quadrifolia L., Sp. Pl. (1753) 1099. W Asia, China, Europe, Japan, Korea, Mongolia. Reported for the Philippines by Moody (1989) as a weed in rice, but no specimens cited.

Matoniaceae

Matonia R.Br.

1. Matonia foxworthyi Copel., PJS 3 (1908) 343. Borneo, Moluccas, New Guinea, Peninsular Malaysia, Philippines. MINDORO, SIBUYAN. Conservation status: Endangered (DAO 2017-11). Photos

Phanerosorus Copel.

1. Phanerosorus major Diels, Notizbl. Bot. Gart. Berlin-Dahlem 11 (1932) 311. New Guinea, Moluccas, Philippines. SAMAR. Conservation status: Critically Endangered (DAO 2017-11). Photos

Nephrolepidaceae

Nephrolepis Schott

1. Nephrolepis biserrata (Sw.) Schott, Gen. Fil. (1834) pl.3; --Aspidium biserratum Sw., J. Bot. (Schrader) 1800 (1801) 32. --Tectaria fraxinea Cav., Descr. Pl. (1801) 250; --Type: Nee s.n. (BM, fragm., MA), Philippines. --Nephrolepis macrophylla C.Presl, Epimel. Bot. (1851) 43; --Type: Cuming 22 (PRC, holo; BM, iso), Philippines. Africa, Americas, ?Andaman Isls, Australia, Bangladesh, Bismarck Arch, Borneo, Cambodia, China, Hong Kong, India, Indian Ocean, Japan, Java, Lesser Sunda Isls, Myanmar, New Caledonia, New Guinea, Nicobar Isls, Pacific, Peninsular Malaysia, Ryukyu Isls, Solomon Isls, Sri Lanka, Sulawesi, Sumatra, Taiwan, Thailand, Vietnam, Philippines. MINDANAO: Bukidnon, PALAWAN.
2. Nephrolepis brownii (Desv.) Hovenkamp & Miyam., Blumea 50 (2005) 293; --Nephrodium brownii Desv., Mem. Soc. Linn. Paris 6, 2 (1827) 252. --Nephrolepis acutangula C.Presl, Tent. Pterid. (1836) 79. --Nephrolepis pubescens Copel., PJS 81 (1952) 12; --Type: BS 11539 (MICH), Philippines. --Nephrolepis multiflora (Roxb.) Jarrett ex C.V.Morton, Contr. U.S. Natl. Herb. 38 (19740 309; --Davallia multiflora Roxb., Calcutta J. Nat. Hist. 4 (1844) 515. Australia, Bangladesh, Bismarck Arch, Borneo, Cambodia, China, India, Japan, Java, Laos, Lesser Sunda Isls, Myanmar, New Caledonia, New Guinea, Pacific, Peninsular Malaysia, Ryukyu Isls, Singapore, Solomon Isls, Sri Lanka, Sulawesi, Taiwan, Thailand, Vietnam, Philippines. MINDANAO, PALAWAN.
   
3. Nephrolepis cordifolia (L.) C.Presl, Tent. Pterid. (1836) 79; --Polypodium cordifolium L., Sp. Pl. 2 (1753) 1089. --Nephrolepis auriculata (L.) Trimen, J. Linn. Soc. Bot. 24 (1887) 152; --Polypodium auriculatum L., Sp. Pl. 2 (1753) 1088. Australia, Bangladesh, Bhutan, Borneo, Cambodia, China, India, Indian Ocean, Japan, Java, Laos, Lesser Sunda Isls, Myanmar, Nepal, New Caledonia, Palau Isls, Peninsular Malaysia, Ryukyu Isls, South Korea, Sri Lanka, Sulawesi, Taiwan, Thailand, Tibet, Vietnam, Philippines. MINDANAO: Bukidnon. Photos
4. Nephrolepis falcata (Cav.) C.Chr., Dansk Bot. Ark. 9 (1937) 15; --Tectaria falcata Cav., Descr. Pl. (1802) 250; --Type: Née s.n. (MA, holo; BM, iso), Philippines. --Nephrolepis barbata Copel., Fragm. Fl. Philipp. 3 (1905) 178; --Type: Copeland 1286 (MICH), Philippines. Myanmar, Sri Lanka, Sumatra, Vietnam, Philippines. LUZON, MINDANAO: Bukidnon, NEGROS, SAMAR. Photos
5. Nephrolepis hirsutula (G.Forst.) C.Presl, Tent. Pterid. (1836) 79; --Polypodium hirsutulum G.Forst., Fl. Ins. Austr. (1786) 81. Australia, Bismarck Arch, Borneo, Java, Lesser Sunda Isls, Moluccas, Myanmar, New Caledonia, New Guinea, Pacific, Palau Isls, Peninsular Malaysia, Solomon Isls, Sulawesi, Sumatra, Vietnam, Philippines. MINDANAO: Bukidnon, PALAWAN.
6. Nephrolepis radicans (Burm.) Kuhn, Ann. Mus. Bot. Lugd.-Bat. 4 (1869) 285; --Polypodium radicans Burm., Fl. Ind. (1768) 233. --Nephrolepis clementis Christ, PJS 3 c (1908) 272; --Type: Clemens 920 (P), Philippines. --Nephrolepis volubilis (Fée) J.Sm., J. Bot. (Hooker) 3 (1841) 413; --Lepidonevron volubile Fée, Mém. Foug., 5. Gen. Fil. (1852) 301; --Type: Cuming 37 (P, holo; BM, iso), Philippines. Bangladesh, Borneo, Cambodia, India, Java, Laos, Lesser Sunda Isls, Myanmar, New Caledonia, New Guinea, Peninsular Malaysia, Singapore, Sumatra, Thailand, Vietnam, Philippines.
7. Nephrolepis undulata (Afzel. ex Sw.) J.Sm., Bot. Mag. 72 (Companion) (1846) 35 bis; --Aspidium undulatum Afzel. ex Sw., J. Bot. (Schrader) 1800 (1801) 32. --Nephrolepis glabra Copel., PJS 1 (Suppl. 2) (1906) 146; --Type: Copeland 1819 (MICH, holo; KYO, P, SING, iso), Philippines. Africa, Americas, Bangladesh, India, Indian Ocean, Myanmar, Nepal, Thailand, Philippines. LUZON.

Possibly present:

1. Nephrolepis acutifolia (Desv.) Christ, Verh. Naturf. Ges. Basel 11 (1895) 243; --Lindsaea acutifolia Desv., Mem. Soc. Linn. Paris 6, 2 (1827) 312. Africa, Australia, Borneo, Indian Ocean, Java, Lesser Sunda Isls, Moluccas, Myanmar, New Guinea, Pacific, Palau Isls, Peninsular Malaysia, Singapore, Sulawesi, Sumatra, Thailand, Vietnam, ?Philippines.
2. Nephrolepis falciformis J.Sm., Ferns Brit. For. (1866) 287. Borneo, Cambodia, China, India, Laos, Maldives, Moluccas, Myanmar, New Guinea, Peninsular Malaysia, Singapore, Sulawesi, Sumatra, Thailand, Vietnam, ?Philippines.

Oleandraceae

Oleandra Cav.

1. Oleandra cumingii J.Sm., J. Bot. (Hooker) 4 (1842) 413; --Type: Cuming 60 (K, holo; B, BM, SING, US, iso), Luzon. --Oleandra macrocarpa C.Presl, Epimel. Bot. (1851) 41; --Fée, Mém. Foug., 5. Gen. Filic. (1852) 304; --Type: Cuming 60 p.p. (PRC, holo teste Holttum; B, BM, K, SING, US, iso), Luzon. --Oleandra scandens Copel., PJS 46 (1931) 218; --Type: Elmer 6513 (US, holo), Baguio. China, Hong Kong, Laos, Lesser Sunda Isls, Peninsular Malaysia, Thailand, Philippines. LUZON, PALAWAN. Photos
2. Oleandra musifolia Blume, Enum. Pl. Javae (1828) 141 (as 'musaefolium'); --Type: from Indonesia. --Oleandra benguetensis Copel., PJS 46 (1931) 217; --Type: Elmer 6286 (US, holo), Luzon, Baguio. Australia, China, India, Java, Laos, Lesser Sunda Isls, Sri Lanka, Sulawesi, Thailand, Vietnam, Philippines. LUZON.
3. Oleandra neriiformis Cav., Anales Hist. Nat. 2 (1799) 115 (as “neriformis”); --Type: Née s.n. (holo, MA 476029, teste Gariletti), Luzon, Mauban. --Oleandra herrei Copel., PJS 81 (1952) 12; --Type: Herre s.n. (UC, holo). --Oleandra maquilingensis Copel., PJS 46 (1931) 217; --Type: Matthew s.n. (MICH 1210440, lecto). --Oleandra colubrina sensu Copel., FFP 1 (1958) 181, pro parte. --Oleandra nitida (Copel.) Copel., FFP (1958) 181; --Oleandra colubrina var. nitida Copel., PJS 3 c (1908) 33; --Type: Copeland 1474 (US, lecto), Mindanao, Mt. Apo. Bismarck Arch, Borneo, Cambodia, Java, Lesser Sunda Isls, Moluccas, New Guinea, Pacific, Peninsular Malaysia, Solomon Isls, Sulawesi, Sumatra, Thailand, Vietnam, Philippines. MINDANAO: Bukidnon, PALAWAN. Photos
4. Oleandra sibbaldii Grev., Ann. and Mag. Nat. Hist. Ser. 2 1 (1848) 327; --Oleandra cumingii var. sibbaldii (Grev.) Baker, Syn. Fil. (Hooker & Baker) (1868) 303.
   1868; --Type: Sibbald s.n. (E, holo), French Polynesia, Tahiti. Borneo, Moluccas, New Guinea, Pacific, Solomon Isls, Sulawesi, Philippines. MINDANAO: Bukidnon. Photos

Ophioglossaceae

Botrychium Sw.

1. Botrychium daucifolium Wall. ex Hook. & Grev., Icon. Fil. (1829) t.161; --Copel., FFP 1 (1958) 20; --Coritico & Amoroso, Nature Conservation Research 5 (2020) 82; --Sceptridium daucifolium (Wall. ex Hook. & Grev.) Lyon, Bot. Gaz. 40 (1905) 457. Bhutan, Borneo, China, India, Japan, Java, Lesser Sunda Isls, Moluccas, Myanmar, Nepal, Pacific, Peninsular Malaysia, Ryukyu Isls, Sri Lanka, Sulawesi, Sumatra, Vietnam, Philippines. MINDANAO: Bukidnon, Misamis Occidental, North Cotabato. Conservation status: Vulnerable (DAO 2017-11). Photos  
2. Botrychium lanuginosum Wall. ex Hook. & Grev., Icon. Fil. (1828) t.79; --Copel., FFP 1 (1958) 19; --Japanobotrychum lanuginosum (Wall. ex Hook. & Grev.) M.Nishida ex Tagawa, J. Jap. Bot. 33 (1958) 201. Bhutan, China, India, Java, Lesser Sunda Isls, Myanmar, Nepal, New Guinea, Sri Lanka, Sulawesi, Sumatra, Taiwan, Thailand, Tibet, Vietnam, Philippines. Conservation status: Endangered (DAO 2017-11).

Helminthostachys Kaulf.

1. Helminthostachys zeylanica (L.) Hook., Genera (1840) t.47; --Copel., FFP 1 (1958) 20; --Holttum, Rev. Fl. Mal. 2 (1954) 42; --Coritico & Amoroso, Nature Conservation Research 5 (2020) 82; --Osmunda zeylanica L., Sp. Pl. 2 (1753) 1063. Australia, Bangladesh, Bhutan, Bismarck Arch, Borneo, Cambodia, China, India, Japan, ?Java, Laos, Lesser Sunda Isls, Moluccas, Moluccas, Myanmar, Nepal, New Caledonia, New Guinea, Nicobar Isls, Pacific, Palau Isls, Peninsular Malaysia, Ryukyu Isls, Solomon Isls, Sri Lanka, Sulawesi, Sumatra, Taiwan, ?Thailand, Vietnam, Philippines. MINDANAO: Misamis Occidental, PALAWAN. Throughout the Philippines. Terrestrial in open brushy places near sea level on damp ground. Conservation status: Critically Endangered (DAO 2017-11). Photos  

Ophioderma (Blume) Endl.

1. Ophioderma intermedium (Hook.) Nishida, Bull. Nation. Sc. Mus. Tokyo ser. 2, 4 (1959) 335; --Ophioglossum intermedium Hook., Icon. Pl. 10 (1854) t.995; --Copel., FFP 1 (1958) 18. Australia, Borneo, Java, New Guinea, Sumatra, Philippines. Photos
2. Ophioderma pendulum (L.) C.Presl, Suppl. Tent. Pterid. (1845) 56; --Coritico & Amoroso, Nature Conservation Research 5 (2020) 82; --Ophioglossum pendulum L., Sp. Pl., ed. 2. 2 (1763) 1518. Africa, Australia, Bangladesh, Bismarck Arch, Borneo, Cambodia, China, India, Indian Ocean, Japan, Java, Laos, Lesser Sunda Isls, Moluccas, New Guinea, Nicobar Isls, Pacific, Peninsular Malaysia, Ryukyu Isls, Singapore, Solomon Isls, South Korea, Sri Lanka, Sulawesi, Sumatra, Taiwan, Thailand, Vietnam, Philippines. MINDANAO: Bukidnon, Davao Oriental, Misamis Occidental, North Cotabato, PALAWAN.
3. Ophioderma subsessile Amoroso & Coritico, PJS 150 S1 (2020) 216; --Type: Amoroso & Nobleza 18966 (PNH, holo; CMUH, BRIT, iso), Mindanao: Bukidnon Prov, San Fernando, Mt Pantaron Range, 820m, 20-Aug-2020. Endemic to the Philippines. MINDANAO: Bukidnon. Conservation status: Critically Endangered (Amoroso et al. 2020).
   

Ophioglossum L.

1. Ophioglossum costatum R.Br., Prodr. (1810) 163. Africa, Australia, Bangladesh, India, Indian Ocean, Peninsular Malaysia, Sri Lanka, Sulawesi, Sumatra, Tahiti, Thailand, Philippines.
2. Ophioglossum gramineum Willd., Nov. Act. Acad. Erf. 2 (1802) 18. Africa, Australia, Borneo, Cambodia, India, Java, Moluccas, Myanmar, New Guinea, Sri Lanka, Thailand, Vietnam, Philippines.
3. Ophioglossum parvifolium Hook. & Grev., Bot. Misc. 3 (1833) 218. --Ophioglossum nudicaule var. macrorrhizum (Kunze) Clausen, Mem. Torrey Bot. Cl. 19 (1938) 150; --Copel., FFP 1 (1958) 18; --Ophioglossum macrorrhizum Kunze, Farrnkr. 1 (1837) 57. Africa, Australia, Bismarck Arch, China, India, Indian Ocean, Java, Lesser Sunda Isls, Moluccas, Myanmar, Nepal, New Caledonia, New Guinea, Pacific, Palau Isls, Peninsular Malaysia, Singapore, Solomon Isls, Sri Lanka, Sulawesi, Sumatra, Thailand, Vietnam, Volcano Isls, Philippines.
4. Ophioglossum pendulum L., Sp. Pl. ed. 2 (1763) 1518; --Copel., FFP 1 (1958) 19; --Holttum, Rev. Fl. Mal. 2 (1954) 40. Old World tropics from Madagascar through tropical Asia to Polynesia. Throughout the Philippines. Epiphytic in everwet rainforests. Conservation status: Endangered (DAO 2017-11). Photos.
5. Ophioglossum petiolatum Hook., Exot. Fl. (1823) pl.56; --Copel., FFP 1 (1958) 17; --Chandra et al. Taiwania 53 (2008) 203. Australia, Bangladesh, Bhutan, Bismarck Arch, Borneo, China, India, Japan, Java, Lesser Sunda Isls, Moluccas, Myanmar, Nepal, New Caledonia, New Guinea, Pacific, Pakistan, Palau Isls, Peninsular Malaysia, Solomon Isls, Sri Lanka, Sumatra, Taiwan, Thailand, Tibet, Vietnam, Philippines. MINDANAO: Bukidnon.
6. Ophioglossum ramosii Copel., PJS 56 (1935) 97; FFP 1 (1958) 18; --Barcelona et al. PJS 142 (2013) 65; --Type: Ramos BS 14771, Mindanao: Camiguin de Mindanao. Endemic to the Philippines. CAMIGUIN, MINDANAO: Bukidnon (photos). Note by P.B. Pelser (8-Oct-19): here considered distinct from Ophioglossum reticulatum (Barcelona et al. 2013). Photos
7. Ophioglossum reticulatum L., Sp. Pl. (1753) 1063; --Copel., FFP 1 (1958) 17. --Ophioglossum nudicaule L.f. var. grandifolium Clausen, Mem. Torr. Bot. Club 19 (1938) 151; --Copel., FFP 1 (1958) 18. Pantropic. MINDANAO: Bukidnon. Probably throughout the Philippines. Terrestrial in humid mostly shaded sites. Only present during the wet season, 0-1600m. Photos

Osmundaceae

Osmunda L.

1. Osmunda herbacea Copel., PJS 40 (1929) 291; FFP 1 (1958) 31. Endemic to the Philippines. MINDANAO.

Plenasium C.Presl

1. Plenasium banksiifolium (C.Presl) C.Presl, Tent. Pterid. (1836) 110; --Coritico & Amoroso, Nature Conservation Research 5 (2020) 82; --Osmunda banksiifolia (C.Presl) Kuhn, Ann. Lugd.-Bat. 4 (1869) 299; --Copel., FFP 1 (1958) 30. --Osmunda bromeliifolia (C.Presl) Copel., PJS 2 c (1907) Bot. 147; --Copel., FFP 1 (1958) 30. Bonin Isls, Borneo, China, Japan, Java, Russian Far East, Ryukyu Isls, Sulawesi, Taiwan, Philippines. LUZON, MINDANAO: Bukidnon, Davao Oriental, Misamis Occidental, North Cotabato, MINDORO, PANAY. Conservation status: Other Threatened Species (DAO 2017-11, as Osmunda banksiifolium). Photos

Plagiogyriaceae

Plagiogyria (Kunze) Mett.

1. Plagiogyria adnata (Blume) Bedd., Ferns Brit. India (1865) pl.51; --Lomaria adnata Blume, Enum. Pl. Javae 2 (1828) 205. Bhutan, Borneo, China, India, Japan, Java, Myanmar, Peninsular Malaysia, Ryukyu Isls, Sumatra, Taiwan, Thailand, Vietnam, Yakushima Isl, Philippines. LUZON.
2. Plagiogyria egenolfioides (Baker) Copel., J. Straits Branch Roy. Asiat. Soc. 63 (1912) 72; --Lomaria egenolfioides Baker, Kew Bull. 7 (1894).
   
   • var. latipinna (Copel.) X.C.Zhang & Noot., Blumea 42 (1997) 484; --Plagiogyria tuberculata var. latipinna Copel. PJS 2 c (1907) 133. --Plagiogyria tuberculata Copel., PJS 1 (Suppl. 2) (1906) 153. Borneo, Moluccas, Sulawesi, Philippines. MINDANAO: Bukidnon. Photos
3. Plagiogyria euphlebia (Kunze) Mett., Abh. Senckenberg. Naturf. Ges. (Plagiogyria) 2 (1858) 274; --Lomaria euphlebia Kunze, Bot. Zeitung (Berlin) 6 (1848) 521. --Plagiogyria christii Copel., PJS 1 (Suppl. 2) (1906) 153. China, India, Japan, Myanmar, Nepal, Ryukyu Isls, South Korea, Taiwan, Vietnam, Philippines. MINDANAO: Bukidnon. Photos  
4. Plagiogyria falcata Copel., PJS 2 (Suppl. 2) (1907) 133. China, Taiwan, Philippines. LUZON, MINDORO.
5. Plagiogyria glauca (Blume) Mett., Abh. Senckenberg. Naturf. Ges. (Plagiogyria) 2 (1858) 273; --Lomaria glauca Blume, Enum. Pl. Javae (1828) 204. --Plagiogyria nana Copel., PJS 4 (1909) 114; --Plagiogyria glauca var. nana (Copel.) C.Chr., Brittonia 2 (1937) 284. Bismarck Arch, Borneo, China, India, Java, Lesser Sunda Isls, Moluccas, Myanmar, New Guinea, Solomon Isls, Sulawesi, Sumatra, Taiwan, Tibet, Philippines. LUZON, MINDANAO: Bukidnon. Photos
6. Plagiogyria pycnophylla (Kunze) Mett., Farngatt. 2 (1858) 272; --Lomaria pycnophylla Kunze, Bot. Zeitung (Berlin) 6 (1848) 143. Bhutan, Borneo, China, India, Java, Myanmar, Nepal, New Guinea, Sulawesi, Sumatra, Taiwan, Thailand, Tibet, Vietnam, Philippines. LUZON, MINDANAO: Bukidnon. Photos
7. Plagiogyria stenoptera (Hance) Diels, Nat. Pflanzenfam. 1 (1899) 282; --Blechnum stenopterum Hance, J. Bot. 21 (1883) 268. China, Japan, Taiwan, Vietnam, Philippines. LUZON.

Polypodiaceae

including Drynariaceae, Grammitidaceae, Loxogrammaceae, Platyceriaceae

Acrosorus Copel.

1. Acrosorus friderici-et-pauli (Christ) Copel., PJS 1 (Suppl. 2) (1906) 159; --Davallia friderici-et-pauli Christ, Verh. Naturf. Ges. Basel 11 (1895) 202. --Acrosorus exaltatus Copel., PJS 1 (Suppl. 2) (1906) 159. --Acrosorus merrillii Copel., PJS 2 c (1907) 136. --Acrosorus symmetricus Copel., PJS 81 (1952) 44. --Acrosorus triangularis (Scort. ex Bedd.) Copel., PJS 3 c (1909) 347; --Polypodium triangulare Scort. ex Bedd., J. Bot. (1887) 324, nom. illeg., auct. non Gmel. Peninsular Malaysia, Borneo, New Guinea, Thailand, Moluccas, Sulawesi, Vietnam, Philippines. MINDORO, NEGROS.

Possibly present:

1. Acrosorus streptophyllus (Baker) Copel., PJS 56 (1935) 480; --Polypodium streptophyllum Baker, J. Bot. 17 (1879) 42. Borneo, Moluccas, New Guinea, Peninsular Malaysia, Sulawesi, Vietnam, ?Philippines. ?MINDANAO. Recorded for the Philippines by Price (1972); some material so identified belongs to Acrosorus friderici-et-pauli. Occurrence in the Philippines needs confirmation.

Aglaomorpha = Drynaria (Bory) J.Sm.

 Anapausia C.Presl  = Leptochilus

Arthromeris (T.Moore) J.Sm. = Selliguea

Belvisia Mirbel = Lepisorus

Bosmania Testo

Last edited by Julie F. Barcelona, 22 August 2019

1. Bosmania membranacea (D.Don) Testo, Syst. Bot. 44 (2019) 8; --Microsorum membranaceum (D.Don) Ching, Bull. Fan Mem. Inst. Biol. 4 (1933) 309; --Polypodium membranaceum D.Don, Prodr. Fl. Nepal. (1825) 2; --Type: from Nepal. --Microsorum schneideri (Christ) Copel., Gen. Fil. (1947) 197; --Polypodium schneideri Christ, Bull. Herb. Boissier 6 (1898) 835. Bhutan, China, India, Jammu & Kashmir, Japan, Laos, Myanmar, Nepal, Sri Lanka, Sulawesi, Taiwan, Thailand, Tibet, Vietnam, Philippines. LUZON: Benguet.

Calymmodon C.Presl

Last edited by Julie F. Barcelona, 7 September 2020

1. Calymmodon clavifer (Hook.) T.Moore, Index Fil. 10 (1861) 219; --Grammitis clavifer Hook. Bismarck Arch, Borneo, Moluccas, New Guinea, Sulawesi, Philippines.
2. Calymmodon cucullatus (Nees & Blume) C.Presl, Tent. Pterid. (1836) 204; --Polypodium cucullatum Nees & Blume, Nova Acta Regiae Soc. Sci. Upsal. 11 (1823) 121. --Calymmodon ordinatus Copel., PJS 34 (1927) 267. Australia, Bismarck Arch, Borneo, Japan, Java, ?Lesser Sunda Isls, Moluccas, New Caledonia, New Guinea, New Guinea, Pacific, Peninsular Malaysia, Solomon Isls, Sulawesi, Sumatra, Taiwan, Thailand, Vietnam, Philippines. MINDANAO: Bukidnon. Photos
3. Calymmodon gracilis (Fée) Copel., PJS 34 (1927) 266; --Plectopteris gracilis Fée, Mém. Foug. 5 (1852) 230. ?Australia, Borneo, Moluccas, Peninsular Malaysia, Sulawesi, Taiwan, ?Thailand, Vietnam, Philippines. PALAWAN. Photos
4. Calymmodon muscoides (Copel.) Copel., PJS 34 (1927) 264; --Polypodium muscoides Copel., LPB 3 (1910) 839. Borneo, Philippines. MINDANAO. Photos

Christopteris Copel. = Drynaria

Chrysogrammitis Parris

1. Chrysogrammitis glandulosa (J.Sm.) Parris, Kew Bull. 53 (1998) 912; --Ctenopteris glandulosa J.Sm., Hist. Fil. (1875) 185. --Ctenopteris merrittii (Copel.) Tagawa, Acta Phytotax. Geobot. 9 (1940) 210; --Polypodium merrittii Copel., PJS 1(Suppl. 4) (1906) 255. Borneo, Peninsular Malaysia, Sri Lanka, Sumatra, Taiwan, Philippines. MINDORO. Photos
2. Chrysogrammitis musgraviana (Baker) Parris, Kew Bull. 53 (1998) 914; --Polypodium musgravianum Baker, J. Bot. 28 (1890) 108; --Chen et al., Phytotaxa 443 (2020) 122; --Type: from New Guinea. --Xiphopteris apoensis (Copel.) Copel., Gen. Fil. (1947) 215; --Polypodium apoense Copel., PJS 56 (1935) 105. Bismarck Arch, Borneo, Moluccas, New Guinea, Peninsular Malaysia, Solomon Isls, Sulawesi, Vanuatu, Vietnam, Philippines. LUZON: Rizal.

Ctenopteris Blume = Chrysogrammitis, Ctenopterella, Dasygrammitis,  Prosaptia, Tomophyllum

  Christiopteris Copel. = Drynaria

 Colysis C.Presl = Leptochilus

  Crypsinus C.Presl = Selliguea

Ctenopterella Parris

1. Ctenopterella blechnoides (Grev.) Parris, Gard. Bull. Sing. 58 (2007) 235; --Ctenopteris blechnoides (Grev.) Wagner & Grether, Univ. Calif. Publ. Bot. 23 (1948) 61; --Grammitis blechnoides Grev. -- Ann. Mag. Nat. Hist. II. 1 (1848) 320. Australia, Bismarck Arch, Borneo, Cambodia, China, India, Java, ?Lesser Sunda Isls, Micronesia, Moluccas, New Caledonia, New Guinea, Pacific Isls, Palau Isls, Peninsular Malaysia, Singapore, Solomon Isls, Sri Lanka, Sulawesi, Sumatra, Thailand, Philippines. PALAWAN. Photos
2. Ctenopterella denticulata (Blume) Parris, Gard. Bull. Singapore 58 (2007) 235; --Grammitis denticulata Blume, Fl. Javae Fil. (1828) 121. --Ctenopteris solida Kunze in Zoll., Syst. Verz. (1854) 37. Borneo, Java, Lesser Sunda Isls, Moluccas, New Guinea, Sulawesi, ?Vietnam, Philippines.
3. Ctenopterella pediculata (Baker) Parris, Gard. Bull. Sing. 58 (2007) 237; --Ctenopteris pediculata (Baker) Copel., Fern Fl. Philip. 3 (1960) 530; --Polypodium pediculatum Baker, Syn. Fil. (Hooker & Baker) (1868) 455. Borneo, Philippines. MINDANAO.

Dasygrammitis Parris

1. Dasygrammitis fuscata (Blume) Parris, Gard. Bull. Sing. 58 (2007) 239; --Ctenopteris fuscata (Blume) Kunze. Borneo, Peninsular Malaysia, Sulawesi, Sumatra, Philippines.
2. Dasygrammitis malaccana (Baker) Parris, Gard. Bull. Singapore 58 (2007) 239; --Polypodium malaccanum Baker, Ann. Bot. (Oxford) 8 (1894) 129. --Ctenopteris matutumensis Copel., FFP 3 (1960) 528, anglice, non rite publice; --Type: Copeland s.n. (MICH, holo), Mindanao: Mt Matutum. --Ctenopteris pachycaulis (Copel.) Copel., Fern Fl. Philip. 3 (1960) 529; --Polypodium pachycaulum Copel., PJS 56 (1935) 105; --Type: Copeland s.n. (MICH, UC), Mindanao: Mt Apo. Peninsular Malaysia, Philippines. MINDANAO.
3. Dasygrammitis mollicoma (Nees & Blume) Parris, Gard. Bull. Sing. 58 (2007) 239; --Ctenopteris mollicoma (Nees & Blume) Kunze, Bot. Zeitung (Berlin) 4 (1846) 425; --Polypodium mollicomum Nees & Blume, Nova Acta 11 (1923) 121. Borneo, Java, Lesser Sunda Isls, Moluccas, New Guinea, Peninsular Malaysia, Sri Lanka, Sulawesi, Sumatra, Taiwan, Thailand, Vietnam, Philippines. Photos

Dendroconche Copel.

1. Dendroconche linguiforme (Mett.) Testo, Sundue & Field, Syst. Bot. 44 (2019) 11; --Microsorum linguiforme (Mett.) Copel., Univ. Calif. Publ. Bot. 16 (1929) 111; --Polypodium linguiforme Mett., Ann. Mus. Bot. Lugduno-Batavi 2 (1866) 228; --Type: from Indonesia. Bismarck Arch, Borneo, Moluccas, New Guinea, Pacific, Solomon Isls, Sulawesi, Sumatra, Philippines.

Dendroglossa C.Presl = Leptochilus

 Diblemma J.Sm. = Microsorum

 Drymoglossum C.Presl = Pyrrosia

Drynaria (Bory) J.Sm., nom. cons.

Last edited by Julie F. Barcelona, 11 July 2021

1. Drynaria cornucopia (Copel.) Alderw., Bull. Dépt. Agric. Indes Néerl. 21 (1908) 8; --Aglaomorpha cornucopia (Copel.) M.C.Roos, Blumea 31 (1985) 153; --Coritico & Amoroso, Nature Conservation Research 5 (2020) 82; --Thayeria cornucopia Copel., PJS 1 (1906) Suppl. 165, t.28. Endemic to the Philippines. LUZON: Abra, Benguet, Kalinga, Mountain Province, MINDANAO: Bukidnon, Davao, Misamis Occidental, North Cotabato, Zamboanga. Conservation status: Vulnerable (DAO 2017-11; as Aglaomorpha cornucopia). Photos
2. Drynaria descensa Copel., PJS 3 c (1908) 36; --Aglaomorpha descensa (Copel.) Hovenkamp & S.Linds., Gard. Bull. Singapore 69 (2017) 150; --Type: Copeland 2061, but also labelled Pteridophyta Philippinensia Exsiccatae 42 (MICH, lecto; B, BM, G, K, L, MU, NSW, S, UC), Luzon: Nueva Ecija. Endemic to the Philippines. LUZON: Nueva Ecija. Photos
3. Drynaria heraclea (Kunze) T.Moore, Index Fil. (1862) 346; --Aglaomorpha heraclea (Kunze) Copel., Univ. Calif. Publ. Bot. 16 (1929) 117; --Coritico & Amoroso, Nature Conservation Research 5 (2020) 82; --Drynariopsis heraclea (Kunze) Ching, Sunyatsenia 5 (1940) 262; --Polypodium heracleum Kunze, Bot. Zeitung (Berlin) 6 (1848) 117. Bismarck Arch, Borneo, Java, Lesser Sunda Isls, Moluccas, New Guinea, Peninsular Malaysia, Solomon Isls, Sulawesi, Sumatra, Thailand, Philippines. MINDANAO: Bukidnon, Misamis Occidental, North Cotabato. Conservation status: Vulnerable (DAO 2017-11; as Aglaomorpha heraclea). Photos
4. Drynaria meyeniana (Schott) Christenh., Phytotaxa 230 (2015) 299; --Aglaomorpha meyeniana Schott, Gen. Fil. (1834) pl.20. Taiwan, Philippines. CATANDUANES, CEBU, LUZON, MINDORO. Conservation status: Vulnerable (DAO 2017-11, as Aglaomorpha meyeniana). Photos
5. Drynaria pilosa (J.Sm. ex Kunze) Christenh., Phytotaxa 230 (2015) 299; --Aglaomorpha pilosa (J.Sm. ex Kunze) Copel., PJS 6 c (1911) 141; --Dryostachyum pilosum J.Sm. ex Kunze, Die Farrnkräuter in kolorirten Abbildungen naturgetreu erläutert und beschrieben 1 (1844) 139. Moluccas, Sulawesi, Philippines. LUZON, MINDANAO, NEGROS. Conservation status: Vulnerable (DAO 2017-11; as Aglaomorpha pilosa). Photos
6. Drynaria quercifolia (L.) J.Sm., J. Bot. (Hooker) 3 (1841) 398; --Aglaomorpha quercifolia (L.) Hovenkamp & S. Linds., Gard. Bull. Singapore 69 (2017) 152; --Polypodium quercifolium L., Sp. Pl. 2 (1753) 1087; --Type: from India. India and China, across Malesia to tropical Australia and Fiji, Philippines. MINDANAO: Bukidnon. Throughout the Philippines. A very common epiphyte or lithophyte in seasonal areas, in open forests, about settlements, etc., usually at low elevation. Photos
7. Drynaria rigidula (Sw.) Bedd., Ferns Brit. India (1869) pl.314; --Aglaomorpha rigidula (Sw.) Hovenkamp & S. Linds., Gard. Bull. Singapore 69 (2017) 153; --Polypodium rigidulum Sw., J. Bot. (Schrader) 1800 (1801) 26; --Type: from Java. Australia, Bismarck Arch, Cambodia, China, Java, Laos, Lesser Sunda Isls, Moluccas, Myanmar, New Caledonia, New Guinea, Pacific, Peninsular Malaysia, Solomon Isls, Sulawesi, ?Sulawesi, Sumatra, Thailand, Vietnam, Philippines. Photos
8. Drynaria sagitta (Christ) Christenh., Phytotaxa 230 (2015) 299; --Aglaomorpha sagitta (Christ) Hovenkamp & S.Linds., Gard. Bull. Singapore 69 (2017) 153; --Coritico & Amoroso, Nature Conservation Research 5 (2020) 82; --Christopteris sagitta (Christ) Copel., Fragm. Fl. Philipp. (1905) 188; --Selliguea sagitta (Christ) Fraser-Jenk., Taxon. Revis. Indian Subcontinental Pteridophytes (2008) 45; --Polypodium sagittum Christ, Bull. Herb. Boissier 6 (1898) 199; --Type: Loher s.n. (P, lecto), Luzon: La Trinidad. --Christopteris copelandii Christ, Bull. Herb. Boissier II. 6. (1906) 990. Endemic to the Philippines. LUZON, MINDANAO: Bukidnon, Misamis Occidental, North Cotabato. Conservation status: Vulnerable (DAO 2017-11 as Selliguea sagitta (Christ) Fraser-Jenk.). Photos
9. Drynaria sparsisora (Desv.) T.Moore, Index Fil. (1862) 348; --Aglaomorpha sparsisora (Desv.) Hovenkamp & S.Linds., Gard. Bull. Singapore 69 (2017) 153; --Polypodium sparsisorum Desv., Ges. Naturf. Freunde Berlin Mag. Neuesten Entdeck. Gesammten Naturk. 5 (1811) 315; --Type: from New Guinea. Australia, Bangladesh, Bismarck Arch, Borneo, Cambodia, Java, Lesser Sunda Isls, Moluccas, Myanmar, New Guinea, Peninsular Malaysia, Singapore, Solomon Isls, Sri Lanka, Sulawesi, Sumatra, Thailand, Vietnam, Philippines. PALAWAN. Photos
10. Drynaria speciosa (Blume) Christenh., Phytotaxa 230 (2015) 299; --Aglaomorpha speciosa (Blume) M.C.Roos, Phylogen. Syst. Dryn. (1985) 244; --Coritico & Amoroso, Nature Conservation Research 5 (2020) 82; --Photinopteris speciosa (Blume) C.Presl, Abh. Königl. Böhm. Ges. Wiss., ser. 5 6 (1851) 624; --Lomaria speciosa Blume, Enum. Pl. Javae 2 (1828) 202. --Aglaomorpha acuminata (Willd.) Hovenkamp, Fl. Males., Ser. 2, Pterid. 3 (1998) 13; --Photinopteris acuminata (Willd.) C.V.Morton, Contr. U.S. Natl. Herb. 38 (1963) 31; --Acrostichum acuminatum Willd., Sp. Pl. 5 (1810) 116. Borneo, Cambodia, China, Java, Laos, Peninsular Malaysia, Singapore, Sulawesi, Sumatra, Thailand, Vietnam, Philippines. MINDANAO: Davao Oriental. Conservation status: Vulnerable (DAO 2017-11; as Aglaomorpha acuminata). Photos
    
11. Drynaria aglaomorpha Christenh., Phytotaxa 230 (2015) 299; --Aglaomorpha splendens (J.Sm.) Copel., PJS 6 c (1911) 141; --Coritico & Amoroso, Nature Conservation Research 5 (2020) 82. --Dryostachyum splendens J.Sm., J. Bot. (Hooker) 3 (1841) 399. Sulawesi, Philippines. LUZON, MINDANAO: Bukidnon, North Cotabato, NEGROS. Conservation status: Vulnerable (DAO 2017-11; as Aglaomorpha splendens). Photos

Drynariopsis (Copel.) Ching = Drynaria

Goniophlebium (Blume) C.Presl

1. Goniophlebium benguetense (Copel.) Copel., Fern Fl. Philip. 3 (1961) 461; --Schellolepis benguetensis (Copel.) Pic.-Serm., Webbia 28 (1973) 470; --Polypodium benguetense Copel., PJS 1 (Suppl. 2) (1906) 256. Moluccas, Philippines. LUZON, MINDANAO. Photos
2. Goniophlebium coadunatum Barcelona & M.G.Price, Fern Gaz. 15 (1999) 261. Endemic to the Philippines.
3. Goniophlebium mengtzeense (Christ) Rödl-Linder, PJS 116 (1987) 154; --Polypodium mengtzeense Christ, Bull. Herb. Boissier 6 (1898) 869. Cambodia, China, India, Laos, Taiwan, Thailand, Vietnam, Philippines.
4. Goniophlebium percussum (Cav.) Wagner & Grether, Occ. Pap. Bishop Mus. 19 (1948) 88; --Schellolepis percussa (Cav.) Pic.Serm., Webbia 28 (1973) 470; --Cyathea percussa Cav., Descr. Pl. (1803) 548. Australia, Borneo, Java, Moluccas, New Guinea, Peninsular Malaysia, Singapore, Sulawesi, Sumatra, Thailand, Philippines.
5. Goniophlebium persicifolium (Desv.) Bedd., Ferns br. Ind. Correct. (1870); --Schellolepis persicifolia (Desv.) Pic.-Serm., Webbia 28 (1973) 470; --Polypodium persicifolium Desv., Berl. Mag. 5 (1811) 316. Borneo, China, Java, Lesser Sunda Isls, Moluccas, New Guinea, Pacific, Peninsular Malaysia, Solomon Isls, Sulawesi, Sumatra, Thailand, Vietnam, Philippines. MINDANAO: Bukidnon. Photos
6. Goniophlebium pseudoconnatum (Copel.) Copel., Fern Fl. Philip. 3 (1960) 462; --Polypodium pseudoconnatum Copel., PJS 1 (Suppl. 2) (1906) 161. Borneo, Moluccas, New Guinea, Sulawesi, Philippines. LUZON, MINDANAO: Bukidnon. Photos
7. Goniophlebium subauriculatum (Blume) C.Presl, Tent. Pterid. (1836) 186; --Polypodium subauriculatum Blume, Enum. Pl. Javae 2 (1828) 133. --Goniophlebium tomentellum (C.Chr.) Copel., Fern Fl. Philip. 3 (1960) 461; --Schellolepis tomentella (C.Chr.) Pic.-Serm., Webbia 28 (1973) 470; --Polypodium tomentellum C.Chr., Index Filic. (1906) 570. --Goniophlebium integrum Copel., PJS 81 (1952) 42. Australia, Borneo, Cambodia, China, India, Java, Laos, Lesser Sunda Isls, Moluccas, Myanmar, New Caledonia, New Guinea, New Guinea, Peninsular Malaysia, Singapore, Solomon Isls, Sulawesi, Sumatra, Thailand, Vietnam, Philipines. LUZON, MINDANAO: Bukidnon. Photos
8. Goniophlebium terrestre Copel., PJS 56 (1935) 106; --Schellolepis terrestris (Copel.) M.G.Price, Kalikasan 3 (1974) 178. Endemic to the Philippines. LUZON: ?Isabela, Laguna, Quirino. Conservation status: Critically Endangered (DAO 2017-11). Photos
   

 Grammitis auctt. Males., non Sw. = Oreogrammitis

Lecanopteris Reinw.

Last edited by Julie F. Barcelona, 11 October 2020

1. Lecanopteris deparioides (Cesati) Baker, J. Bot. 19 (1881) 366; --NPfl. 328; --Coritico & Amoroso, Nature Conservation Research 5 (2020) 82. --Lecanopteris curtisii Baker, J. Bot. (1881) Icon. Plant, pl. 1607; --H Gay, Gard. Bull. Sing. 45 (1993) 301. --Lecanopteris pumila Copel., Polyp. Philipp. (1905) 134; Fern Fl. Philip. 3 (1960) 499, non Blume (1829.)-- Lecanopteris philippinensis Alderw., Bull. Jard. Bot. Buitenz. Ser. 2, no. 1 (1911) 8. Sumatra, Peninsular Malaysia, Philippines to the Moluccas. MINDANAO: Bukidnona, Lanao del Sur, Davao, Davao Oriental, Misamis Occidental. 800-1200m. Conservation status: Endangered (DAO 2017-11). Photos
2. Lecanopteris luzonensis Hennipman, Blumea 32 (1987) 311. Endemic to the Philippines. LUZON: Pangasinan, Pampanga, Bataan (Mt Mariveles), Laguna, Quezon, Camarines Norte. Conservation status: Endangered (DAO 2017-11). Photos
3. Lecanopteris sarcopus (Teijsm. & Binn.) Copel., Univ. Calif. Publ. Bot. 16 (1929) 123; --Polypodium sarcopum Teijsm. & Binn., Tijdschr. Nederl. Ind. 29 (1867) 241. --Lecanopteris lomarioides (J.Sm.) Copel., Univ. Calif. Publ. Bot. 16 (1929) 123, nom. inval.; --Drynaria lomarioides J.Sm., J. Bot. 3 (1841) 397, nom. inval. Sulawesi, Philippines. LUZON: Nueva Ecija, Nueva Vizcaya. Conservation status: Endangered (DAO 2017-11).
   
4. Lecanopteris sinuosa (Wall. ex Hook.) Copel., Gen. Fil. (1947) 205; --Coritico & Amoroso, Nature Conservation Research 5 (2020) 82; --Polypodium sinuosum Wall. ex Hook., Sp. Fil. 5 (1863) 61. Australia, Bismarck Arch, Borneo, Cambodia, Java, Lesser Sunda Isls, Moluccas, New Guinea, Peninsular Malaysia, Singapore, Solomon Isls, Sulawesi, Sumatra, Thailand, Vanuatu, Vietnam, Philippines. MINDANAO: Davao Oriental, PALAWAN. Conservation status: Endangered (DAO 2017-11). Photos

Lemmaphyllum C.Presl

1. Lemmaphyllum carnosum (J.Sm. ex Hook.) C.Presl, Epimel. Bot. (1851) 158; -- Drymoglossum carnosum (Wall.) J.Sm. ex Hook., Gen. Fil. (1841) t.78A. Photos
   • var. carnosum. --?Lemmaphyllum spathulatum (C.Presl) C.Presl, Epimel. Bot. (1851) 158. ?Cambodia, China, India, ?Laos, Myanmar, Thailand, Vietnam, Philippines. LUZON

Lepisorus (J.Sm.) Ching

1. Lepisorus accedens (Blume) Hosok., Trans. Nat. Hist. Soc. Formosa 31 (1941) 477; --Lemmaphyllum accedens (Blume) Donk, Reinwardtia 2 (1954) 409; --Weatherbya accedens (Blume) Copel., Gen. Fil. (1947) 191; --Polypodium accedens Blume, Enum. Pl. Javae 2 (1828) 121. Australia, Bismarck Arch, Borneo, Fiji, Java, Lesser Sunda Isls, Micronesia, Moluccas, New Guinea, Peninsular Malaysia, Samoa, Solomon Isls, Sulawesi, Sumatra, Thailand, Philippines. MINDANAO: Bukidnon. Photos
   
2. Lepisorus longifolius (Blume) Holttum, Rev. Fl. Malaya 2 (1955) 151; --Pleopeltis longifolia Blume, Enum. Pl. Javae 2 (1828) Add.; --Paragramma longifolia (Blume) T.Moore, Ind. Fil. 1 (1857) 32. Borneo, Java, Myanmar, ?New Guinea, Peninsular Malaysia, Sulawesi, Sumatra, Vietnam, Philippines. PALAWAN. Photos
3. Lepisorus mucronatus (Fée) Li Wang, Bot. J. Linn. Soc. 162 (2010): 35; --Belvisia mucronata (Fée) Copel., Gen. Fil. (1947) 192; --Hymenolepis mucronata Fée, Gen. Fil. (1851-1852) 82. Australia, Bismarck Arch, Borneo, Cambodia, China, Java, Laos, Lesser Sunda Isls, Moluccas, New Caledonia, New Guinea, Pacific, Palau Isls, Peninsular Malaysia, Solomon Isls, Sri Lanka, Sulawesi, Sumatra, Taiwan, Thailand, Vietnam, Philippines. MINDANAO: Bukidnon, PALAWAN.
4. Lepisorus platyrhynchos (Kunze) Li Wang, Bot. J. Linn. Soc. 162 (2010) 35; --Coritico & Amoroso, Nature Conservation Research 5 (2020) 82; --Belvisia platyrhynchos (Kunze) Copel., Gen. Fil. (1947) 192; --Hymenolepis platyrhynchos Kunze, Farrnkräuter 1 (1842) 111. --Belvisia glauca (Copel.) Copel., Gen. Fil. (1947) 192; --Hymenolepis platyrhynchos Kunze var. glauca Copel., LPB 3 (1910) 847; --Hymenolepis glauca (Copel.) C.Chr., Dansk Bot. Ark. 6 (1929) 62. New Guinea, Philippines. LUZON, MINDANAO: Agusan del Norte, Bukidnon, Davao, North Cotabato, MINDORO. Conservation status: Endangered (DAO 2017-11). Photos
5. Lepisorus spicatus (L.f.) Li Wang, Bot. J. Linn. Soc. 162 (2010) 35; --Belvisia spicata (L.f.) Mirb. ex Copel., Gen. Fil. (1947) 192; --Acrostichum spicatum L.f., Suppl. (1781) 444. --Belvisia revoluta (Blume) Copel., Gen. Fil. (1947) 192; --Hymenolepis revoluta Blume, Enum. Pl. Javae 2 (1828) 201. Africa, Borneo, Indian Ocean, Java, Lesser Sunda Isls, Moluccas, New Caledonia, New Guinea, Pacific, Peninsular Malaysia, Solomon Isls, Sri Lanka, Sulawesi, Sumatra, Thailand, Vietnam, Philippines. MINDANAO: Bukidnon, PALAWAN.
6. Lepisorus thunbergianus (Kaulf.) Ching, Bull. Fan Mem. Inst. Biol. 4 (1933) 88; --Pleopeltis thunbergiana Kaulf., Wesen Farrenk. (1827) 113. China, Hawaii, India, Jammu & Kashmir, Japan, Laos, Myanmar, Nepal, Ryukyu Isls, South Korea, Taiwan, Tibet, Vietnam, Philippines. LUZON.
7. Lepisorus validinervis (Kunze) Li Wang, Bot. J. Linn. Soc. 162 (2010) 36; --Belvisia validinervis (Kunze) Copel., Gen. Fil. (1947) 192; --Hymenolepis validinervis Kunze, Bot. Zeitung (Berlin) 6 (1848) 122. Bismarck Arch, Borneo, Java, Lesser Sunda Isls, Moluccas, New Guinea, Peninsular Malaysia, Sulawesi, Sumatra, Vanuatu, Philippines. Photos

Leptochilus Kaulf.

1. Leptochilus axillaris (Cav.) Kaulf., Enum. Filic. (1824) 147; --Acrostichum axillare Cav., Ann. Hist. Nat. 1 (1799) 101. Bangladesh, Borneo, China, India, Java, Lesser Sunda Isls, Moluccas, Myanmar, New Guinea, Nicobar Isl, Peninsular Malaysia, Sulawesi, Sumatra, Thailand, Vietnam, Philippines. PALAWAN. 
2. Leptochilus bolsteri (Copel.) Parris, Fern Gaz. 20 (2018) 305; --Colysis bolsteri (Copel.) Copel., Fern Fl. Philip. 3 (1960) 489; --Polypodium bolsteri Copel., PJS 1(Suppl. 4) (1906) 257. Borneo, Philippines. MINDANAO.
   
3. Leptochilus decurrens Blume, Enum. Pl. Javae 2 (1828) 206; --Anapausia decurrens (Blume) C.Presl, Epimel. Bot. (1851) 186. Bhutan, Bonin Isl, Borneo, ?Cambodia, China, Christmas Isl, India, Japan, Java, Laos, Lesser Sunda Isls, Moluccas, Myanmar, Nepal, New Guinea, Peninsular Malaysia, South Korea, Sri Lanka, Sulawesi, Sumatra, Taiwan, Thailand,  Vietnam, Philippines. MINDANAO: Bukidnon.
4. Leptochilus dolichopterus (Copel.) Fraser-Jenk. & Amoroso, Taxon. Revis. Indian Subcontinental Pteridophytes (2008) 62; --Polypodium dolichopterum Copel., PJS 1 (Suppl. 2) (1906) 162. Endemic to the Philippines.
   
5. Leptochilus ellipticus (Thunb.) Noot., Blumea 42 (1997) 283; --Colysis elliptica (Thunb.) Ching, Bull. Fan Mem. Inst. Biol. 4 (1933) 333; --Polypodium ellipticum Thunb., Fl. Jap. (Thunberg) (1784) 335. Australia, Bhutan, China, Hong Kong, India, Japan, Laos, Myanmar, Nepal, Ryukyu Isls, South Korea, Taiwan, Thailand, Tibet, Vietnam, Philippines. LEYTE, LUZON, MINDANAO, SAMAR. Photos
6. Leptochilus fluviatilis (Lauterb.) Liang Zhang & Li Bing Zhang, Phytotaxa 374 (2018) 173; --Leptochilus macrophyllus var. fluviatilis (Lauterb.) Noot., Blumea 42 (1997) 289; --Polypodium fluviatile Lauterb., Bot. Jahrb. Syst. 44 (1910) 507; --Type: from Borneo. Borneo, Sulawesi, Philippines.
   
7. Leptochilus hemionitideus (C.Presl) Noot., Blumea 42 (1997) 285; --Selliguea hemionitidea C.Presl, Tent. Pterid. (1836) 216. Bangladesh, Bhutan, China, India, Japan, Myanmar, Nepal, Ryukyu Isls, Taiwan, Thailand, Tibet, Vietnam, Philippines. Photos
8. Leptochilus insignis (Blume) Fraser-Jenk., Taxon. Revis. Indian Subcontinental Pteridophytes (2008) 62; --Microsorum insigne (Blume) Copel., Univ. Calif. Publ. Bot. 16. (1929) 112; --Polypodium insigne Blume, Enum. Pl. Javae 2 (1828) 127. --Microsorum decurrens (Brack.) Copel., Fern Fl. Philip. 3 (1960) 481; --Drynaria decurrens Brack., U.S. Expl. Exped., Filic. 16 (1854) 48. --Microsorum palmatum auct. non (Blume) Fée. Bangladesh, Bhutan, Borneo, ?Cambodia, China, India, Japan, Java, ?Laos, Lesser Sunda Isls, Myanmar, Nepal, Nicobar Isls, Peninsular Malaysia, Ryukyu Isls, Sri Lanka, Sumatra, Taiwan, Thailand, Tibet, Vietnam, Philippines. PALAWAN. Photos
   
9. Leptochilus longissimus (Blume) L.Y.Kuo, Taiwania 64 (2019) 378; --Phymatosorus longissimus (Blume) Pic.Serm., Webbia 28 (1973) 459; --Polypodium longissimum Blume, Enum. Pl. Javae 2 (1828) 127. --Microsorum rubidum (J.Sm.) Copel., Gen. Fil. (1947) 197; --Drynaria rubida J.Sm., J. Bot. (Hooker) 3 (1841) 397. Bangladesh, Bismarck Arch, China, Hong Kong, India, Japan, Java, Moluccas, Myanmar, Palau Isls, Peninsular Malaysia, Ryukyu Isls, Singapore, Sulawesi, Sumatra, Taiwan, Thailand, Vietnam, Philippines. Photos
   
10. Leptochilus macrophyllus (Blume) Noot., Blumea 42 (1997) 286; --Colysis macrophylla (Blume) C.Presl, Epimel. Bot. (1851) 147; --Grammitis macrophylla, Enum. Pl. Javae 2 (1828) 119. Bismarck Arch, Borneo, Java, Lesser Sunda Isls, Moluccas, New Guinea, Nicobar Isls, Peninsular Malaysia, Solomon Isls, Sulawesi, Sumatra, Thailand, Philippines. MINDANAO: Bukidnon, PALAWAN. Photos
11. Leptochilus minor Fée, Mém. Foug., 2. Hist. Acrostich. (1845) 87; --Dendroglossa minor (Fée) Copel., Gen. Fil. (1947) 199; --Colysis minor (Fée) M.G.Price, Kalikasan 3 (1974) 176. Borneo, India, Peninsular Malaysia, Sulawesi, Sumatra, Thailand, Vietnam, Philippines. LUZON. Photos
12. Leptochilus pedunculatus (Hook. & Grev.) Fraser-Jenk., Taxon. Revis. Indian Subcontinental Pteridophytes (2008) 62; --Leptochilus macrophyllus (Blume) Noot. var. pedunculatus (Hook. & Grev.) Noot., Blumea 42 (1997) 290; --Ceterach pedunculatum Hook. & Grev., Icon. Filic. (1827) t. 5. --Colysis membranacea (Blume) C.Presl, Epimel. Bot. (1851) 147; --Grammitis membranacea Blume, Enum. Pl. Javae 2 (1828) 118. Bangladesh, China, India, Java, Langkawi Isl, Laos, Myanmar, Nepal, Peninsular Malaysia, Sri Lanka, Sumatra, Thailand, Vietnam, Philippines.
    
13. Leptochilus pothifolius (D.Don) Fraser-Jenk., Taiwania 53 (2008) 181; --Colysis pothifolia (D.Don) C.Presl, Epimel. Bot. (1851) 148; --Hemionitis pothifolia D.Don, Prodr. Fl. Nepal. (1825) 13. Bhutan, Bonin Isl, Cambodia, China, Hong Kong, India, Japan, Laos, Myanmar, Nepal, Ryukyu Isls, ?Taiwan, Thailand, Vietnam, Philippines.
14. Leptochilus pteropus (Blume) Fraser-Jenk., Taxon. Revis. Indian Subcontinental Pteridophytes (2008) 62; --Microsorum pteropus (Blume) Copel., Univ. Calif. Publ. Bot. 16 (1929) 112; --Polypodium pteropus Blume, Enum. Pl. Javae (1828) 125, Addenda 3.
    
    • ssp. pteropus. Bangladesh, Bhutan, Borneo, China, Hong Kong, India, Java, Laos, Lesser Sunda Isls, Moluccas, Myanmar, Nepal, New Guinea, Peninsular Malaysia, Philippines, Ryukyu Isls, Sumatra, Taiwan, Thailand, Vietnam, Philippines.

Loxogramme (Blume) C.Presl

1. Loxogramme antrophyoides (Baker) C.Chr., Mitt. Inst. Hamburg 7 (1928) 163; --Acrostichum antrophyoides Baker, J. Linn. Soc., Bot. 22 (1886) 231. Borneo, Java, Moluccas, Sulawesi, Sumatra, Philippines.
2. Loxogramme avenia (Blume) C.Presl, Tent. Pterid. (1836) 215; --Grammitis avenia Blume, Enum. Pl. Javae 2 (1828) 117. --Loxogramme malayana Copel., PJS C 11 (1916) 46. Cambodia, China, Japan, Java, Lesser Sunda Isls, Micronesia, Moluccas, Myanmar, Peninsular Malaysia, Sulawesi, Sumatra, Thailand, Vietnam, Philippines. MINDANAO: Bukidnon, PALAWAN. Photos
3. Loxogramme chinensis Ching, Sinensia 1 (1929) 13. Bhutan, China, India, Myanmar, Nepal, Taiwan, Thailand, Tibet, Vietnam, Philippines.
4. Loxogramme conferta Copel, Polyp. Philipp. (1905) 68. Endemic to the Philippines. BOHOL, MINDANAO, SAMAR.
5. Loxogramme dimorpha Copel., LPB 2 (1908) 407. Endemic to the Philippines. LUZON, MINDANAO, NEGROS.
6. Loxogramme forbesii Copel, PJS C 9 (1914) 232. Borneo, Sumatra, Philippines.
7. Loxogramme parallela Copel., Fragm. Fl. Philipp. (1905) 182. Borneo, India, Japan, Java, Sri Lanka, Philippines. LUZON, MINDANAO. Photos
8. Loxogramme scolopendrioides (Gaudich.) C.V.Morton, Contr. U.S. Natl. Herb. 38 (1973) 242; --Grammitis scolopendrioides Gaudich., Voy. Uranie (1828) 310. --Loxogramme scolopendrina (Bory) C.Presl, Tent. Pterid. (1836) 215; --Grammitis scolopendrina Bory, Dup. Voy. Bot. 2 (1828) 257. Borneo, Java, Moluccas, Myanmar, New Guinea, Peninsular Malaysia, Solomon Isls, Sulawesi, Sumatra, Thailand, Philippines. Photos
9. Loxogramme subselliguea (Baker) Alston, Journ. of Bot. 77 (1939) 289; --Polypodium subselligueum Baker, J. Bot. 28 (1890) 107. --Loxogramme major Copel., PJS 56 (1935) 106. Bismarck Arch, New Guinea, Sulawesi, Philippines.

Possibly present:

1. Loxogramme remotefrondigera (Hayata) C.Chr., Ind. Fil., Suppl. 2 (1917) 27; --Polypodium remote-frondigerum Hayata, Ic. Pl. Formos. 5 (1915) 1323. Taiwan, ?Philippines.

Micropolypodium Hayata

1. Micropolypodium pulogense (Copel.) A.R.Sm., Novon 2 (1992) 422; --Xiphopteris pulogensis (Copel.) Copel., Gen. Fil. (1947) 215; --Polypodium pulogense Copel., PJS C 6 (1911) 148. Endemic to the Philippines. LUZON.

Microsorum Link

1. Microsorum congregatifolium (Alderw.) Holttum, Rev. Fl. Malaya 2 (1955) 178; --Pleopeltis congregatifolia Alderw., Bull. Jard. Bot. Buitenzorg, sér. 3, 2 (1920) 166. --Microsorum mindanaense (Christ) Copel., Gen. Fil. (1947) 196; --Polypodium mindanaense Christ, Bull. Herb. Boissier, sér. 2, 6 (1906) 994. Borneo, Peninsular Malaysia, Sumatra, Philippines. MINDANAO: Bukidnon. Photos
2. Microsorum heterocarpum (Blume) Ching, Bull. Fan Mem. Inst. Biol. 4 (1933) 295; --Polypodium heterocarpum Blume, Fl. Javae Fil. (1829) 167. Borneo, Java, Lesser Sunda Isls, Moluccas, Peninsular Malaysia, Sulawesi, Sumatra, Thailand, Philippines. LUZON, MINDANAO, PALAWAN. Photos
3. Microsorum heterolobum (C.Chr.) Copel, Gen. Fil. (1947) 196; --Polypodium heterolobum C.Chr., Index Filic. (1906) 532. Endemic to the Philippines. LUZON. Probably a hybrid.
4. Microsorum longissimum J.Sm. ex Fée, Mem. Foug., Gen. Filic. (1850-1852) 268. --Microsorum sablanianum (Christ) Copel., Gen. Fil. (1947) 196; --Polypodium sablanianum Christ, PJS C 2 (1907) 177. Borneo, Philippines. LUZON. Photos
5. Microsorum monstrosum (Copel.) Copel., Gen. Fil. (1947) 196; --Polypodium monstrosum Copel., LPB 1 (1906) 78. Endemic to the Philippines. LUZON. Photos
6. Microsorum pentaphyllum (Baker) Copel., Gen. Fil. (1947) 196; --Polypodium pentaphyllum Baker, Ann. Bot. (Oxford) 5 (1891) 478. --Microsorum curranii (Copel.) Copel., Gen. Fil. (1947) 196; --Polypodium curranii Copel., PJS 4 c (1909) 114. Endemic to the Philippines. Photos
7. Microsorum punctatum (L.) Copel., Univ. Calif. Publ. Bot. 16 (1929) 111; --Acrostichum punctatum L., Sp. Pl., ed. 2. 2 (1763) 1524.  --Microsorum validum (Copel.) Ching, Bull. Fan Mem. Inst. Biol. 4 (1933) 295; --Polypodium validum Copel., Fragm. Fl. Philipp. (1905) 191. Africa, Andaman Isls, Bangladesh, Borneo, Cambodia, China, India, Indian Ocean, Java, Laos, Lesser Sunda Isls, Micronesia, Moluccas, Myanmar, Nepal, New Caledonia, New Guinea, Nicobar Isls, Pacific, Palau Isls, Peninsular Malaysia, Singapore, Solomon Isls, Sri Lanka, Sulawesi, Sumatra, Taiwan, Thailand, Vietnam, Philippines. MINDANAO: Bukidnon, PALAWAN. Photos
8. Microsorum samarense (J.Sm.) Bosman, Leiden Bot. Ser. 14 (1991) 102; --Diblemma samarensis J.Sm., J. Bot. (Hooker) 3 (1841) 399. --Microsorum tenuilore (Kunze ex Mett.) Copel., Univ. Calif. Publ. Bot. 16 (1929) 113; --Polypodium tenuilore Kunze ex Mett., Pol. 86 (1857) n. 150. Endemic to the Philippines. Photos
9. Microsorum sarawakense (Baker) Holttum, Rev. Fl. Malaya 2 (1955) 175; --Polypodium sarawakense Baker, J. Linn. Soc., Bot. 22 (1886) 228. --Microsorum rizalense Copel., PJS 81 (1952) 42. Borneo, Java, Peninsular Malaysia, Sumatra, Thailand, Philippines. Conservation status: Vulnerable (DAO 2017-11). Photos
10. Microsorum zippelii (Blume) Ching, Bull. Fan Mem. Inst. Biol. 4 (1933) 308; --Polypodium zippelii Blume, Fl. Javae Fil. (1829) 172. Bhutan, Borneo, Cambodia, China, India, Java, Laos, Lesser Sunda Isls, Myanmar, New Guinea, Peninsular Malaysia, Sulawesi, Sumatra, Thailand, Vietnam, Philippines. LUZON. Photos

Possibly present:

1. Microsorum musifolium (Blume) Copel., Univ. Calif. Publ. Bot. 16 (1929) 112; --Polypodium musifolium Blume, Enum. Pl. Javae 2 (1828) 134. Borneo, Lesser Sunda Isls, Myanmar, New Guinea, Sumatra, Thailand, ?Philippines.

 Neocheiropteris Christ

Last edited by Julie F. Barcelona, 1 June 2019

No Photos

1. Neocheiropteris ensata (Thunb.) Ching, Bull. Fan Mem. Inst. Biol. 4 (1933) 109; --Microsorum ensatum (Thunb.) H.Ito, J. Jap. Bot. 11 (1935) 96; --Polypodium ensatum Thunb., Trans. Linn. Soc. London 2 (1794) 341. China, India, Japan, South Korea, Taiwan, Vietnam, Philippines. LUZON: Mountain Province.

Oreogrammitis Copel.

1. Oreogrammitis adspersa (Blume) Parris, Gard. Bull. Sing. 58 (2007) 255; --Grammitis adspersa Blume, Fl. Javae Filices (1830) 115. --Polypodium sessilifolium Hook., Sp. Fil. 4 (1863) 168, nom. illeg., non Liebm.; --Type: Cuming 222 (K, lecto; BM, CGE, isolecto), Luzon. --Polypodium paucisorum Copel., PJS 2 c (1907) 137; --Type: Merrill 5964 (MICH, lecto), Mindoro: Mt Halcon. Bismarck Arch, Borneo, China, Fiji, Java, Lesser Sunda Isls, Moluccas, New Guinea, Peninsular Malaysia, Solomon Isls, Sulawesi, Sumatra, Taiwan, Thailand, Vietnam, Philippines. LUZON, MINDORO, PALAWAN.
2. Oreogrammitis beddomeana (Alderw.) T.C.Hsu, Taiwania 64 (2019) 379; --Radiogrammitis beddomeana (Alderw.) Parris, Gard. Bull. Singapore 58 (2007) 241; --Polypodium beddomeanum Alderw., Bull. Jard. Bot. Buitenz. 2, 28 (1918) 39; --Type: from Sri Lanka. --Radiogrammitis alepidota (M.G.Price) Parris, Gard. Bull. Sing. 58 (2007) 241; --Grammitis alepidota M.G.Price, Philip. Agriculturist 57 (1973) 34; --Type: Price 2149 (PNH, holo; K, L, US, iso), Luzon: Mt Banahaw. Sri Lanka, Taiwan, Vietnam, Philippines. LUZON: Ilocos Norte, Cagayan, Quezon, MINDANAO: Bukidnon, PALAWAN. Photos
   
3. Oreogrammitis caespitosa (Blume) Parris, Gard. Bull. Sing. 58 (2007) 257; --Grammitis caespitosa Blume, Enum. Pl. Javae 2 (1828) 115. Borneo, Java, Moluccas, New Guinea, Peninsular Malaysia, Sumatra, Taiwan, Philippines.
4. Oreogrammitis congener (Blume) Parris, Gard. Bull. Sing. 58 (2007) 258; --Grammitis congener Blume, Enum. Pl. Javae 2 (1828) 115. --Grammitis brevipila Copel., Brittonia 1 (1931) 69; --Type:
   Williams 1581 (NY, holo; MICH, UC, iso), Luzon: La Trinidad. --Grammitis duplopilosa Copel., Brittonia 1 (1931) 69; --Type: Williams 1582 (NY, holo; UC, iso), Luzon: Mt Santo Tomas. Borneo, Cambodia, Java, Lesser Sunda Isls, Moluccas, Peninsular Malaysia, Sulawesi, Sumatra, Taiwan, Thailand, Vietnam, Philippines. LUZON: Benguet.
5. Oreogrammitis curtisii (Baker) Parris & Sundue, Phytotaxa 436 (2020) 47; --Themelium curtisii (Baker) Parris, Kew Bull. 59 (2004) 225; --Ctenopteris curtisii (Baker) Copel., PJS 81 (1953) 103; --Polypodium curtisii Baker, J. Bot. (1881) 367; --Type: from Indonesia. --Oreogrammitis blechnifrons (Hayata) T.C.Hsu, Taiwania 64 (2019) 379; --Themelium blechnifrons (Hayata) Parris, Kew Bull. 59 (2004) 224; --Polypodium decrescens var. blechnifrons Hayata, Ic. Pl. Formos. 4 (1914) 245; --Type: from Taiwan. Borneo, Sulawesi, Sumatra, Taiwan, Philippines. Photos
   
6. Oreogrammitis dolichosora (Copel.) Parris, Gard. Bull. Sing. 58 (2007) 258; --Grammitis dolichosora (Copel.) Copel., PJS 80 (1952) 188; --Polypodium dolichosorum Copel., PJS 1(Suppl. 2) (1906) 159; --Type: Copeland 1524 (MICH, lecto; K, NSW, UC, isolecto), Mindanao: Davao. Bismarck Arch, Borneo, Moluccas, New Guinea, Sulawesi, Philippines. MINDANAO: Bukidnon, Davao.
7. Oreogrammitis fasciata (Blume) Parris, Gard. Bull. Sing. 58 (2007) 259; --Grammitis fasciata Blume, Enum. Pl. Javae 2 (1828) 116. Borneo, Java, New Guinea, Sulawesi, Sumatra, Philippines.
8. Oreogrammitis fenicis (Copel.) Parris, Gard. Bull. Sing. 58 (2007) 259; --Grammitis fenicis Copel., Univ. Calif. Publ. Bot. 18 (1942) 224; --Type: Ramos BS 80147 (MICH, holo; K, PNH lost, SING, UC,
   iso), Batanes. Endemic to the Philippines. BATANES.
9. Oreogrammitis halconensis (Baker) Parris & Sundue, Phytotaxa 436 (2020) 48; --Themelium halconense (Baker) Parris, Kew Bull. 59 (2004) 225; --Ctenopteris halconensis (Copel.) Copel., FFP 3 (1960) 533; --Grammitis halconensis (Copel.) Christenh., Global Flora 4 (2018) 50; --Polypodium halconense Copel., PJS 2 c (1907) 138; --Type: Merrill 5976 (MICH, lecto), Mindoro: Mt Halcon. Borneo, Vietnam, Philippines. MINDORO.
   
10. Oreogrammitis jagoriana (Mett. ex Kuhn) Parris & Sundue, Phytotaxa 436 (2020) 49; --Radiogrammitis jagoriana (Mett. ex Kuhn) Parris, Gard. Bull. Sing. 58 (2007) 242; --Grammitis jagoriana (Mett. ex Kuhn) Tagawa, Acta Phytotax. Geobot. 10 (1941) 284; --Polypodium jagorianum Mett. ex Kuhn, Linn. 36 (1869) 128; --Type: Jagor 835 (B, lecto; BM, K, isolecto), Luzon. --Polypodium christii Copel., Polyp. Philipp. (1905) 118; --Type: Copeland 1520 (MICH, lecto; BM, KYO, US, isolecto), Mindanao: Mt Apo. Borneo, Solomon Isls, Sulawesi, Thailand, Vietnam, Philippines. BATANES, MINDANAO.
11. Oreogrammitis knutsfordiana (Baker) Parris, Gard. Bull. Sing. 58 (2007) 262; --Grammitis knutsfordiana (Baker) Copel., Univ. Calif. Publ. Bot. 18 (1942) 224; --Polypodium knutsfordianum Baker, J. Bot. 28 (1890) 107. --Polypodium setosum var. calvum Copel., PJS 2 c (1907) 136; --Type: Merrill 5965 (MICH), Mindoro: Mt Halcon. Bismarck Arch, Borneo, Fiji, Moluccas, New Guinea, Peninsular Malaysia, Solomon Isls, Sulawesi, Sumatra, Philippines. MINDORO.
12. Oreogrammitis locellata (Baker) Parris, Gard. Bull. Sing. 58 (2007) 262; --Grammitis locellata (Baker) Copel., Occ. Pap. Bishop Mus. 15 (1939) 86; --Polypodium locellatum Baker, J. Bot. 28 (1890) 108. --Grammitis stenocrypta Copel., PJS 46 (1931) 220. Borneo, Java, Moluccas, New Guinea, Sulawesi, Sumatra, Philippines.
13. Oreogrammitis loheriana (Christ) Parris & Sundue, Phytotaxa 436 (2020) 49; --Themelium loherianum (Christ) Parris, Kew Bull. 59 (2004) 225; --Grammitis loheriana (Christ) Copel., PJS 80 (1952) 213; --Polypodium loherianum Christ, Bull. Herb. Boiss. 6 (1898) 197; --Type: Loher 1131 (P, lecto; K, isolecto), Luzon: Mountain Province, Mt Data, Mar-1897. Endemic to the Philippines. LUZON: Mountain Province.
    
14. Oreogrammitis marivelesensis (Copel.) Parris, Gard. Bull. Sing. 58 (2007) 263; --Grammitis marivelesensis Copel., Univ. Calif. Publ. Bot. 18 (1942) 224; --Type: Copeland 215 (MICH, holo), Luzon: Mt Mariveles. Taiwan, Philippines. LUZON: Bataan.
15. Oreogrammitis merrillii (Baker) Parris, Gard. Bull. Sing. 58 (2007) 263; --Grammitis merrillii (Copel.) Copel., PJS 80 (1952) 224; Polypodium merrillii Copel., Fragm. Fl. Philipp. (1905) 188; --Type: Merrill 754 (US, lecto; PNH lost, isolecto), Palawan: Iwahig River. Moluccas, New Guinea, Philippines. PALAWAN.
16. Oreogrammitis oncobasis (M.G.Price) Parris, Gard. Bull. Sing. 58 (2007) 264; --Grammitis oncobasis M.G.Price, Contr. Univ. Michigan Herb. 17 (1990) 274; --Type: Price 3376 (MICH, holo), Luzon: Mt Burnay. Endemic to the Philippines. LUZON: Abra.
17. Oreogrammitis padangensis (Baker) Parris, Gard. Bull. Sing. 58 (2007) 264; --Grammitis padangensis (Baker) Copel., PJS 80 (1952) 212; --Polypodium padangense Baker, J. Bot. 18 (1880) 213. --Polypodium durum Copel., LPB 3 (1910) 837; --Type: Elmer 11689 (MICH, lecto), Mindanao: Mt Calelan. Bismarck Arch, Borneo, Java, Lesser Sunda Isls, Moluccas, New Guinea, Sumatra, Philippines. MINDANAO: Davao.
18. Oreogrammitis pilosiuscula (Blume) Parris, Gard. Bull. Singapore 58 (2007) 265; --Grammitis pilosiuscula Blume, Enum. Pl. Javae 2 (1828) 115. Borneo, Java, Sumatra, Philippines.
19. Oreogrammitis reinwardtii (Blume) Parris, Gard. Bull. Sing. 58 (2007) 266; --Grammitis reinwardtii Blume, Enum. Pl. Javae 2 (1828) Add. Australia, Bismarck Arch, Borneo, Java, Lesser Sunda Isls, Moluccas, New Guinea, Peninsular Malaysia, Solomon Isls, Sri Lanka, Sulawesi, Sumatra, Taiwan, Thailand, Vietnam, Philippines. PALAWAN.
20. Oreogrammitis scabristipes (Baker) Parris, Gard. Bull. Sing. 58 (2007) 267; --Grammitis scabristipes (Baker) Copel., Univ. Calif. Publ. Bot. 18 (1942) 223; --Polypodium scabristipes Baker, J. Bot. 28 (1890) 108. --Grammitis bulbotricha (Copel.) Copel., PJS 46 (1931) 219; --Polypodium bulbotrichum Copel., PJS 40 (1929) 309; --Copeland’s Pteridophyta Philippensia Exsiccata 136 (MICH, lecto; BM, BO, K, SING, UC, isolecto), Luzon: Mt Pulog. --Grammitis limapes Copel., 46 (1931) 218. Borneo, Java, Lesser Sunda Isls, Moluccas, New Guinea, Sulawesi, Sumatra, Vanuatu, Philippines. LUZON.
21. Oreogrammitis setigera (Blume) T.C.Hsu, Taiwania 64 (2019) 379; --Radiogrammitis setigera (Blume) Parris, Gard. Bull. Singapore 58 (2007) 244; --Polypodium setigerum Blume, Enum. Pl. Javae 2 (1828) 123. --Grammitis intromissa (Christ) Parris, Fern Gaz. 12 (1981) 180; --Polypodium intromissum Christ, Verh. Naturf. Ges. Basel 11 (1896) 440. --Polypodium heanophyllum Copel., PJS 40 (1929) 310; --Type: Copeland’s Pteridophyta Philippensis Exsiccata 158 (MICH, lecto; BM, K, SING, UC, isolecto), Mindanao: Mt Matutum. Borneo, Java, Lesser Sunda Isls, Moluccas, New Guinea, Solomon Isls, Sulawesi, Sumatra, Taiwan, Philippines. MINDANAO: Bukidnon. Photos
    
22. Oreogrammitis subpinnatifida (Blume) Parris & Sundue, Phytotaxa 436 (2020) 51; --Radiogrammitis subpinnatifida (Blume) Parris, Gard. Bull. Singapore 58 (2007) 245; --Xiphopteris subpinnatifida (Blume) Copel., Gen. Fil. (1947) 215; --Grammitis subpinnatifida Blume, Fl. Javae Fil. (1828) 118; --Type: from Java. Borneo, Java, New Guinea, Peninsular Malaysia, Sulawesi, Sumatra, Philippines. LUZON.
    
23. Oreogrammitis sumatrana (Baker) Parris, Gard. Bull. Sing. 58 (2007) 269; --Grammitis sumatrana (Baker) Copel., PJS 66 (1935) 105; --Polypodium sumatranum Baker, J. Bot. 18 (1880) 214 . --Polypodium pleiosoroides Copel., Fragm. Fl. Philipp. (1905) 189; --Type: Copeland 1011 (PNH, ?lost), Mindanao: Mt Apo. Borneo, Java, Moluccas, New Guinea, Sulawesi, Sumatra, Philippines. MINDANAO.
24. Oreogrammitis taxodioides (Baker) Parris & Sundue, Phytotaxa 436 (2020) 51; --Themelium taxodioides (Baker) Parris, Kew Bull. 52 (1997) 739; --Ctenopteris taxodioides (Baker) Copel., Gen. Fil. (1947) 219; --Polypodium taxodioides Baker, J. Bot. 17 (1879) 42; --Type: from Borneo. --Polypodium pulcherrimum Copel., LPB 3 (1910) 841; --Type: Elmer 11355 (MICH, lecto; BM, BO, E, K, US, isolecto), Mindanao: Davao, Mt Calelan, Aug-1909. Borneo, Moluccas, New Guinea, Solomon Isls, Sulawesi, Philippines. MINDANAO: Davao.
25. Oreogrammitis tenuisecta (Blume) T.C.Hsu, Taiwania 64 (2019) 379; --Themelium tenuisectum (Blume) Parris, Kew Bull. 52 (1997) 740; --Ctenopteris tenuisecta (Blume) J.Sm., Hist. Fil. (1875) 185; --Polypodium tenuisectum Blume, Enum. Pl. Javae 2 (1828) 134. Java, Moluccas, Myanmar, New Guinea, Pacific, Peninsular Malaysia, Solomon Isls, Sumatra, Taiwan, Thailand, Philippines. Photos
26. Oreogrammitis torricelliana (Brause) Parris, Gard. Bull. Sing. 58 (2007) 270; --Grammitis torricelliana (Brause) Parris, Fern Gaz. 12 (1981) 180; --Polypodium torricellianum Brause, Engl. Jahrb. 49 (1912) 45. --Grammitis microtricha Copel., PJS 56 (1935) 105; --Type: Copeland s.n. (MICH, lecto), Mindanao: Mt Apo. New Guinea, Solomon Isls, Philippines. MINDANAO: Bukidnon.
27. Oreogrammitis yoderi (Copel.) Parris & Sundue, Phytotaxa 436 (2020) 51; --Themelium yoderi (Copel.) Parris, Kew Bull. 52 (1997) 739; --Ctenopteris yoderi (Copel.) Copel., Gen. Fil. (1947) 219; --Polypodium yoderi Copel., PJS 1 (Suppl. 2) (1906) 161; --Type: Yoder s.n. (MICH, lecto), Panay: Antique Prov, Mt Madiaas, Apr-1905. Moluccas, New Guinea, Solomon Isls, Philippines. PANAY.

Photinopteris J.Sm. = Drynaria

 Phymatodes C.Presl = Microsorum

 Phymatopsis J.Sm. = Selliguea

 Phymatopteris Pic.Serm. = Selliguea

 Phymatosorus Pic.Serm.

Last edited by Julie F. Barcelona, 7 September 2020

Photos

1. Phymatosorus commutatus (Blume) Pic.-Serm., Webbia 28 (1973) 459; --Microsorum commutatum (Blume) Copel., Gen. Fil. (1947) 196; --Polypodium commutatum Blume, Enum. Pl. Javae 2 (1828). --Microsorum phanerophlebium (Copel.) Copel., Gen. Fil. (1947) 196; --Polypodium phanerophlebium Copel., PJS 1 (Suppl. 2) (1906) 163. Bismarck Arch, Borneo, Java, Lesser Sunda Isls, Moluccas, New Guinea, Pacific, Solomon Isls, Sulawesi, Sumatra, Philippines. LUZON, MINDANAO: Bukidnon, PALAWAN. Photos
2. Phymatosorus membranifolius (R.Br.) S.G.Lu, Guihaia 19 (1999) 27; --Microsorum membranifolium (R.Br.) Ching, Bull. Fan Mem. Inst. Biol. Bot. 10 (1941) 239; --Polypodium membranifolium R.Br., Prodr. Fl. Nov. Holland. (1810) 147. --Polypodium plukenetii C.Presl, Reliq. Haenk. 1 (1825) 21. --Microsorum nigrescens (Blume) Copel., Occ. Pap. Bishop Mus. 14 (1938) 74; --Phymatosorus nigrescens (Blume) Pic.-Serm., Webbia 28 (1973) 459; --Polypodium nigrescens Blume, Enum. Pl. Javae 2 (1828) 126. Andaman Isls, Australia, Bangladesh, Bismarck Arch, Borneo, Cambodia, China, India, Java, Lesser Sunda Isls, Micronesia, Moluccas, New Guinea, Pacific, Peninsular Malaysia, Singapore, Solomon Isls, Sri Lanka, Sulawesi, Sumatra, Taiwan, Thailand, Vietnam, Philippines. PALAWAN. Photos
3. Phymatosorus scolopendria (Burm.f.) Pic.-Serm., Webbia 28 (1973) 460; --Microsorum scolopendria (Burm.f.) Copel., Univ. Calif. Publ. Bot. 16 (1929) 112; --Polypodium scolopendria Burm.f., Fl. Ind. (1768) 232. Tropical Africa to Polynesia and Australia, Philippines. MINDANAO: Bukidnon. Throughout the Philippines. Epiphytic or lithophytic in exposed sites and disturbed forests at low elevations, even on coastal bluffs subjected to salt spray. Common. Photos

Platycerium Desv.

1. Platycerium coronarium (König ex Müller) Desv., Mém. Soc. Linn. Paris 6 (1827) 213; --Coritico & Amoroso, Nature Conservation Research 5 (2020) 80; --Osmunda coronaria König ex Müller, Naturforsch. Halle 21 (1785) 107. Borneo, Cambodia, Laos, Myanmar, Peninsular Malaysia, Singapore, Sumatra, Thailand, Vietnam, Philippines. LUZON, MINDANAO: Bukidnon, Davao Oriental. High canopy epiphyte in lowland primary dipterocarp forests. Also cultivated as an ornamental. Conservation status: Critically Endangered (DAO 2017-11). Photos
2. Platycerium grande (A.Cunn. ex Fée) J.Sm., J. Bot. (Hooker) 3 (1841) 402; --Alcicornium grande (A. Cunn. ex Fée) Underw., Bull. Torrey Bot. Cl. 32 (1905) 594. Endemic to the Philippines. ?MASBATE, MINDANAO: Zamboanga, Lanao, Davao (Mt Apo). Also reported by Copeland (1960) from S LUZON (Mt Cristobal), possibly in error. Majestic crown epiphyte in forest, lowland to 1000m. Also cultivated as an ornamental. Conservation status: Critically Endangered (DAO 2017-11). Photos

 Pleopeltis Humb. & Bonpl. = Lepisorus

Podosorus Holttum [Endemic to the Philippines]

1. Podosorus angustatus Holttum, Kew Bull. 20 (1966) 455; --Type: H. Gutierrez 61-142. A monotypic Philippine endemic genus known only from the type (Gutierrez, H. 61-142), Luzon: Isabela Prov, San Mariano, Brgy Disulap, along creek in Dipterocarp forest. Endemic to the Philippines. LUZON. Only from the type. Conservation status: Critically Endangered (DAO 2017-11).

Prosaptia C.Presl

1. Prosaptia alata (Blume) Christ, Ann. Jard. Bot. Buitenzorg 2, 5 (1905) 127; --Davallia alata Blume, Enum. Pl. Javae 2 (1828) 230. --Prosaptia toppingii Copel., PJS 1 (Suppl. 2) (1906) 158. Borneo, Cambodia, India, Java, Moluccas, New Guinea, Pacific, Peninsular Malaysia, Solomon Isls, Sri Lanka, Sulawesi, Sumatra, Thailand, Vietnam, Philippines.
2. Prosaptia ancestralis Copel., LPB 3 (1910) 385. Endemic to the Philippines. MINDANAO.
3. Prosaptia barathrophylla (Baker) M.G.Price, Contr. Univ. Michigan Herb. 17 (1990) 276; --Polypodium barathrophyllum Baker, J. Bot. (1891) 107. Borneo, Cambodia, China, Peninsular Malaysia, Sulawesi, Thailand, Vietnam, Philippines. PALAWAN. Philippine material with the following two names are referable to this species: Prosaptia khasyana (Hook.) C.Chr. & Tardieu and Ctenopteris khasyana (Hook.) Holttum. 
4. Prosaptia celebica (Blume) Tagawa & K.Iwats., Acta Phytotax. Geobot. 24 (1969) 61; --Ctenopteris celebica (Blume) Copel., Univ. Calif. Publ. Bot. 18 (1942) 225. --Ctenopteris pseudoarticulata (Copel.) Tagawa, Acta Phytotax. Geobot. 16 (1955) 51; --Polypodium pseudoarticulatum Copel., PJS 1 (Suppl. 2) (1906) 160. Borneo, Java, New Guinea, Peninsular Malaysia, Sulawesi, Sumatra, Taiwan, Thailand, Philippines. MINDANAO: Bukidnon. Photos
5. Prosaptia contigua (G.Forst.) C.Presl., Tent. Pterid. (1836) 166; --Trichomanes contiguum G.Forst., Prod. (1786) 84. --Prosaptia pinnatifida C.Presl, Tent. Pterid. (1836) 166. Australia, Bismarck Arch, Borneo, China, India, Java, Lesser Sunda Isls, Moluccas, New Caledonia, New Guinea, Pacific, Peninsular Malaysia, Ryukyu Isls, Solomon Isls, Sri Lanka Thailand, Sulawesi, Sumatra, Taiwan, Philippines. MINDANAO: Bukidnon. Photos
6. Prosaptia cryptocarpa Copel., PJS 1(Suppl. 2) (1906) 158. Endemic to the Philippines. MINDANAO. Photos
7. Prosaptia davalliacea (F.Muell. & Baker) Copel., Univ. Calif. Publ. Bot. 12 (1931) 404; --Polypodium davalliaceum F.Muell. & Baker, J. Bot. (1890) 108. --Prosaptia linearis Copel., PJS 4 c (1909) 115. Borneo, Moluccas, New Guinea, Peninsular Malaysia, Sulawesi, Philippines. LUZON. Photos
8. Prosaptia micropora (Copel.) Parris, Kew Bull. 65 (2010) 123; --Ctenopteris microcarpa Copel., FFP 3 (1960) 531. Endemic to the Philippines. MINDANAO.
9. Prosaptia multicaudata (Copel.) Parris, Kew Bull. 50 (1995) 166; --Ctenopteris multicaudata (Copel.) Copel., PJS 81 (1953) 112; --Polypodium multicaudatum Copel., PJS 1 (Suppl. 2) (1906) 160. Borneo, Philippines. MINDANAO.
10. Prosaptia negrosensis (Copel.) M.G.Price, Contr. Univ. Michigan Herb. 17 (1990) 276; --Ctenopteris negrosensis (Copel.) Copel., FFP 3 (1960) 527; --Polypodium negrosense Copel., LPB 2 (1908) 409. --Ctenopteris spongiosa (Copel.) Copel., FFP 3 1960 527; --Polypodium spongiosum Copel., PJS 40 (1929) 308. Endemic to the Philippines. NEGROS.
11. Prosaptia nutans (Blume) Mett., Reise Novara 1 (1870) 214; --Ctenopteris nutans (Blume) J.Sm., Hist. Fil. (1875) 185; --Polypodium nutans Blume, Enum. Fil. (1828) 128. Borneo, Java, Moluccas, New Guinea, Peninsular Malaysia, Sulawesi, Sumatra, Taiwan, Philippines.
12. Prosaptia obliquata (Blume) Mett., Novara Reise Bot. Exped. 1 (1870) 214; --Ctenopteris obliquata (Blume) Copel., PJS 81 (1952) 111; --Polypodium obliquatum Blume, Enum. Pl. Javae 2 (1828) 128. Borneo, China, India, Java, Lesser Sunda Isls, Moluccas, New Guinea, Peninsular Malaysia, Sri Lanka, Sulawesi, Sumatra, Taiwan, Thailand, Vietnam, Philippines. MINDANAO: Bukidnon, PALAWAN. Photos
13. Prosaptia pectinata T.Moore, Index Fil. (1861) 298. --Prosaptia leysii (Baker) Ching, Bull. Fan Mem. Inst. Biol. Bot. 10 (1941) 242; --Polypodium leysii Baker, J. Bot. (1879) 66. Borneo, Java, Moluccas, Peninsular Malaysia, Sulawesi, Sumatra, Taiwan, Thailand, Philippines.
14. Prosaptia urceolaris (Hayata) Copel., PJS 40 (1929) 311; --Polypodium urceolare Hayata, Ic. Pl. Formos. 6 (1916) 324. Taiwan, Philippines.
15. Prosaptia venulosa (Blume) M.G.Price, Contr. Univ. Michigan Herb. 16 (1987) 197; --Ctenopteris venulosa (Blume) Kunze, Bot. Zeit. (Berlin) (1846) 425; --Polypodium venulosum Blume, Enum. Fil. (1828) 128. Borneo, Java, Lesser Sunda Isls, Moluccas, New Guinea, Sulawesi, Philippines. Photos

Pteropsis Desv. = Pyrrosia

Pyrrosia Mirbel

1. Pyrrosia lanceolata (L.) Farw., Amer. Midl. Naturalist 12 (1931) 245; --Acrostichum lanceolatum L., Sp. Pl. 2 (1753) 1067. --Pyrrosia varia (Kaulf.) Farw., Amer. Midl. Naturalist 12 (1931) 302. Africa, Andaman Isls, Australia, Bangladesh, Bangladesh, Bhutan, Borneo, Cambodia, China, India, Indian Ocean, Java, Laos, Lesser Sunda Isls, Moluccas, Myanmar, Nepal, New Guinea, Nicobar Isls, Pacific, Palau Isls, Peninsular Malaysia, Ryukyu Isls, Singapore, Solomon Isls, Sri Lanka, Sulawesi, Sumatra, Taiwan, Thailand, Tibet, Vietnam, Philippines. PALAWAN. Epiphytic or lithophytic, usually at low elevation. Photos
2. Pyrrosia longifolia (Burm.f.) C.V.Morton, J. Wash. Acad. Sci. 36 (1946) 168; --Acrostichum longifolium Burm.f., Fl. Indica (1768) 228. --Pyrrosia acrostichoides (G.Forst.) Ching, Bull. Chin. Bot. Soc. 1 (1935) 69; --Polypodium acrostichoides G.Forst., Prod. (1786) 81. Australia, Bismarck Arch, Borneo, Cambodia, China, Java, Laos, Lesser Sunda Isls, Moluccas, Myanmar, New Caledonia, New Guinea, Peninsular Malaysia, Singapore, Solomon Isls, Sulawesi, Sumatra, Tahiti, Thailand, Vietnam, Philippines. PALAWAN. Photos
3. Pyrrosia nummularifolia (Sw.) Ching, Bull. Chin. Bot. Soc. 1 (1935) 47; --Acrostichum nummularifolium Sw., Syn. Fil. 191 (1806) 419. Bangladesh, Bhutan, Borneo, ?Cambodia, China, India, ?Laos, Lesser Sunda Isls, Myanmar, Peninsular Malaysia, Sulawesi, Thailand, Vietnam, Philippines. PALAWAN. Photos
4. Pyrrosia piloselloides (L.) M.G.Price, Kalikasan 3 (1974) 176; --Drymoglossum piloselloides (L.) C.Presl, Tent. Pterid. (1836) 227; --Pteropsis piloselloides (L.) Desv., Mém. Soc. Linn. Paris 6 (1827) 218; --Pteris piloselloides L., Sp. Pl. (ed. 2) 2 (1763) 1530. Andaman Isls, Bangladesh, Bismarck Arch, Borneo, ?Cambodia, China, India, Japan, Java, Laos, Lesser Sunda Isls, Moluccas, New Guinea, Peninsular Malaysia, Singapore, Sulawesi, Sumatra, Thailand, Vietnam, Philippines. Throughout the Philippines. Epiphytic on tree trunks and branches or epilithic on outcrops and boulders along rivers and streams at low elevation. Common. Photos
5. Pyrrosia porosa (C.Presl) Hovenkamp, Blumea 30 (1984) 208. Bhutan, China, India, Japan, Laos, Myanmar, Nepal, Sri Lanka, Taiwan, Thailand, Tibet, Vietnam, Philippines. LUZON. Photos
6. Pyrrosia rasamalae (Racib.) K.H.Shing, Amer. Fern J. 73 (1983) 78; --Polypodium rasamalae Racib., Pterid. Buit. (1899) 99. --Pyrrosia lanuginosa (Giesenh.) Copel., FFP 3 (1960) 472; --Niphobolus lanuginosus Giesenh., Niph. (1901) 190. Borneo, India, Java, Laos, Lesser Sunda Isls, Myanmar, Peninsular Malaysia, Sumatra, Thailand, Philippines. LUZON. Photos
7. Pyrrosia samarensis (C.Presl) Ching, Bull. Chin. Bot. Soc. 1 (1935) 49; --Gyrosorium samarense C.Presl., Epim. Bot. (1851) 140. --Pyrrosia philippinensis Copel., PJS 81 (1952) 43, nom. superfl. Endemic to the Philippines. Photos
8. Pyrrosia sphaerosticha (Mett.) Ching, Bull. Chin. Bot. Soc. 1 (1935) 62; --Polypodium sphaerostichum Mett., Abh. Senckenberg. Naturf. Ges. (Polypodium) (1857) 130. Lesser Sunda Isls, Moluccas, Sulawesi, Philippines. MINDANAO: Bukidnon. Photos
9. Pyrrosia splendens (C.Presl) Ching, Bull. Chin. Bot. Soc. 1 (1935) 68. Endemic to the Philippines. BOHOL, LEYTE, LUZON, MINDANAO, SAMAR. Conservation status: Vulnerable (DAO 2017-11). Photos

Radiogrammitis Parris = Oreogrammitis

 Schellolepis Pic.-Serm. = Goniophlebium

Scleroglossum Alderw.

1. Scleroglossum minus (Fée) C.Chr., Gard. Bull. Straits Settlem. 4 (1929) 29; --Vittaria minor Fée, Mém. Foug. 3 (1851) 23. Borneo, Moluccas, New Guinea, Peninsular Malaysia, Solomon Isls, Sulawesi, Sumatra, Philippines. PALAWAN.
   
2. Scleroglossum pusillum (Blume) Alderw., Bull. Jard. Bot. Buitenzorg Ser. 2, 7 (1912) 39; --Vittaria pusilla Blume, Enum. Fil. (1828) 199. Bismarck Arch, Borneo, Java, Lesser Sunda Isls, Moluccas, New Guinea, Pacific, Peninsular Malaysia, Solomon Isls, ?Sri Lanka, Sulawesi, Sumatra, Thailand, Vietnam, Philippines. MINDANAO: Bukidnon, NEGROS, PALAWAN. Photos
3. Scleroglossum sulcatum (Mett. ex Kuhn) Alderw., Bull. Jard. Bot. Buitenzorg Ser. 2, 7 (1912) 39; --Vittaria sulcata, Mett. ex Kuhn, Linn. 36 (1869) 68. --Scleroglossum intermedium (Copel.) Copel., PJS 56 (1935) 480; --Monogramma intermedia Copel., PJS 1, Suppl. 4 (1906) 255. Borneo, China, India, ?Java, Moluccas, New Caledonia, New Guinea, Pacific, Peninsular Malaysia, Solomon Isls, Sri Lanka, Sulawesi, Sumatra, Taiwan, Thailand, Vietnam, Philippines.

Selliguea Bory

1. Selliguea albidosquamata (Blume) Parris, Pl. Mt. Kinabalu (1992) 152; --Crypsinus albidosquamatus (Blume) Copel., Gen. Fil. (1947) 207; --Polypodium albidosquamatum Blume, Enum. Pl. Javae 2 (1828) 132. Borneo, Java, Lesser Sunda Isls, Moluccas, New Guinea, Peninsular Malaysia, Solomon Isls, Sulawesi, Sumatra, Philippines. MINDANAO: Bukidnon. Photos
2. Selliguea caudiformis (Blume) J.Sm., Ferns Brit. For. (1866) 97; --Polypodium caudiforme Blume, Enum. Pl. Javae (1829) 122. --Selliguea calophlebia (Copel.) Copel., FFP 3 (1960) 508; --Polypodium calophlebium Copel., PJS 2 c (1907) 140. Moluccas, Sulawesi, Philippines. PALAWAN.
3. Selliguea elmeri (Copel.) Ching, Sunyatsenia 5 (1940) 260; --Selliguea feei Bory var. elmeri (Copel.) Tagawa, (1956); --Polypodium elmeri Copel., Fragm. Fl. Philip. (1905) 191. Endemic to the Philippines. LUZON, MINDANAO: Bukidnon. Photos
4. Selliguea enervis (Cav.) Ching, Bull. Fan Mem. Inst. Biol. Bot. 10 (1941) 239; --Crypsinus enervis (Cav.) Copel., Gen. Fil. (1947) 207; --Polypodium enerve Cav., Descr. Pl. (1802) 245. Bismarck Arch, Borneo, Java, Lesser Sunda Isls, Moluccas, New Guinea, Peninsular Malaysia, Solomon Isls, Sulawesi, Sumatra, Thailand, Vietnam, Philippines. PALAWAN. Photos
5. Selliguea glauca (J.Sm. ex Brack.) Hovenkamp, Blumea 43 (1998) 56; --Crypsinus glaucus (J.Sm. ex Brack.) Copel., Gen. Fil. (1947) 206; --Drynaria glauca J. Sm. ex Brack., U.S. Expl. Exp. 16 (1854) 54. Endemic to the Philippines. Photos
6. Selliguea hastata (Thunb.) Fraser-Jenk., Taxon. Revis. Indian Subcontinental Pteridophytes (2008) 44; --Crypsinus hastatus (Thunb.) Copel., Gen. Fil. (1947) 206;  --Phymatopteris hastata (Thunb.) Pic.-Serm., Webbia 28 (1973) 462; --Polypodium hastatum Thunb., Linn. Syst. Veg. ed. 14 (1784) 935. China, Japan, North Korea, Siberia, South Korea, Taiwan, Tibet, Philippines. LUZON: Benguet. Epiphytic or lithophytic, c. 2400m. Photos
   
7. Selliguea heterocarpa (Blume) Blume, Enum. Pl. Javae (1828) 125; --Grammitis heterocarpa Blume, Enum. Pl. Jav. 2 (1829) 11. Borneo, Java, Lesser Sunda Isls, Moluccas, New Guinea, Peninsular Malaysia, Singapore, Sulawesi, Sumatra, Thailand, Philippines.
8. Selliguea lagunensis (Christ) Hovenkamp, Blumea 43 (1998) 55; --Crypsinus lagunensis (Christ) Copel., Gen. Fil. (1947) 206; --Phymatopteris lagunensis (Christ) Pic.-Serm., Webbia 28 (1973) 463; --Polypodium lagunense Christ, Bull. Herb. Boiss. 6 (1898) 201. Endemic to the Philippines. LUZON. Photos
9. Selliguea moulmeinensis (Bedd.) X.C.Zhang & L.J.He, Taxon 67 (2018) 244; --Pleopeltis moulmeinensis Bedd., Handb. Ferns Brit. India (1867) t.205. --Arthromeris proteus (Copel.) Tagawa, J. Jap. Bot. 27 (1952) 218; --Polypodium proteus Copel., PJS 1 (Suppl. 2) (1906) 164. --Arthromeris mairei (Brause) Ching, Sunyatsenia 6 (1941) 5; --Polypodium mairei Brause, Hedwigia 64 (1914) 208. ?Bhutan, China, India, Myanmar, Nepal, Tibet, Philippines. LUZON. Conservation status: Other Threatened Species (DAO 2017-11 as Arthromeris mairei).
   
10. Selliguea oodes (Kunze) Hovenkamp, Blumea 43 (1998) 89; --Crypsinus oodes (Kunze) Copel., Gen. Fil. (1947) 207; --Polypodium oodes Kunze, Bot. Zeit. (Berlin) (1846) 421. Borneo, Sulawesi, Philippines. LUZON. Photos
11. Selliguea pyrolifolia (Goldm.) Hovenkamp, Blumea 43 (1998) 58; --Crypsinus pyrolifolius (Goldm.) Copel., Gen. Fil. (1947) 206; --Polypodium pyrolifolium Goldm., Nova Acta 19, Suppl. 1 (1843) 453. Endemic to the Philippines. MINDANAO: Bukidnon. Photos
12. Selliguea stenophylla (Blume) Parris, Pl. Mt. Kinabalu (1992) 151; --Crypsinus stenophyllus (Blume) Copel., Gen. Fil. (1947) 206; --Polypodium stenophyllum Blume, Enum. Fil. (1828) 124. Borneo, Java, Lesser Sunda Isls, Moluccas, New Guinea, Peninsular Malaysia, Sulawesi, Sumatra, Thailand, Philippines. Photos
13. Selliguea taeniata (Sw.) Parris, Pl. Mt. Kinabalu (1992) 152; --Crypsinus taeniatus (Sw.) Copel., Gen. Fil. (1947) 206; --Phymatopteris taeniata (Sw.) Pic.-Serm., Webbia 28 (1973) 465; --Polypodium taeniatum Sw., Schrad., J. Bot. 1800 (1801) 26. --Crypsinus ramosii Copel., PJS 81 (1952) 43. Borneo, Brunei, Java, Lesser Sunda Isls, Moluccas, New Guinea, Peninsular Malaysia, Sulawesi, Sumatra, Philippines. MINDANAO: Bukidnon, PALAWAN. Photos
14. Selliguea taeniophylla (Copel.) Parris, Pl. Mt. Kinabalu (1992) 152; --Polypodium taeniophyllum Copel., PJS 7 (1912) 65. --Selliguea lateritia (Baker) Hovenkamp, Blumea 43 (1998) 71; --Polypodium lateritium Baker, Ann. Bot. 5 (1891) 476. Borneo, Cambodia, New Guinea, Peninsular Malaysia, Sulawesi, Sumatra, Thailand, Vietnam, Philippines.
15. Selliguea triloba (Houtt.) M.G.Price, Blumea 43 (1998) 57; --Crypsinus trilobus (Houtt.) Copel., Gen. Fil. (1947) 206; --Phymatopteris triloba (Houtt.) Pic.-Serm., Webbia 28 (1973) 465; --Polypodium trilobum Houtt., Natuurl. Hist. 14 (1783) 148. --Selliguea matutumensis Copel., PJS 81 (1952) 44. Borneo, Cambodia, China, Java, Laos, Moluccas, Peninsular Malaysia, Sumatra, Thailand, Vietnam, Philippines. MINDANAO: Bukidnon.
16. Selliguea whitfordii (Copel.) Hovenkamp, Blumea 43 (1998) 60; --Crypsinus whitfordii (Copel.) Copel., Gen. Fil. (1947) 206; --Polypodium whitfordii Copel., PJS 1 (Suppl. 4) (1906) 256. --Crypsinus subdimorphus Copel., PJS 81 (1952) 43. Endemic to the Philippines. Photos

Possibly present:

1. Selliguea feei Bory, Dict. Class d'Hist. Nat. 6 (1824) 588. China, Java, Lesser Sunda Isls, Moluccas, Sulawesi, Sumatra, ?Philippines.

Themelium (T.Moore) Parris = Oreogrammitis

Thylacopteris Kunze ex J.Sm.

1. Thylacopteris papillosa (Blume) Kunze ex J.Sm., Hist. Fil. (1875) 88; --Polypodium papillosum Blume, Enum. Pl. Javae 2 (1828) 131. Borneo, India, Java, Lesser Sunda Isls, Moluccas, Peninsular Malaysia, Sulawesi, Sumatra, Philippines. PALAWAN. Photos

Tomophyllum (E.Fourn.) Parris

1. Tomophyllum macrum (Copel.) Parris, Gard. Bull. Sing. 58 (2007) 247; --Ctenopteris macra (Copel.) Copel., FFP 3 (1960) 529; --Polypodium macrum Copel., Fragm. Fl. Philip. (1905) 189; --Type: Copeland 1016 (MICH, lecto), Mindanao: Mt Apo. Endemic to the Philippines. MINDANAO, PALAWAN.
2. Tomophyllum millefolium (Blume) Parris, Gard. Bull. Sing. 58 (2007) 247; --Ctenopteris millefolia (Blume) Copel., Gen. Fil. (1947) 219; --Polypodium millefolium Blume, Enum. Fil. (1828) 134. Borneo, Java, Moluccas, New Guinea, Sulawesi, Sumatra, Philippines.
3. Tomophyllum minutum (Blume) Parris, Gard. Bull. Sing. 58 (2007) 247; --Ctenopteris minuta (Blume) Holttum, Rev. Fl. Mal. 2 (1955) 228; --Polypodium minutum Blume, Enum. Fil. (1828) 130. Borneo, Java, Philippines. LUZON, MINDANAO: Bukidnon.
4. Tomophyllum repandulum (Kunze ex Mett.) Parris, Gard. Bull. Singapore 58 (2007) 248; --Polypodium repandulum Kunze ex Mett., Abh. Senckenberg. Naturf. Ges. (Polypodium) 50 (1857) n.54. Borneo, Moluccas, New Guinea, Solomon Isls, Sri Lanka, Sulawesi, Vietnam, Philippines.
5. Tomophyllum subsecundodissectum (Zoll.) Parris, Gard. Bull. Sing. 58 (2007) 249; --Ctenopteris subsecundodissecta (Zoll.) Copel., PJS 81 (1953) 114; --Polypodium subsecundodissectum Zoll., Syst. Verz. 38 (1854) 48; --Type: Merrill 5971 (MICH, lecto), Mindoro: Mt Halcon. Bismarck Arch, Borneo, Java, Moluccas, New Caledonia, New Guinea, Philippines. MINDANAO: Bukidnon, MINDORO. Photos

Possibly present:

1. Tomophyllum inconspicuum (Blume) Parris, Gard. Bull. Singapore 58 (2007) 247; --Polypodium inconspicuum Blume, Enum. Fil. (1828) 130. Borneo, Java, Palau Isls, Peninsular Malaysia, ?Philippines.

Xiphopterella Parris

1. Xiphopterella nudicarpa (P.M.Zamora & Co) Parris, Fern Gaz. 20 (2015) 47; --Acrosorus nudicarpus P.M.Zamora & Co, Nat. Appl. Sc. Bull. Univ. Philipp. 32 (1980) 47; --Xiphopteris nudicarpa (P.M.Zamora & Co) Parris in Parris et al., Pl. Mt Kinabalu 1 (1992) 77. --Grammitis palasaba M.G.Price, Contr. Univ. Mich. Herb. 16 (1987) 196. Borneo, Philippines. PALAWAN. Conservation status: Vulnerable (DAO 2017-11).

Psilotaceae

Psilotum L.

1. Psilotum complanatum Sw., Syn. Fil. (1806) 188. Americas, Australia, Bismarck Arch, Indian Ocean, Moluccas, New Guinea, Nicobar Isls, Pacific, Palau Isls, Peninsular Malaysia, Singapore, Solomon Isls, Sulawesi, Sumatra, Thailand, Vietnam, Philippines. MINDANAO: Bukidnon, Davao Oriental, Misamis Occidental. Conservation status: Endangered (DAO 2017-11). Photos
2. Psilotum nudum (L.) Griseb., Prodr. Aethéogam. (1805) 112; --Coritico & Amoroso, Nature Conservation Research 5 (2020) 82; --Lycopodium nudum L., Sp. Pl. 2 (1753) 1100. --Psilotum triquetrum Sw., Schrad. J. Bot. 1800 (2) 1801. Africa, Americas, Andaman Isls, Australia, Bangladesh, China, India, Japan, Laos, Lesser Sunda Isls, Maldives, Middle East, Moluccas, Myanmar, Nepal, New Caledonia, New Guinea, Nicobar Isls, Pacific, Pakistan, Palau Isls, Peninsular Malaysia, Singapore, Solomon Isls, South Korea, Sri Lanka, Sulawesi, Taiwan, Thailand, Vietnam, Philippines. MINDANAO: Bukidnon, Davao Oriental, North Cotabato. Conservation status: Vulnerable (DAO 2017-11). Photos

Tmesipteris Bernh.

1. Tmesipteris zamorarum Gruezo & Amoroso, Asia Life Sci. 21 (2012) 656 (as "zamorae"); --Coritico & Amoroso, Nature Conservation Research 5 (2020) 82; --Type: WS Gruezo WM 14493 (CAPH, holo; WGRZ, iso), Luzon: Quezon, Mt. Banahaw, 19-Mar-1982, 1200m. --Tmesipteris lanceolata non Dang, sensu Bierhorst, J. Linn. Soc. Bot. 67, Suppl. 1 (1973) 46. --Tmesipteris tannensis auctt. non (Spreng.) Bernh. Endemic to the Philippines. LUZON: Benguet (Mt Pulog), Mountain Province, Quezon, MINDANAO: Bukidnon, Davao, Misamis Occidental, North Cotabato, South Cotabato, MINDORO: Mt Halcon. 1000-2900m. Conservation status: Endangered (DAO 2017-11). Photos

Pteridaceae 

Including Acrostichaceae, Adiantaceae, Antrophyaceae, Ceratopteridaceae, Cheilanthaceae, Hemionitidaceae, Parkeriaceae, Sinopteridaceae, Taenitidaceae, Vittariaceae

Acrostichum L.

1. Acrostichum aureum L., Sp. Pl. 2 (1753) 1069. --Parahemionitis arifolia (Burm.f.) Panigr., Amer. Fern J. 83 (1993) 90; --Hemionitis arifolia (Burm. f.) T.Moore, Ind. Fil. (1859) 114. Africa, Americas, Andaman Isls, Bangladesh, Bismarck Arch, Cambodia, China, Hong Kong, India, Indian Ocean, Lesser Sunda Isls, Myanmar, New Caledonia, New Guinea, Nicobar Isls, Pacific, Palau Isls, Peninsular Malaysia, Ryukyu Isls, Singapore, Solomon Isls, Sri Lanka, Sulawesi, Taiwan, Thailand, Vietnam, Philippines. PALAWAN. Photos
2. Acrostichum speciosum Willd., Sp. Pl., ed. 4 5 (1810) 117. Andaman Isls, Australia, Bismarck Arch, China, Indian Ocean., Lesser Sunda Isls, Moluccas, New Caledonia, New Guinea, Nicobar Isls, Pacific, Palau Isls, Peninsular Malaysia, Singapore, Solomon Isls, ?Sri Lanka, Sulawesi, Thailand, Vietnam, Philippines. PALAWAN. Photos

Adiantum L.

1. Adiantum capillus-veneris L., Sp. 2 (1753) 1096. Very widespread, including in the Philippines. Photos
2. Adiantum caudatum L., Mant. Pl. (1771) 308. Bangladesh, Bhutan, Cambodia, China, India, Laos, Lesser Sunda Isls, Nepal, New Caledonia, New Guinea, Peninsular Malaysia, Sri Lanka, Sulawesi, Taiwan, Thailand, Vanuatu, Vietnam, Philippines. PALAWAN. Photos
3. Adiantum ciliatum Blume, Enum. Pl. Javae 2 (1828) 215. --Adiantum malesianum Ghatak, Bull. Bot. Surv. India 5 (1963) 73. Borneo, China, Moluccas, Myanmar, Peninsular Malaysia, Sulawesi, Sumatra, Taiwan, Vietnam, Philippines. PALAWAN. Photos
   
4. Adiantum cupreum Copel., LPB 4 (1911) 1152. Endemic to the Philippines. SIBUYAN. Conservation status: Vulnerable (DAO 2017-11).
5. Adiantum diaphanum Blume, Enum. Pl. Javae 2 (1828) 215. --Adiantum setulosum J.Sm., Companion Bot. Mag. 72 (1846) 22. Australia, Borneo, China, Japan, Lesser Sunda Isls, Moluccas, New Caledonia, New Guinea, Pacific, Ryukyu Isls, Solomon Isls, Sulawesi, Taiwan, Vietnam, Philippines. Photos
   
6. Adiantum edgeworthii Hook., Sp. Fil. 2 (1851) 14. Bhutan, China, India, Japan, Lesser Sunda Isls, Myanmar, Nepal, peninsular Malaysia, Taiwan, Thailand, Tibet, Vietnam, Philippines.
7. Adiantum flabellulatum L., Sp. 2 (1753) 1095. Bangladesh, Cambodia, China, India, Japan, Laos, Lesser Sunda Isls, Myanmar, Peninsular Malaysia, Ryukyu Isls, Singapore, Sri Lanka, Taiwan, Thailand, Vietnam, Yakushima Isl, Philippines.
8. Adiantum hispidulum Sw., J. Bot. (Schrader) 1800 (1801) 82. Africa, Australia, China, India, Indian Ocean, Java, Lesser Sunda Isls, New Caledonia, Pacific, Palau Isls, Peninsular Malaysia, Singapore, Sri Lanka, Sulawesi, Taiwan, Philippines. Photos  
9. Adiantum hosei Baker, J. Bot. 26 (1888) 324; --Coritico et al., PJS 149 (2020) 782. Borneo, Sulawesi, Philippines. MINDANAO: Bukidnon.
   
10. Adiantum mindanaoense Copel., PJS 1 (Suppl. 2) (1906) 154. Moluccas, New Guinea, Sulawesi, Philippines. MINDANAO. Conservation status: Endangered (DAO 2017-11). Photos
11. Adiantum monosorum Baker, Syn. Fil. (Hooker & Baker), ed. 2 (1874) 472. --Adiantum scabripes Copel., PJS, C 7 (1912) 55. Moluccas, Solomon Isls, Sulawesi, Philippines. MINDANAO. Conservation status: Vulnerable (DAO 2017-11). Photos 
    
12. Adiantum philippense L., Sp. Pl. 2 (1753) 1094. Photos
    
    • ssp. philippense. Africa, Andaman Isls, Australia, Bangladesh, Bhutan, Cambodia, China, India, Indian Ocean, Laos, Lesser Sunda Isls, Moluccas, Myanmar, Nepal, Pacific, Palau Isls, Peninsular Malaysia, ?Solomon Isls, Sri Lanka, Sulawesi, Taiwan, Thailand, Vietnam, Philippines. PALAWAN.
    • ssp. intermedium, S.C.Verma & Fraser-Jenk., Taxon. Revis. Indian Subcontinental Pteridophytes (2008). Bhutan, Bismarck Arch, China, Fiji, Guam, India, Myanmar, Nepal, New Guinea, Solomon Isls, Sulawesi, Thailand, Vietnam, Philippines.
13. Adiantum soboliferum Wall. ex Hook., Sp. Fil. 2 (1851) 13. Africa, China, Taiwan, Myanmar, Thailand, Laos, ?Cambodia, India, Vietnam, Peninsular Malaysia, Lesser Sunda Isls, Madagascar, Philippines. LUZON, PALAWAN. Photos
14. Adiantum stenochlamys Baker, Ann. Bot. (Oxford) 5 (1891) 209. --Adiantum opacum Copel., PJS 1 (Suppl. 4) (1906) 255. Andaman Isls, Borneo, Borneo, Cambodia, Laos, Peninsular Malaysia, Singapore, Sulawesi, Sumatra, Thailand, Vietnam, Philippines. PALAWAN. Conservation status: Vulnerable (DAO 2017-11). Photos
15. Adiantum tenerum Sw., Prodr. (1788) 135. Introduced from South America, not native to the Philippines. Photos

Aleuritopteris Fée

1. Aleuritopteris albomarginata (C.B.Clarke) Ching, Hong Kong Naturalist 10 (1941) 199; --Cheilanthes albomarginata C.B.Clarke, Trans. Linn. Soc. London, Bot. 1 (1880) 456. Bhutan, China, India, Myanmar, Nepal, Pakistan, Thailand, Tibet, Vietnam, Philippines. Photos
2. Aleuritopteris dubia (Hope) Ching, Hongkong Nat. 10 (1941) 200; --Cheilanthes dubia Hope, J. Bombay Nat. Hist. Soc. 12 (1899) 528. --Aleuritopteris rufa auct. non (D.Don) Ching sensu Copel. Photos
   
3. Aleuritopteris formosana (Hayata) Tagawa, Acta Phytotax. Geobot. 14 (1952) 191; --Cheilanthes formosana Hayata, Enum. Pl. Formosa (1906) 61. Bhutan, China, India, Myanmar, Nepal, Pakistan, Taiwan, Thailand, Tibet, Philippines.
4. Aleuritopteris krameri (Franch. & Sav.) Ching, Hongkong Nat. 10 (1941) 202; --Cheilanthes krameri Franch. & Sav., Enum. Pl. Jap. 2 (1879) 212. --Aleuritopteris farinosa auct. non (Forssk.) Fée. Japan, Thailand, Taiwan, Philippines.

Antrophyum Kaulf.

1. Antrophyum callifolium Blume, Enum. Pl. Javae 2 (1828) 111. --Antrophyum nanum Fée, Mém. Foug. 4 (1851) 44. --Antrophyum alatum Brack., U.S. Expl. Exped., Filic. 16 (1854) 64. Andaman Isls, Australia, Bismarck Arch, Borneo, Cambodia, China, Laos, Lesser Sunda Isls, Moluccas, New Caledonia, New Guinea, Nicobar Isls, Pacific, Palau Isls, Peninsular Malaysia, Singapore, Solomon Isls, Sulawesi, Sumatra, Taiwan, Thailand, Vietnam, Philippines. PALAWAN.
   
2. Antrophyum latifolium Blume, Fl. Javae 2 (1829) 75. Borneo, Java, Moluccas, New Guinea, Sulawesi, Sumatra, Philippines. Photos  
3. Antrophyum ledermannii Hieron., Bot. Jahrb. Syst. 56 (1920) 175. Moluccas, New Guinea, Philippines.
4. Antrophyum parvulum Blume, Pl. Jav. (1828) 110. --Antrophyum immersum (Bory ex Willd.) Mett., Ann. Mus. Bot. Lugd.-Bat. 4 (1868) 171; --Hemionitis immersa Bory ex Willd., Sp. 5 (1810) 127. Africa, Borneo, ?Cambodia, China, India, Indian Ocean, Japan, Java, ?Laos, Lesser Sunda Isls, Nicobar Isls, Peninsular Malaysia, Singapore, Sulawesi, Sumatra, Taiwan, Thailand, Vietnam, Philippines. PALAWAN. Photos
5. Antrophyum plantagineum (Cav.) Kaulf., Enum. Filic. (1824) 197; --Hemionitis plantaginea Cav., Descr. Pl. (1802) 260. Australia, Bangladesh, Bismarck Arch, India, Java, Lesser Sunda Isls, Moluccas, New Caledonia, New Guinea, Pacific, Palau Isls, Peninsular Malaysia, Solomon Isls), Sri Lanka, Sulawesi, Sumatra, Philippines. MINDANAO: Bukidnon. Photos
6. Antrophyum reticulatum (G.Forst.) Kaulf., Enum. Filic. (1824) 198; --Hemionitis reticulata G.Forst., Fl. Ins. Austr. (1786) 79.  --Antrophyum clementis Christ, PJS 2 c (1907) 175. Andaman Isls, Bhutan, Borneo, China, India, Laos, Lesser Sunda Isls, Moluccas, Myanmar, Nepal, Nicobar Isls, Pacific, Peninsular Malaysia, Sri Lanka, Sulawesi, Tibet, Vietnam, Philippines. PALAWAN. Photos
7. Antrophyum semicostatum Blume, Enum. Pl. Javae 2 (1828) 110. Borneo, Fiji, Lesser Sunda Isls, Moluccas, Peninsular Malaysia, Solomon Isls, Sulawesi, Sumatra, Philippines. Photos
8. Antrophyum sessilifolium (Cav.) Spreng., Syst. Veg. 4 (1827) 67; --Hemionitis sessilifolia Cav., Descr. Pl. (1802) 261. Borneo, Sulawesi, Taiwan, Philippines. Photos
9. Antrophyum williamsii Benedict, Amer. Fern J. 1 (1911) 72. Endemic to the Philippines. LUZON. Photos

Austrogramme E.Fourn.

1. Austrogramme luzonica (Alderw.) M.Kato,  J. Fac. Sci. Univ. Tokyo, Bot. 14 (1988) 174; --Taenitis luzonica (Alderw.) Holttum, Kew Bull. 30 (1975) 335; --Syngramma luzonica Alderw., Bull. Jard. Bot. Buitenzorg Ser. 2I, 2 (1920) 174. Endemic to the Philippines. Photos

Calciphilopteris Yesilyurt & H.Schneid.

1. Calciphilopteris wallichii (J.Sm.) Yesilyurt & H.Scnheid., Phytotaxa 7 (2010) 57; --Doryopteris wallichii J.Sm., Journal of Botany 4 (1841) 404; --Type: Cuming 238 (K, holo; BM, C, G, L, iso). Endemic to Philippines. LUZON. Note by J.F. Barcelona (15-Aug-14): Philippine specimens under Doryopteris ludens auct. non. (Wall. ex Hook) J.Sm. refer to this species. Photos 

Ceratopteris Brongn.

1. Ceratopteris oblongiloba Masuyama & Watano, Acta Phytotax. Geobot. 61 (2010) 84; --Type from Indonesia. Cambodia, Java, Sumatra, Thailand, Philippines. LUZON: Rizal.
2. Ceratopteris thalictroides (L.) Brongn., Bull. Sci. Soc. Philom. Paris 8 (1821) 186; --Batuyong et al., Biodiversitas 21 (2020) 3975; --Acrostichum thalictroides L., Sp. Pl. 2(1753) 1070; --Type from Sri Lanka. --Naias obvoluta Blanco, Fl. Filip. Ed. 2 (1845) 460. --Ceratopteris siliquosa (L.) Copel., PJS 6 (1935) 107; --Acrostichum siliquosum L., Sp. Pl. 2 (1753) 1070. Asia, Oceania, Central America. LUZON: Ilocos Norte, Zambales, MINDANAO: Davao, SAMAR. Conservation status: Endangered (DAO 2017-11). Photos

Excluded:

1. Ceratopteris gaudichaudii var. vulgaris Masuyama & Watano; Acta Phytotax. Geobot. 61 (2010) 79. Mistakenly reported for the Philippines by Kinosian et al. (2020) (S.P. Kinosian, pers. comm. 8-Sep-20).

Cheilanthes Sw. = Oeosporangium p.p.

Coniogramme Fée

1. Coniogramme fraxinea (D.Don) Diels in Engl. & Prantl, Nat. Pflanzenfam. 1 (1899) 262; --Diplazium fraxineum D.Don, Prodr. Fl. Nepal. (1825) 12. --Coniogramme macrophylla (Blume) Hieron., Hedwigia 57 (1916) 291; --Coniogramme javanica var. macrophylla Blume, Enum. Pl. Javae 2 (1828) 113. Bhutan, Borneo, ?Cambodia, China, India, Java, Java, Laos, Lesser Sunda Isls, Moluccas, Myanmar, Nepal, Pacific, Peninsular Malaysia, Solomon Isls, Sulawesi, Sumatra, Taiwan, Thailand, Tibet, Vietnam, Philippines. Photos
2. Coniogramme serrulata (Blume) Fée, Mém. Foug., 5. Gen. Filic. 167 (1850) t. 14 B. f. 2; --Gymnogramma serrulatum Blume, Enum. Pl. Jav. (1828) 113. --Coniogramme subcordata Copel., LPB 3 (1910) 823.Bhutan, China, India, Jammu & Kashmir, Japan, Java, Korea, Laos, Myanmar, Nepal, Pakistan, Russia, Sri Lanka , Taiwan, Thailand, Tibet, Vietnam, Philippines. Photos

Copelandiopteris Stone = Pteris

Doryopteris J.Sm.

1. Doryopteris kirkii (Hook.) Alston, Bol. Soc. Brot. sér. 2, 30 (1956) 14; --Coritico & Amoroso, Nature Conservation Research 5 (2020) 82; --Doryopteris concolor var. kirkii (Hook.) T.C.E.Fr., Wiss. Erg. d. Schwed. Rhodesia-Kongo Exp. 1911-12, Bot. 1 (1941) 4; --Cheilanthes kirkii Hook., Sec. Cent. Ferns (1861) t. 81. --Doryopteris cuspidata Copel., PJS 38 (1929) 148. Africa, Australia, Borneo, China, India, Indian Ocean, Lesser Sunda Isls, New Caledonia, New Guinea, Pacific, Sri Lanka, Sulawesi, Taiwan, Vietnam, Philippines. MINDANAO: Davao Oriental. Conservation status: Endangered (DAO 2017-11, as Doryopteris concolor). Photos

Haplopteris C.Presl

Last edited by Julie F. Barcelona, 17 May 2021

Photos

1. Haplopteris alternans (Copel.) S.Linds. & C.W.Chen, Gard. Bull. Singapore 66 (2014) 170; --Vittaria alternans Copel., PJS 1 (Suppl. 2) (1906) 157. --Vittaria hecistophylla Copel., PJS 81 (1952) 45. Sulawesi, Philippines. MINDANAO: Bukidnon, NEGROS. Photos
2. Haplopteris anguste-elongata (Hayata) E.H.Crane, Syst. Bot. 22 (1998) 514; --Vittaria anguste-elongata Hayata, Icon. Pl. Formosan. 6 (1916) 161. China, Ryukyu Isls, Taiwan, Philippines.
3. Haplopteris capillaris (Copel.) C.W.Chen, S.Linds. & K.T.Yong, Syst. Bot. 44 (2019) 491; --Vaginularia capillaris (Copel.) C.Chr., Index Filic., Suppl. Tert. (1934) 193; --Monogramma capillaris Copel., PJS 6c (1911) Bot. 147; --Type: Merrill 6961 (PNH, holo, lost; MICH, lecto; NY, P, isolecto), Negros: Mt Kanlaon, Apr-1910. Borneo, Taiwan, Philippines. NEGROS.
   
4. Haplopteris dareicarpa (Hook.) S.Linds. & C.W.Chen, Gard. Bull. Singapore 66 (2014) 171; --Monogramma dareicarpa Hook., Sp. Fil. 5 (1864) 121. ?Australia, Borneo, New Guinea, Peninsular Malaysia, Singapore, Solomon Isls, Philippines. Photos
   
5. Haplopteris elongata (Sw.) E.H.Crane, Syst. Bot. 22 (1998) 514; --Vittaria elongata Sw., Syn. Fil. 109 (1806) 302. --Vittaria intermedia Blume, Enum. Pl. Javae 2 (1828) 199. --Vittaria philippinensis Christ, Bull. Herb. Boissier II. 6 (1906) 1007. --?Vittaria crispomarginata Christ, Bull. Herb. Boissier II. 6 (1907) 1007; --Type: Loher s.n. (BM, P), Luzon: Makiling. Africa, Andaman Isls, Australia, Bangladesh, Bismarck Arch, Borneo, Cambodia, China, India, Indian Ocean, Laos, Lesser Sunda Isls, Moluccas, Myanmar, New Caledonia, New Guinea, Pacific, Peninsular Malaysia, Singapore, Solomon Isls, Sri Lanka, Sulawesi, Sumatra, Thailand, Tibet, Vietnam, Philippines. LUZON, PALAWAN. Note by J.F. Barcelona (8-Aug-2020): It is presently unclear if Vittaria crispomarginata is taxonomically distinct from Haplopteris elongata. Photos
6. Haplopteris ensiformis (Sw.) E.H.Crane, Syst. Bot. 22 (1998) 514; --Vittaria ensiformis Sw., Ges. Nat. Fr. Berl. Neu. Schr. 2 (1799) 134. --Vittaria incurvata Cav., Descr. (1802) 270. --Vittaria lineata auct. non Smith. --Vittaria pachystemma Christ, PJS 2 c (1907) 174. Africa, Australia, ?Bangladesh, Bismarck Arch, Borneo, Cambodia, China, India, Indian Ocean, Java, Laos, Lesser Sunda Isls, Moluccas (Seram), New Guinea, Pacific, Palau Isls, Peninsular Malaysia, Ryukyu Isls, Singapore, Solomon Isls, Sulawesi, Sumatra, Taiwan, Thailand, Vietnam, Philippines. MINDANAO: Bukidnon, PALAWAN. Photos
7. Haplopteris mindanaoensis S.Linds. & C.W.Chen, Gard. Bull. Singapore 70 (2018) 184; --Type: L.-Y. Kuo 3659 (PNH, holo; CMUH, SING, TAIF, iso), Mindanao: Mt Kitanglad, 18-Dec-2012. Endemic to the Philippines. MINDANAO: Bukidnon.
8. Haplopteris scolopendrina (Bory) C.Presl, Tent. Pterid. (1836) 141; --Vittaria scolopendrina (Bory) Thwaites, Enum. pl. Zeyl. (1864) 381; --Pteris scolopendrina Bory, Voy. 2 (1804) 323. --Vittaria subcoriacea Christ, PJS 2 c (1907) 175. Borneo, Indian Ocean, Moluccas, New Guinea, Pacific, Peninsular Malaysia, Solomon Isls, Sri Lanka, Sulawesi, Sumatra, Thailand, Philippines. MINDANAO: Bukidnon, PALAWAN. Photos
9. Haplopteris stenophylla (Copel.) C.W.Chen & S.Linds., Kew Bull. 76 (2021); --Vittaria stenophylla Copel., PJS 40 (1929) 312; --Type: Copeland 122 (MICH, lecto; BO, K, M, MU, NY, SING, UC, US, isolecto), Luzon: Benguet Prov, Mt Santo Tomas, May-1909, 2300m. Endemic to the Philippines. LUZON: Abra, Benguet, Kalinga, SAMAR.
   
10. Haplopteris taeniophylla (Copel.) E.H.Crane, Syst. Bot. 22 (1998) 514; --Vittaria taeniophylla Copel., PJS 1 (Suppl. 2) (1906) 157. Bhutan, Cambodia, China, India, Japan, Laos, Myanmar, Nepal, Sulawesi, Thailand, Tibet, Vietnam, Philippines. LUZON.
11. Haplopteris zosterifolia (Willd.) E.H.Crane, Syst. Bot. 22 (1998) 514; --Vittaria zosterifolia Willd., Sp. Pl. 5 (1810) 406. --Vittaria copelandii Alderw., Bull. Jard. Bot. Buitenzorg Ser. 2, VII (1912) 28. --Vittaria merrillii Christ, PJS 2 (1907) 174. Bismarck Arch, Bonin Isls, Borneo, Cambodia, China, India Andaman Isls, Indian Ocean, Java, Laos, Lesser Sunda Isls, Moluccas, Myanmar, Nepal, New Caledonia, New Guinea, Nicobar Isls, Pacific, Ryukyu Isls, Solomon Isls, South Korea, Sri Lanka, Sulawesi, Taiwan, Thailand, Vietnam, Philippines.

Mickelopteris Fraser-Jenk.

Last edited by Julie F. Barcelona, 19 June 2019

1. Mickelopteris cordata (Roxb. ex Hook. & Grev.) Fraser-Jenk., Annot. Checkl. Ind. Pterid. 1 (2016) 247; --Parahemionitis cordata (Roxb. ex Hook. & Grev.) Fraser-Jenk., New Sp. Syndr. Indian Pteridol. (1997) 187; --Hemionitis cordata Roxb. ex Hook. & Grev., Ic. Fil. (1828) t. 64. Bangladesh, Cambodia, China, India, Laos, Lesser Sunda Isls, Myanmar, Peninsular Malaysia, Sri Lanka, Taiwan, Thailand, Vietnam, Philippines. Photos

Monogramma Comm. ex Schkuhr. = Haplopteris

Oeosporangium Vis.

1. Oeosporangium belangeri (Bory) Fraser-Jenk., Annot. Checkl. Ind. Pterid. 1 (2016) 255; --Cheilanthes belangeri (Bory) C.Chr., Index Filic. (1905) 172; --Pteris belangeri Bory, Voy. Indes Or. 2 (1833) 44. Bangladesh, Borneo, Cambodia, China, India, Laos, Myanmar, Nepal, Peninsular Malaysia, Thailand, Vietnam, Philippines. LUZON.
2. Oeosporangium chusanum (Hook.) Fraser-Jenk., Annot. Checkl. Ind. Pterid. 1 (2016) 256; --Cheilanthes mysurensis var. chusana (Hook.) Christ, Ind. (1901) 293; --Cheilanthes chusana Hook., Sp. Fil. 2 (1852) 95. China, Japan, Laos, North Korea, South Korea, Sulawesi, Taiwan, Vietnam, Philippines. LUZON.
   
3. Oeosporangium tenuifolium (Burm.f.) Fraser-Jenk. & Pariyar, Annot. Checkl. Ind. Pterid. 1 (2016) 261; --Cheilanthes tenuifolia (Burm.f.) Sw., Syn. Fil. (1806) 129 Andaman Isls, Australia, Bangladesh, Borneo, Cambodia, China, India, Laos, Lesser Sunda Isls, Moluccas, Nepal, New Caledonia, Pacific, Pakistani Kashmir, Palau Isls, Peninsular Malaysia, Singapore, Solomon Isls, Sri Lanka, Taiwan, Thailand, Vietnam, Philippines. PALAWAN. Photos

Insufficiently known:

1. Cheilanthes javensis (Desv.) T.Moore, Index Fil. (1861) 244; --Cincinalis javensis Desv., Berl. Mag. 5 (1811) 329. --Cheilanthes densa (J.Sm.) Fée, Mém. Foug. 5 (1852) 156; --Notholaena densa J.Sm., J. Bot. (Hooker) 3 (1841) 395; --Type: Cuming 282 (K, MICH, UC, US), Luzon. --Cheilanthes dilimanensis P.M.Zamora, Kalikasan 3 (1974) 193; --Type: Zamora 71-886 (PUH, holo; MICH, iso), Luzon: Quezon City, Diliman, 1971. Note by J.F. Barcelona (18-Aug-20): It is presently unclear to which species of Oeosporangium Philippine specimens of this taxon belong.

Onychium Kaulf.

1. Onychium japonicum (Thunb.) Kunze, Bot. Zeitung (Berlin) 6 (1848) 507. Bonin Isl, China, Japan, Java, Lesser Sunda Isls, Ryukyu Isls, South Korea, Taiwan, Thailand, Philippines. Photos  
2. Onychium siliculosum (Desv.) C.Chr., Index Filic. fasc. 8 (1906) 469; --Pteris siliculosa Desv., Mag. Neuesten Entdeck. Gesammten Naturk. Ges. Naturf. Freunde Berlin 5 (1811) 324. ?Andaman Isls, Andaman Isls, Australia, Bangladesh, Bhutan, Bismarck Arch, Cambodia, China, India, Java, LaosThailand, Lesser Sunda Isls, Myanmar, Nepal, New Guinea, Peninsular Malaysia, Sulawesi, Taiwan, Tibet, Vietnam, Philippines. PALAWAN. Photos

Paragymnopteris K.H.Shing

1. Paragymnopteris vestita (Wall. ex C.Presl) K.H.Shing, Indian Fern J. 10 (1994) 230. 1994; --Gymnopteris vestita (Wall. ex C.Presl) Underw., Bull. Torrey Bot. Club 29 (1902) 627; --Gymnogramma vestita Wall. ex C.Presl., Tent. Pterid. (1836) 218. Bhutan, China, India, Myanmar, Nepal, Pakistan, Taiwan, Thailand, Tibet, Philippines.

Parahemionitis Panigrahi = Mickelopteris (Philippines)

Pityrogramma Link [Not native]

1. Pityrogramma calomelanos (L.) Link, Handb. Gewachs. 3 (1833) 20; --Acrostichum calomelanos L., Sp. Pl. 2 (1753) 1072. Americas. MINDANAO: Bukidnon. Not native to the Philippines.  Photos

Pteris L.

1. Pteris amoena Blume, Enum. Pl. Jav. (1828) 210. --Pteris purpureorhachis Copel., PJS 12 (1917) 48. Borneo, China, India, Japan, Java, Laos, Myanmar, Solomon Isls, Taiwan, Thailand, Tibet, Vietnam, Yakushima Isl, Philippines.
   
2. Pteris arisanensis Tagawa, Acta Phytotax. Geobot. 6 (1936) 102. Bhutan, China, Hong Kong, India, Myanmar, Nepal, Sri Lanka, Taiwan, Thailand, Tibet, Vietnam, Philippines.
   
3. Pteris armata C.Presl, Reliq. Haenk. 1 (1825) 56. Endemic to the Philippines. PALAWAN. Photos
4. Pteris biaurita L., Sp. Pl. 2 (1753) 1076. Photos 
   
   • ssp. fornicata Fraser-Jenk., Taxon. Revis. Indian Subcontinental Pteridophytes (2008) 116. --Pteris pacifica Hieron., Hedwigia 66 (1914) 355. --Pteris cumingii Hieron., Hedwigia 66 (1914) 358. Andaman Isls, Australia, Bangladesh, Bhutan, Bismarck Arch, Borneo, Cambodia, China, India, Indian Ocean, Laos, Lesser Sunda Isls, Moluccas, Myanmar, Nepal, Pacific, Palau Isls, Peninsular Malaysia, Solomon Isls, Sri Lanka, Sulawesi, Sumatra, Taiwan, Thailand, Tibet, Vietnam, Philippines. MINDANAO: Bukidnon, PALAWAN.
   • ssp. walkeriana Fraser-Jenk. & Rajkumar, Taxon. Revis. Indian Subcontinental Pteridophytes (2008) 115. Andaman Isls, Bangladesh, Bhutan, China, India, Java, Laos, Myanmar, Nepal, Thailand, Tibet, Philippines.
5. Pteris blumeana J.Agardh, Rec. Pterid. (1839) 22. Bangladesh, Bismarck Arch, Cambodia, ?China, India, Java, Laos, Moluccas, Myanmar, New Guinea, Solomon Isls, Sulawesi, Thailand, Vietnam, Philippines. Photos  
6. Pteris brevis Copel., PJS 81 (1952) 7. Endemic to the Philippines. BASILAN. Conservation status: Vulnerable (DAO 2017-11).
7. Pteris calocarpa (Copel.) M.G.Price, Kew Bull. 29 (1975) 727; --Copelandiopteris calocarpa (Copel.) Stone, Webbia 28 (1973) 493; --Schizostege calocarpa Copel., PJS 1 (1906) 155. Endemic to the Philippines. MINDANAO: Zamboanga. Conservation status: Critically Endangered (DAO 2017-11).
8. Pteris cretica L., Mant. Pl. 1 (1767) 130. --Pteris stenophylla sensu Copel., FFP 1 (1958) 129, auct. non Hook. & Grev. Photos
   
   • ssp. cretica. Africa, Bangladesh, Bhutan, China, Europe, India, Indian Ocean, Japan, Middle East, Myanmar, Nepal, Pacific, Pakistan, ?Peninsular Malaysia, Ryukyu Isls, South Korea, Sri Lanka, SW Asia, Taiwan, Thailand, Tibet, Vietnam, Philippines. PALAWAN.
   • ssp. laeta (Wall. ex Ettingsh.) Fraser-Jenk., Taxon. Revis. Indian Subcontinental Pteridophytes (2008) 100; --Pteris laeta Wall. ex Ettingsh., Farnkr. Jetztw. (1865) 96. Afghanistan, Azerbaijan, Bhutan, Cambodia, China, India, Iran, Japan, Laos, Lesser Sunda Isls, Myanmar, Nepal, Pakistan, Sri Lanka, Sulawesi, Taiwan, Tibet, Turkey, Vietnam, Philippines.
9. Pteris dataensis Copel., PJS 81 (1952) 7. Endemic to the Philippines. LUZON. Conservation status: Vulnerable (DAO 2017-11).
10. Pteris distans J.Sm., J. Bot. 3 (1841) 405. Endemic to the Philippines. LUZON. Possibly a hybrid or a bizarre freak.
11. Pteris edanoi Copel., PJS 81 (1952) 8, as 'edanyoi'. Endemic to the Philippines. LUZON.
12. Pteris elmeri Christ. ex Copel., PJS 81 (1952) 7. Endemic to the Philippines. LUZON.
13. Pteris endoneura M.G.Price, Kew Bull. 29 (1975) 727; --Copelandiopteris endoneura (M.G.Price) Stone, Kalikasan 5 (1976) 330. Endemic to the Philippines. LUZON: Camarines Norte, Quezon, SAMAR. Conservation status: Endangered (DAO 2017-11). Photos
14. Pteris ensiformis Burm., Fl. Ind. (1768) 230. Photos
    • var. ensiformis. --Pteris mutilata auct. non L. Bangladesh, Bhutan, Borneo, Cambodia, China, India, Laos, Lesser Sunda Isls, Moluccas, Myanmar, Nepal, New Caledonia, New Guinea, Nicobar Isls, Pacific, Palau Isls, Peninsular Malaysia, Ryukyu Isls, Singapore, Solomon Isls Bismarck Arch, Sri Lanka, Sulawesi, Taiwan, Thailand, Vietnam, Philippines.
    • var. victoriae Hort. Not native to the Philippines. Probably best treated as a garden cultivar.
15. Pteris glaucovirens Goldm. ex Hieron., Hedwigia 66 (1914) 339. Borneo, Philippines. LUZON.
16. Pteris heteromorpha Fée, Mém. Foug. 5 (1852) 127. Borneo, China, India, Laos, Myanmar, Sulawesi, Sulawesi, Thailand, Vietnam, Philippines. LUZON. Photos  
17. Pteris khasiana Hieron., Hedwigia 55 (1914) 364. Photos
    
    • ssp. fauriei (Hieron.) Fraser-Jenk., Annot. Checkl. Ind. Pterid. 1 (2016) 292; --Pteris fauriei Hieron., Hedwigia 55 (1914) 345; --Hatusima, Mem. Fac. Kagoshima Univ. 5 (1966) 20. --Pteris rigidula Copel., FFP 1 (1958) 137. China, Japan, Micronesia, Ryukyu Isls, South Korea, Taiwan, Vietnam, Philippines.
18. Pteris loheri Copel., PJS 81 (1952) 8. Endemic to the Philippines. LUZON.
19. Pteris longipinnula Wall. ex J.Agardh, Recens. Spec. Pter. (1839) 19. Bangladesh, Borneo, China, India, Japan, Lesser Sunda Isls, Moluccas, Myanmar, Nepal, Peninsular Malaysia, Sulawesi, Thailand, Philippines. Photos
20. Pteris luzonensis Hieron., Hedwigia 66 (1914) 370. Endemic to the Philippines. LUZON, MINDORO.
21. Pteris macgregorii Copel., PJS 81 (1952) 9. Endemic to the Philippines. LUZON. Conservation status: Vulnerable (DAO 2017-11).
22. Pteris melanocaulon Fée, Gen. (1850-52) 127. Sulawesi, Philippines. PALAWAN. Photos
23. Pteris melanorhachis Copel., PJS 81 (1952) 9. Endemic to the Philippines. LUZON.
24. Pteris mertensioides Willd., Sp. Pl. (1810) 394. --Pteris gardneri (Fée) Hook., Sp. Fil. 2 (1858) 206; --Litobrochia gardneri Fée, Gen. (1850-52) 139. --Pteris patens Hook., Sp. Fil. 2 (1858) 177, nom. illeg. auct. non Kunze. Andaman Isls, Bhutan, Borneo, Borneo, India, Java, Moluccas, Pacific, Peninsular Malaysia, Singapore, Sri Lanka, Sulawesi, Thailand, Philippines. MINDANAO: Bukidnon, PALAWAN. Photos  
25. Pteris micracantha Copel., PJS 81 (1952) 10. Endemic to the Philippines. LUZON. Conservation status: Vulnerable (DAO 2017-11).
26. Pteris mucronulata Copel., PJS 81 (1952) 10. Endemic to the Philippines. PALAWAN. Conservation status: Vulnerable (DAO 2017-11).
27. Pteris opaca (C.Presl) J.Sm. ex Fée, Mém. Foug., 5. Gen. Filic. 242 (1850) 358; --Pycnodoria opaca C.Presl, Abh. Kön. Böhm. Ges. Wiss. 5(6) (Epim. Bot.) (1851) 101. --Pteris taenitis Copel., PJS 7 (1912) 63. Borneo, Moluccas, New Guinea, Sulawesi, Philippines. BOHOL, LUZON: Tayabas, MINDANAO, PALAWAN, SAMAR. Conservation status: Vulnerable (DAO 2017-11 as Pteris taenitis). Photos
28. Pteris oppositipinnata Fée, Mém. Foug. 10 (1865) 17. --Pteris asperula J.Sm. ex Hieron., Hedwigia 66 (1914) 302. --Pteris caesia Copel., PJS 1 (Suppl. 2) (1906) 156. Borneo, Sulawesi, Philippines. PALAWAN. Photos  
29. Pteris pachysora (Copel.) M.G.Price, Kew Bull. 29 (1975) 727; --Copelandiopteris pachysora (Copel.) Stone, Webbia 28 (1973) 493; --Schizostege pachysora Copel., PJS 1 (Suppl. 2) (1906) 155. Endemic to the Philippines. MINDANAO: Zamboanga. Conservation status: Critically Endangered (DAO 2017-11).
30. Pteris pedicellata Copel., PJS 81 (1952) 10. Endemic to the Philippines. LUZON.
31. Pteris pellucens J.Agardh, Rec. Sp. Gen. Pterid. (1839) 32. --Pteris philippinensis Fée, Mém. Foug. 8 (1857) 114; --Pteris longipes D.Don var. philippinensis (Fée) Christ. --Pteris longipes D.Don, Prod. Fl. Nepal. (1825) 15; --Copel., FFP 1 (1958) 133. Bhutan, Borneo, China, India, Laos, Lesser Sunda Isls, Myanmar, Nepal, Sulawesi, Taiwan, Thailand, Vietnam, Philippines. PALAWAN. Photos 
    
32. Pteris pellucida C.Presl, Reliq. Haenk. 1 (1825) 55. Borneo, Java, Lesser Sunda Isls, New Guinea, Philippines. Photos
33. Pteris ramosii Copel., PJS 81 (1952) 11. Endemic to the Philippines. LUZON. Possibly a variant of Pteris philippinensis. Conservation status: Vulnerable (DAO 2017-11).
34. Pteris rangiferina C.Presl ex Miq., Ann. Mus. Bot. Lugd.-Bat. 4 (1868) 95. Borneo, Java, Philippines.
35. Pteris ryukyuensis Tagawa, Acta Phytotax. Geobot. 4 (1935) 204; --Hatusima, Mem. Fac. Kagoshima Univ. 5 (1966) 20. --?Pteris cretica sensu Copel., FFP 1 (1958) 129, quoad pl. ex Batanes Islands, non L. Japan, Ryukyu Isls, Taiwan, Philippines. Photos
36. Pteris schlechteri Brause, Bot. Jahrb. Syst. 49 (1912) 33. Moluccas, New Guinea, Sulawesi, Philippines. LUZON, MINDANAO: Bukidnon, NEGROS. Photos  
37. Pteris semipinnata L., Sp. Pl. 2 (1753) 1076. Bangladesh, Bhutan, Borneo, China, India, Laos, Moluccas, Myanmar, Nepal, Ryukyu Isls, South Korea, Sulawesi, Sumatra, Taiwan, Thailand, Vietnam, Philippines. Photos  
38. Pteris setulosocostulata Hayata, Ic. Pl. Formos. 4 (1914) 241. China, Japan, Taiwan, Philippines.
39. Pteris squamipes Copel., PJS 81 (1952) 11. Endemic to the Philippines. MINDANAO. Conservation status: Vulnerable (DAO 2017-11).
40. Pteris tripartita Sw., J. Bot. (Schrader) 1800 (1801) 67. --Pteris microdictyon (Fée) Hook., Sp. Fil. 2 (1858) 228; --Litobrochia microdictyon Fée, Gen. (1850-52) 138. Africa, Andaman Isls, Australia, Bismarck Arch, Borneo, China, India, Indian Ocean, Java, Laos, Lesser Sunda Isls, Moluccas, New Caledonia, Nicobar Isls, Pacific, Peninsular Malaysia, Singapore, Solomon Isls, Sri Lanka, Sulawesi, Sumatra, ?Taiwan, Thailand, Vietnam, Philippines. MINDANAO: Bukidnon, PALAWAN. Photos  
41. Pteris vittata L. ex Hieron., Hedwigia 64 (1914) 290; --Batuyong et al., Biodiversitas 21 (2020) 3975. Africa, Andaman Isls, Armenia, Atlantic Ocean, Australia, Azerbajian, Bangladesh, Bismarck Arch, Borneo, Cambodia, China, Europe, India, Indian Ocean, Japan, Java, Laos, Lesser Sunda Isls, Middle East, Moluccas, Myanmar, Nepal, New Guinea, Nicobar Isls, Pakistan, Peninsular Malaysia, Ryukyu Isls, Singapore, Solomon Isls, Sri Lanka, Sulawesi, Sumatra, Taiwan, Thailand, Tibet, Vietnam, Philippines. LUZON: Ilocos Norte, MINDANAO: Bukidnon, PALAWAN. Photos
42. Pteris wallichiana J.Agardh., Rec. Sp. Gen. Pterid. (1839) 69. Bhutan, Borneo, China, India, Japan, Java, Laos, Lesser Sunda Isls, Nepal, New Guinea, Peninsular Malaysia, Ryukyu Isls, Sri Lanka, Sulawesi, Taiwan, Thailand, Tibet, Vietnam, Philippines. LUZON. Photos
43. Pteris whitfordii Copel., PJS 1 (Suppl. 4) (1906) 255. Endemic to the Philippines. MINDANAO: Bukidnon, NEGROS. Photos

Possibly present:

1. Pteris cadieri Christ, J. Bot. (Morot) 19 (1905) 72.
   
   • ssp. cadieri. Bangladesh, Cambodia, China, Ryukyu Isls, Taiwan, Thailand, Vietnam, ?Philippines.
2. Pteris grevilleana Wall. ex J.Agardh, Recens. Spec. Pter. (1839) 23.
   
   • ssp. digitata (Baker) Fraser-Jenk., Annot. Checkl. Ind. Pterid. 1 (2016) 289; --Pteris quadriaurita var. digitata Baker, J. Bot. (1879) 40. Borneo, China, Myanmar, Taiwan, Thailand, Vietnam, ?Philippines.
3. Pteris terminalis Wall. ex J.Agardh, Recens. Spec. Gen. Pterid. (1839) 20. --Pteris excelsa Gaudich., Voy. Uranie (1829) 388. Bhutan, Bismarck Arch, Borneo, China, Fiji, Hawaii, India, Japan, Lesser Sunda Isls, Moluccas, Nepal, New Guinea, Pakistan, Peninsular Malaysia, Solomon Isls, South Korea, Sulawesi, Taiwan, Tibet, Vietnam, ?Philippines.

Schizoloma Gaudich. = Taenitis

Syngramma J.Sm.

1. Syngramma alismifolia (C.Presl) J.Sm., Lond. J. Bot. 4 (1845) 168; --Diplazium alismifolium C. Presl, Rel. Haenk. 1 (1825) 49. Borneo, Cambodia, Java, ?New Guinea, Nicobar Isls, Peninsular Malaysia, Singapore, Sulawesi, Sumatra, Thailand, Vietnam, Philippines. MINDANAO: Bukidnon. Photos  
2. Syngramma vittiformis J.Sm., Lond. J. Bot. 4 (1845) 168. Borneo, Sulawesi, Philippines. LUZON, MINDANAO, SAMAR. Photos
3. Syngramma wallichii (Hook.) Bedd., Handb. Ferns Brit. India (1866) t.153; --Gymnogramma wallichii Hook., Sp. 5 (1864) 155. Borneo, Moluccas, Peninsular Malaysia, Sulawesi, Sumatra, Philippines. Photos

Taenitis Willd.

1. Taenitis blechnoides (Willd.) Sw., Syn. Fil. 24 (1806) 220; --Pteris blechnoides Willd., Phyt. (1794) 13. Australia, Bismarck Arch, Borneo, Cambodia, China, ?India, Laos, Lesser Sunda Isls, Moluccas, Myanmar, New Guinea, Nicobar Isls, Palau Isls, Peninsular Malaysia, Singapore, Solomon Isls, Sri Lanka, Sulawesi, Sumatra, Thailand, Vietnam, Philippines. MINDANAO: Bukidnon, PALAWAN. Photos  
2. Taenitis cordata (Gaudich.) Holttum, Amer. Fern J. 50 (1960) 112; --Schizoloma cordatum Gaudich., Ann. Sci. Nat. 3 (1824) 507. Sulawesi, Philippines. Conservation status: Vulnerable (DAO 2017-11). Photos

Vaginularia Fée

Last edited by Julie F. Barcelona, 7 September 2020

Photos 

1. Vaginularia junghuhnii Mett., Fil. Hort. Bot. Lips. (1856) 25. --Vaginularia paradoxa auct. non (Fée) Mett. --Monogramma paradoxa auct. non (Fée) Mett. Borneo, Lesser Sunda Isls, Moluccas, New Guinea, Sri Lanka, Sulawesi, Sumatra, Thailand, Taiwan, Philippines. LUZON: Camarines Sur (photos), Quezon (photos), MINDANAO: Bukidnon, North Cotabato (photos), NEGROS (photos). Photos
2. Vaginularia trichoidea Fée, Mém. Foug. 3 ((1851)) 54; --Monogramma trichoidea (Fée) J.Sm., J. Bot. 3 (1841) 394. Borneo, China, Java, Lesser Sunda Isls, Moluccas, New Guinea, Peninsular Malaysia, Singapore, Sulawesi, Sumatra, Taiwan, Thailand, Philippines. Photos

Vittaria Sm. = Haplopteris p.p.

Pteridryaceae

Pteridrys C.Chr. & Ching

1. Pteridrys microthecia (Fée) C.Chr. & Ching, Bull. Fan Mem. Inst. Biol. 5 (1934) 139; --Aspidium microthecium Fée, Mem. Foug. (1865) 37; --Type: Cuming 13 (RB, holo; BM, E, K, L, P, US, iso), Luzon: 1-Jan-1856. Endemic to the Philippines. LUZON: Mt Banahaw, MINDANAO: Agusan del Norts, Cotabato, Zamboanga, NEGROS, PALAWAN, SAMAR. Photos
2. Pteridrys syrmatica (Willd.) C.Chr. & Ching, Bull. Fan Mem. Inst. Biol. Bot. 5 (1934) 131; --Aspidium syrmaticum Willd., Sp. Pl., ed. 4 5 (1810) 237; --Type: not from the Philippines. --Polypodium oxyodon Baker, J. Bot. 17 (1879) 66; --Type: Burbidge 27 (K, lecto; K, isolecto), Sulu Archipelago. --Dryopteris syrmatica var. petiolosa Christ, PJS 2 Bot. (1907) 210; --Type: Copeland 928 (P, holo), Mindanao: Dist. of Davao, 1-Ap-1904.  Indonesia, Malaysia, Thailand, Philippines. LUZON: Laguna, MINDANAO: Davao, PALAWAN, SULU. Photos

Rhachidosoraceae

Rhachidosorus Ching

Last edited by Julie F. Barcelona, 18 June 2019

No photos

1. Rhachidosorus stramineus (Copel.) Ching, Acta Phytotax. Sin. 9 (1964) 74; --Athyrium stramineum Copel., PJS 3 (1908) 292. Sulawesi, Philippines.

Saccolomataceae

Orthiopteris Copel.

Edited by Pieter B. Pelser, 22 July 2015

Photos

1. Orthiopteris campylura (Kunze) Copel., Fern Fl. Philip. 1 (1958) 87; --Davallia campylura Kunze, Bot. Zeitung (Berlin) 8 (1850) 132; --Saccoloma campylurum (Kunze) Mett. Ann. Sci. Nat. Bot. sér. 4, 15 (1861) 80; --Davallia inaequalis var. minor Hook., Sp. Fil. 1 (1846) 180; --Type: Cuming 119 (K, lecto; K, MICH, isolecto), Luzon. --Saccoloma moluccanum (Blume) Mett. in Kuhn var. stenolobum Christ, Bull. Herb. Boissier, ser. 2 6 (1906) 1005; --Type: Loher s.n. (P, holo), Luzon: Mabacal/Rizal, Mar-1906. Borneo, Fiji, Java, Moluccas, New Guinea, Peninsular Malaysia, Sumatra, Vanuatu, Philippines. LEYTE, LUZON, MINDANAO, NEGROS. Photos
   
   • var. campylura. Moluccas, Papua New Guinea, Philippines. LEYTE, LUZON, MINDANAO, NEGROS.

Saccoloma Kaulf. = Orthiopteris

Salviniaceae (including Azollaceae)

Azolla Lam.

1. Azolla pinnata R.Br., Prodr. Fl. Nov. Holland. (1810) 167. Photos
   • ssp. asiatica R.M.K.Saunders & K.Fowler, J. Linn. Soc. Bot. 109 (1992) 349. ?Afghanistan, Bangladesh, Bhutan, Borneo, China, India, Jammu & Kashmir, Japan, Java, Lesser Sunda Isls, Moluccas, Myanmar, Nepal, New Guinea, Pakistan, Peninsular Malaysia, Ryukyu Isls, South Korea, Sri Lanka, Sulawesi, Sumatra, Taiwan, Thailand, Vietnam, Philippines.

   Possibly present:

1. Azolla filiculoides Lam., Encycl. 1 (1783) 343. Americas. Not native to the Philippines. Reported for the Philippines by Moody (1989) as a weed in rice, but no specimens cited.
   
2. Azolla rubra R.Br., Prodr. Fl. Nov. Holland. (1810) 167. Australia, ?Indonesia, ?Japan, New Zealand, ?New Guinea, ?Philippines. Photos

Salvinia Seguier [Not native]

1. Salvinia molesta D.Mitch., Brit. Fern Gaz. 10 (1972) 251. Invasive species (CABI 2017). Photos

Schizaeaceae

Actinostachys Wall.

1. Actinostachys digitata (L.) Wall. ex C.F.Reed, Bol. Soc. Broter. Ser. 2, 21 (1948) 130; --Schizaea digitata (L.) Sw., Syn. Fil. (1806) 150; --Copel., FFP 1 (1958) 32; --Acrostichum digitatum L., Sp. Pl. 2 (1753) 1068. Andaman Isls, Australia, Bangladesh, Bonin Isls, Borneo, Cambodia, China, India, Moluccas, Myanmar, New Guinea, Nicobar Isls, Pacific, Palau Isls, Peninsular Malaysia, Ryukyu Isls, Seychelles, Singapore, Solomon Isls, Sri Lanka, Sulawesi, Sumatra, Taiwan, Thailand, Vietnam, Philippines. NEGROS, PALAWAN, PANAY, LUZON: Ilocos Norte. Photos  
2. Actinostachys inopinata (Selling) C.F.Reed, Bol. Soc. Broter. ser. 2, 21 (1948) 130; --Coritico & Amoroso, Nature Conservation Research 5 (2020) 82; --Schizaea inopinata Selling, Svensk. Bot. Tidskr. 40 (1946) 274; --Copel., FFP 1 (1958) 32. Borneo, New Guinea, Palau Isls, Peninsular Malaysia, Sumatra, Philippines. BOHOL, MINDANAO: Davao Oriental. Conservation status: Endangered (DAO 2017-11). Photos
3. Actinostachys minuta Amoroso & Coritico, PhytoKeys 151 (2020) 60; --Type: Amoroso 11213 with Coritico (PNH, holo; BRIT, CMUH, iso), Mindanao: Davao Oriental, San Isidro, Mt Hamiguitan Range Wildlife Sanctuary, 10-Oct-2016. Endemic to the Philippines. MINDANAO: Davao Oriental. Conservation status: Critically Endangered (Amoroso et al. 2020). Photos
4. Actinostachys simplex Amoroso & Coritico, Phytotaxa 508 (2021) 102; --Type: Amoroso 20312 with Nobleza (PNH, holo; CMUH, iso), Mindanao: Davao Oriental, San Isidro, Mt Hamiguitan Range Wildlife Sanctuary, 1-Feb-2021. Conservation status: Critically Endangered (Amoroso et al. 2021). Photos
   

Schizaea Sm.

1. Schizaea dichotoma (L.) Sm., Mém. Acad. Turin 5 (1793) 422; --Copel., FFP 1 (1958) 32; --Acrostichum dichotomum L., Sp. Pl. 2 (1753) 1068. Africa, Andaman Isls, Australia, Bangladesh, Bismarck Arch, Borneo, Borneo, China, India, Indian Ocean, Moluccas, Myanmar, New Caledonia, New Guinea, Nicobar Isls, Pacific, Palau Isls, Peninsular Malaysia, Ryukyu Isls, Singapore, Solomon Isls, Sulawesi, Sumatra, Taiwan, Thailand, Vietnam, Philippines. Photos  
2. Schizaea malaccana Baker, Syn. Fil. (1868) 428; --Coritico & Amoroso, Nature Conservation Research 5 (2020) 82. Australia, Borneo, Moluccas, Myanmar, New Guinea, Peninsular Malaysia, Solomon Isls, Philippines. MINDANAO: Bukidnon, Davao Oriental. Conservation status: Endangered (DAO 2017-11).

Tectariaceae

Aenigmopteris Holttum = Tectaria

Arthropteris J.Sm. ex Hook.f.

1. Arthropteris articulata (Brack.) C.Chr., Index Filic., Suppl. Tert. (1934) 26; --Lastrea articulata Brack., U.S. Expl. Exped., Filic. 16 (1854) 191. --Arthropteris wollastonii (Ridl.) Holttum, Blumea 14 (1966) 229; --Polypodium wollastonii Ridl., Trans. Linn. Soc. London, Bot. 9 (1916) 262. Fiji, Moluccas, New Guinea, Sulawesi, Philippines.
   
2. Arthropteris palisotii (Desv.) Alston, Bol. Soc. Brot. sér. 2, 30 (1956) 6; --Aspidium palisotii Desv., Berl. Mag. 5 (1811) 320. --Arthropteris obliterata (R.Br.) J.Sm, Cat. cult. ferns (1827) 62; --Nephrodium obliteratum R.Br., Prodr. Fl. Nov. Holland. (1810) 148. --Arthropteris glabra Copel., Fragm. Fl. Philipp. (1905) 178. --Arthropteris repens (Brack.) C.Chr., Bull. Bishop Mus. 177 (1943) 48; --Nephrolepis repens Brack., U.S. Expl. Exped., Filic. 16 (1854) 209. --Nephrolepis trichomanoides J.Sm. ex C.Presl, Epimel. Bot. (1851) 44; --Type: Cuming 101 (L, P), Philippines. Africa, Australia, Borneo, China, India, Java, Lesser Sunda Isls, Madagascar, Moluccas, New Caledonia, New Guinea, Pacific, Peninsular Malaysia, Ryukyu Isls, Solomon Isls, Sri Lanka, Sulawesi, Sumatra, Taiwan, Thailand, Vietnam, Philippines. PALAWAN. Photos

Ctenitis (C.Chr.) C.Chr.

1. Ctenitis aciculata (Baker) Ching, Bull. Fan Mem. Inst. Biol., Bot. 8 (1938) 292; --Nephrodium aciculatum Baker, J. Linn. Soc., Bot. 22 (1886) 226. Borneo, Java, New Guinea, Solomon Isls, Sulawesi, Sumatra, Philippines. MINDANAO.
2. Ctenitis atrorubens Holttum, Blumea 31 (1985) 29. --Ctenitis vilis sensu Copel. (1960), pro parte, auct. non (Kunze) Ching. Endemic to the Philippines. Only known from the type.
3. Ctenitis boholensis Holttum, Blumea 31 (1985) 18. Endemic to the Philippines. BOHOL.
4. Ctenitis bulusanica Holttum, Blumea 31 (1985) 19. Endemic to the Philippines. LUZON.
5. Ctenitis decurrentipinnata (Ching) Tardieu & C.Chr., Notul. Syst. (Paris) 7 (1938) 86; --Dryopteris decurrentipinnata Ching, Bull. Fan Mem. Inst. Biol. 2 (1931) 195. China, Vietnam, Philippines. LUZON.
6. Ctenitis erythradenia Holttum, Blumea 31 (1985) 29. --Ctenitis vilis sensu Copel. (1960), pro parte, auct. non (Kunze) Ching. Endemic to the Philippines. LUZON.
7. Ctenitis humilis Holttum, Blumea 31 (1985) 22. Endemic to the Philippines. BOHOL, MINDORO. Photos
8. Ctenitis paleolata Copel., PJS 81 (1952) 24. Endemic to the Philippines. LUZON: Benguet (Pauai). c. 2400m. Known only from the type collection. Conservation status: Critically Endangered (DAO 2017-11).
9. Ctenitis pallens (Brack.) M.G.Price, Kalikasan 12 (1983) 155; --Lastrea pallens Brack., Wilkes. Expl. Exp. Bot. (1854) 197. --Ctenitis rizalensis (Christ) Copel., Gen. Fil. (1947) 125; --Dryopteris rizalensis Christ, Bull. Herb. Boiss. 2, 6 (1906) 1001. Bismarck Arch, Sulawesi, Philippines. LUZON: Rizal, Laguna (Mt Makiling), SAMAR, MINDANAO: Zamboanga (San Ramon), Davao (Mt Apo), NEGROS, PALAWAN. Terrestrial under moist forest shade, 400-1200m.
10. Ctenitis propinqua (J.Sm. ex C.Presl) Copel., Blumea 31 (1985) 18; --Lastrea propinqua J.Sm. ex C.Presl, Abh. Kön. Böhm. Ges. Wiss., ser. 5 6 (1851) 38. Endemic to the Philippines. LUZON: Pampanga (Mt Arayat), Laguna (Mt Makiling), Quezon (Mt Banahaw), Batangas (Mt Makulod).
11. Ctenitis setosa (C.Presl) Holttum, Novit. Bot. Univ. Carol. 1968 (1969) 20; --Lastrea setosa C.Presl, Abh. Königl. Böhm. Ges. Wiss., ser. 5 6 (1851) 400. --Ctenitis vilis sensu Copel. (1960), pro parte, auct. non (Kunze) Ching. Endemic to the Philippines.
12. Ctenitis silvatica Holttum, Blumea 31 (1985) 20. --Ctenitis vilis sensu Copel. (1960), pro parte, auct. non (Kunze) Ching. New Guinea, Philippines. Photos
13. Ctenitis subglandulosa (Hance) Ching, Bull. Fan Mem. Inst. Biol., Bot. 8 (1938) 302; --Alsophila subglandulosa Hance, Ann. Sci. Nat., Bot., sér. 5, 5 (1866) 253. --Ctenitis rhodolepis (Clarke) Ching, Bull. Fan Mem. Inst. Biol., Bot. 8 (1938) 300; --Nephrodium rhodolepis C.B.Clarke, Trans. Linn. Soc. London, Bot. 1 (1880) 526. Bangladesh, Bhutan, Bonin Isls, China, Fiji, India, Japan, Java, Lesser Sunda Isls, Moluccas, Myanmar, New Guinea, Peninsular Malaysia, Ryukyu Isls, Southern Marianas, Taiwan, Thailand, Vietnam, Volcano Isls, Philippines. LUZON, MINDORO, PANAY. Photos
14. Ctenitis subobscura (Christ) Holttum, Fern Gaz. 12 (1984) 320; --Phegopteris subobscura Christ, Bull. Herb. Boissier 6 (1898) 836. Java, Peninsular Malaysia, Sumatra, Thailand, Philippines. MINDANAO.

Excluded

1. Ctenitis subconnexa (Christ) Holttum var. subconnexa and var. alstonii Holttum. A mislocalization to the Philippines by Holttum, Fl. Males. ser. 2, 2 (1991) 28. The locality Batjan is in Halmahera (of the Moluccas).

Heterogonium C.Presl = Tectaria

Pleocnemia C.Presl

1. Pleocnemia brongniartii (Bory) Holttum, Kew Bull. 29 (1974) 350; --Arcypteris brongniartii (Bory) Holttum, Reinwardtia 1 (1951) 195; --Tectaria irregularis var. brongniartii (Bory) Copel. (1907); --Polypodium brongniartii Bory, Voy. Monde 1 (1828) 263. New Guinea, Philippines. LUZON, SIVUYAN. Photos
2. Pleocnemia conjugata (Blume) C.Presl, Abh. Königl. Böhm. Ges. Wiss., ser. 5 6 (1851) 619; --Aspidium conjugatum Blume, Enum. Pl. Javae 2 (1828) 169. Andaman Isls, Borneo, Java, Lesser Sunda Isls, Moluccas, Myanmar, New Guinea, Peninsular Malaysia, Sulawesi, Sumatra, Thailand, Philippines. PALAWAN.
3. Pleocnemia cumingiana C.Presl, Abh. Königl. Böhm. Ges. Wiss., ser. 5 6 (1851) 410. --Tectaria leuzeana sensu Copel. (1960), pro parte. American Samoa, ?Bangladesh, Fiji, Moluccas, Myanmar, Peninsular Malaysia, Sulawesi, Philippines. LUZON: Laguna, Rizal, MINDANAO. Photos
4. Pleocnemia irregularis (C.Presl) Holttum, Kew Bull. 29 (1974) 347; --Arcypteris irregularis (C.Presl) Ching, Sunyatsenia 5 (1940) 251; --Tectaria irregularis (C. Presl) Copel., PJS 2 c (1907) 416; --Polypodium irregulare C.Presl, Reliq. Haenk. 1 (1825) 25. American Samoa, Bangladesh, Bangladesh, Borneo, Cambodia, Fiji, India, Java, Lesser Sunda Isls, Moluccas, Myanmar, New Guinea, Palau Isls, Peninsular Malaysia, Singapore, Solomon Isls, Sulawesi, Sumatra, Thailand, Vietnam, Philippines. PALAWAN. Photos
5. Pleocnemia macrodonta (C.Presl. ex Fée) Holttum, Kew Bull. 29 (1974) 348; --Arcypteris macrodonta (C.Presl) Holttum, Reinwardtia 1 (1951) 194; --Dictyopteris macrodonta C.Presl ex Fée, Gen. Fil. (1852) 267. --Tectaria irregularis sensu Copel. (1960), pro parte. Bismarck Arch, Borneo, New Guinea, Sulawesi, Philippines. LUZON, MINDORO, SIBUYAN. Photos
6. Pleocnemia presliana Holttum, Reinwardtia 1 (1951) 183. Sulawesi, Philippines. LUZON: Cagayan, Aurora, Laguna, PALAWAN (Mt Gantung; Imolnod), LEYTE (Palo), MINDANAO: Davao (Mt Apo, Todaya), Surigao del Norte. Terrestrial, usually in partly shaded area, near sea-level to at least 400m. Photos

Polydictyum C.Presl

1. Polydictyum menyanthidis C.Presl, Abh. Kön. Böhm. Ges. Wiss. 5(6) (Epim. Bot.) (1851) 53; --Tectaria menyanthidis (C.Presl) Copel., PJS 2 (1907) 414 1907. Bismarck Arch, Moluccas, New Guinea, Solomon Isls, Vanuatu, Philippines.

Psomiocarpa C.Presl = Tectaria

Stenosemia C.Presl  = Tectaria, Heterogonium

Tectaria Cav.

1. Tectaria adenophora Copel., LPB 4 (1911) 1151. Endemic to the Philippines. ?LUZON, SIBUYAN. Conservation status: Other Threatened Species (DAO 2017-11).
2. Tectaria aenigma C.W.Chen & C.J.Rothf., J. Pl. Res. 131 (2017) 74; --Aenigmopteris dubia (Copel.) Holttum, Blumea 30 (1984) 4; --Ctenitis dubia (Copel.) Copel., Gen. Fil. (1947) 124; --Dryopteris dubia Copel., LPB 1 (1907) 235. Endemic to the Philippines. LUZON: Quezon (Mt Banahaw de Lucban), PANAY: Iloilo prov. (Callado, photos), NEGROS: Negros Oriental (Cuernos Mtns), LEYTE, CAMIGUIN. Terrestrial, 850-1200m. Photos
   
3. Tectaria angulata (Willd.) Copel., Sarawak Mus. J. 2 (1917) 370; --Polypodium angulatum Willd., Sp. Pl. 5 (1810) 185. --Tectaria trifolia (Alderw.) C.Chr., Bot. Jahrb. Syst. 66 (1934) 49; --Aspidium trifolium Alderw., Bull. Jard. Bot. Buitenzorg 7 (1912) 4. Borneo, Java, Lesser Sunda Isls, Moluccas, New Guinea, Peninsular Malaysia, Singapore, Solomon Isls, Sulawesi, Sumatra, Thailand, Philippines. MINDANAO: Bukidnon, PALAWAN. Photos
4. Tectaria aspidioides (C.Presl) Copel., PJS 38 (1929) 137; --Heterogonium aspidioides C.Presl, Epim. Bot. (1851) 143. ?Borneo, Lesser Sunda Isls, Solomon Isls, Sulawesi, Philippines. LEYTE, LUZON, MINDORO, SAMAR. Photos
5. Tectaria athyriosora M.G.Price, Kalikasan 3 (1974) 113. Endemic to the Philippines. BOHOL, SAMAR, MINDANAO: Surigao del Norte. Photos
6. Tectaria aurita (Sw.) S.Chandra, Kalikasan 12 (1983) 157; --Stenosemia aurita (Sw.) C.Presl, Tent. Pterid. (1836) 237; --Acrostichum auritum Sw., J. Bot. (Schrader) 2 (1801) 12. Borneo, Java, Lesser Sunda Isls, Moluccas, New Guinea, Pacific, Solomon Isls, Sulawesi, Sumatra, Philippines. Photos 
7. Tectaria barberi (Hook.) Copel., PJS 2 (1907) 414; --Polypodium barberi Hook., Sp. Fil. 5 (1864) 100. Borneo, Moluccas, Myanmar, Peninsular Malaysia, Singapore, Sumatra, Thailand, Philippines.
8. Tectaria beccariana (Ces.) C.Chr., Index Filic., Suppl. 3 (1934) 177; --Polypodium beccarianum Ces., Rendiconti Reale Accad. Sci. Fis. (1877) 30. Bismarck Arch, Borneo, Moluccas, New Guinea, Solomon Isls, Sulawesi, Philippines. LUZON, MINDANAO, NEGROS, SAMAR.
9. Tectaria calcarea (C.Presl) Copel., PJS 2 c (1907) 415; --Sagenia calcarea J.Sm., J. Bot. (Hooker) 3 (1841) 410. Endemic to the Philippines. POLILLO, BOHOL, LEYTE. Photos
10. Tectaria crenata Cav., Descr. Pl. (1802) 250. --Tectaria grandifolia sensu Copel. (1960), pro parte. Borneo, Java, Lesser Sunda Isls, Moluccas, Pacific, Palau Isls, Peninsular Malaysia, Solomon Isls, Sulawesi, Sumatra, Thailand, ?Vietnam, Philippines. PALAWAN. Photos
11. Tectaria coadunata (Wall. ex Hook. & Grev.) C.Chr., Contr. U. S. Natl. Herb. 26 (1931) 331; --Aspidium coadunatum Wall. ex Hook. & Grev., Ic, f. (1831) 202. --Tectaria christii Copel., PJS 2 (1907) 1151. Africa, Bangladesh, Bhutan, Borneo, China, India, Laos, Myanmar, Nepal, Sri Lanka, Sulawesi, Taiwan, Thailand, Tibet, Vietnam, Philippines. LUZON: Mountain Province, Benguet, MINDORO: Mindoro Oriental (Mt Halcon). Usually on limestone, c. 1500m. Photos
    
12. Tectaria decurrens (C.Presl) Copel., LPB 1 (1907) 234; --Aspidium decurrens C.Presl, Reliq. Haenk. 1 (1825) 28. --Tectaria peralata Copel., PJS 83 (1954) 98. Bangladesh, Borneo, China, India, Japan, Laos, Moluccas, Myanmar, New Guinea, Nicobar Isls, Pacific, Peninsular Malaysia, Ryukyu Isls, Solomon Isls, Sri Lanka, Sulawesi, Taiwan, Thailand, Vietnam, Philippines. PALAWAN. Photos
13. Tectaria dissecta (G.Forst.) Lellinger, Amer. Fern J. 58 (1968) 156; --Ctenitis dissecta (G.Forst.) Copel., Nov. Fl. Jap. 4 (1938) 91; --Polypodium dissectum G.Forst., Fl. Ins. Austr. (17860 81. Borneo, Java, Lesser Sunda Isls, Moluccas, New Guinea, Pacific, Peninsular Malaysia, Ryukyu Isls, Solomon Isls, Sulawesi, Taiwan, Philippines. PALAWAN. Photos
14. Tectaria dolichosora Copel., PJS 38 (1929) 136. Endemic to the Philippines. LUZON: Cagayan, Quezon (Infanta, Tignoan), Tayabas. Steep slopes near summits.
15. Tectaria griffithii (Baker) C.Chr., Index Filic., Suppl. 3 (1934) 180; --Nephrodium griffithii Baker, Syn. Fil. (1867) 300. Bangladesh, Bhutan, Cambodia, China, India, Laos, Myanmar, peninsular Malaysia, Taiwan, Thailand, Vietnam, Philippines. LUZON to MINDANAO. Photos
16. Tectaria hilocarpa (Fée) M.G.Price, Kalikasan 3 (1974) 175; --Leptochilus hilocarpus Fée, Mém. Foug. 2 (1845) 87. --Hemigramma latifolia (Meyen ex Goldm.) Copel., PJS 2 (1907) 406, excl. Hemionitis zollingeri and other latter names based on this; --Gymnopteris latifolia Meyen ex Goldm., Nov. Actorum Acad. Caes. Leop.-Carol. Nat. Cur. 1 (1843) 460. Borneo, Sulawesi, Philippines. LEYTE, LUZON, MINDANAO, PALAWAN, PANAY. Photos
17. Tectaria laxa (Copel.) M.G.Price, Kalikasan 1 (1972) 37; --Ctenitis laxa Copel., PJS 83 (1954) 98. Endemic to the Philippines. LUZON: Ilocos Norte (Mt Quebrada; Mt Siminublan). 80-1100m. Photos
18. Tectaria lobbii (Hook.) Copel., PJS 10 (1915) 146; --Aspidium lobbii Hook., Sp. Fil. 4 (1862) 59. Borneo, Philippines. BOHOL, MINDANAO: Surigao del Sur. Conservation status: Endangered (DAO 2017-11). Photos
    
    • var. lobbii.
19. Tectaria macleanii (Copel.) S.Y.Dong, Phytotaxa 122 (2013) 62; --Tectaridium macleanii Copel., PJS 30 (1926) 329; --Type: Maclean & Catatan 84 (PNH, holo, lost; MICH, neo; CAS, isoneo), Luzon: Cagayan (West Mountain, Claveria), 29-Dec-1919. --Tectaridium primitivum Copel., PJS 30 (1926) 329; --Wenzel 876 (PNH, holo, lost; F, MO, NY, UC, iso), Leyte: Buenavista (Jaro), 9-Jun-1914. Endemic to the Philippines. LEYTE, LUZON: Cagayan, Camarines Sur, Isabela, Quezon, PANAY, SAMAR. Conservation status: Endangered (DAO 2017-11). Photos
    
20. Tectaria manilensis (C.Presl) Holttum, Indian Fern J. 1 (1984) 36; --Lastreopsis manilensis (C.Presl) M.G.Price, Kalikasan 12 (1983) 155; --Ctenitis manilensis (C.Presl) Holttum, Novit. Bot. Univ. Carol. (1969) 36; --Lastrea manilensis C.Presl., Tent. Pterid. (1836) 76. --Ctenitis loheri (Christ) Copel., FFP 2 (1960) 287; --Athyrium loheri Christ, Bull. Herb. Boissier, sér. 2, 6 (1906) 1001. Myanmar, Peninsular Malaysia, Sulawesi, Thailand, Philippines. LUZON, PALAWAN.
21. Tectaria melanocaulos (Blume) Copel., PJS 2 c (1907) 416; --Aspidium melanocaulon Blume, Enum. Pl. Javae 2 (1828) 161. Andaman Isls, Borneo, Java, Lesser Sunda Isls, Moluccas, New Guinea, Peninsular Malaysia, Solomon Isls, Sulawesi, Sumatra, Thailand, ?Vietnam, Philippines. LUZON, NEGROS.
22. Tectaria membranacea (Hook.) Fraser-Jenk. & Kholia, Annot. Checkl. Ind. Pterid. 2 (2018) 392; --Aspidium membranaceum Hook., Sp. 6 (1864) 105; --Type: Cuming s.n. (K, lecto). --Tectaria devexa (Kunze ex Mett.) Copel., PJS 2 (1907) 415; --Aspidium devexum Kunze ex Mett., Ann. Mus. Bot. Lugd.-Bat. 1 (1864) 237. Australia, Borneo, China, Java, Lesser Sunda Isls, Moluccas, Myanmar, New Guinea, Pacific, Peninsular Malaysia, Ryukyu Isls, Solomon Isls, Sri Lanka, Sulawesi, Sumatra, Taiwan, Thailand, Vietnam, Philippines. Photos
    
23. Tectaria mesodon (Copel.) M.G.Price, Kalikasan 1 (1972) 37; --Ctenitis mesodon (Copel.) Copel., Gen. Fil. (1947) 124; --Dryopteris mesodon Copel., PJS 7 (1912) 54. Endemic to the Philippines. LUZON: Ilocos Norte, MINDANAO: Agusan del Norte (Mt Hilong-hilong), 120m.
24. Tectaria mindanaensis (Holttum) C.W.Chen & C.J.Rothf., J. Pl. Res. 131 (2017) 74; --Aenigmopteris mindanaensis Holttum, Blumea 30 (1984) 7. Endemic to the Philippines. MINDANAO. Conservation status: Vulnerable (DAO 2017-11).
    
25. Tectaria multicaudata (C.B.Clarke) Ching, Sinensia 2 (1931) 20; --Nephrodium multicaudatum C.B.Clarke, Trans. Linn. Soc. London, Bot. 1 (1880) 540. --Tectaria malayensis (Christ) Copel., PJS 2 c (1907) 416; --Aspidium malayense Christ, PJS 2 c (1907) 187. Bangladesh, ?Cambodia, India, Laos, Myanmar, Peninsular Malaysia, Singapore, Taiwan, Thailand, Vietnam, Philippines. LUZON, MINDANAO.
    
26. Tectaria nayarii Mazumdar, Phytotaxa 153 (2014) 298; --Heterogonium pinnatum (Copel.) Holttum, Sarawak Mus. J. 5 (1949) 163; --Stenosemia pinnata Copel., PJS 1 (Suppl.) (1906) 146; --Type: Copeland 1601 (MICH, holo), Mindanao: Zamboanga, San Ramon, 300ft, 8-Jan-1905. Andaman Isls, Borneo, Mariana Isls, Peninsular Malaysia, Solomon Isls, Sulawesi, Sumatra, Thailand, Philippines. MINDANAO: Zamboanga. Photos
27. Tectaria polymorpha (Wall. ex Hook.) Copel., PJS 2 (1907) 413; --Aspidium polymorphum Wall. ex Hook., Sp. Fil. 4 (1862) 54. Bangladesh, Bhutan, Cambodia, China, India, Java, Lesser Sunda Isls, Myanmar, Nepal, Ryukyu Isls, Sri Lanka, Sulawesi, Sumatra, Taiwan, Thailand, Vietnam, Philippines. LUZON, PALAWAN.
28. Tectaria psomiocarpa S.Y.Dong, Phytotaxa 122 (2013) 63; --Psomiocarpa apiifolia (J.Sm. ex Kunze) C.Presl, Abh. Königl. Böhm. Ges. Wiss., ser. 5 6 (1851) 521; --Polybotrya apiifolia J.Sm. ex Kunze Farnkr. 1 (1844) 142; --Type: Cuming 26 (MICH, holo; E, MO, P, iso), Luzon: 1936-1940. Endemic to the Philippines. BOHOL, LUZON: Cagayan, Isabela, Bataan, Rizal, Laguna (Mt Makiling), Quezon, POLILLO, MASBATE, LEYTE, PANAY, SAMAR, MINDANAO: Zamboanga (San Ramon). Terrestrial in forest, from near sea-level to c. 450m. Conservation status: Endangered (DAO 2017-11). Photos
29. Tectaria ramosii (Copel.) Holttum, Fl. Malesiana, Ser. 2, Pterid. 2 (1991) 46; --Ctenitis ramosii Copel., PJS 81 (1952) 24. Endemic to the Philippines. BOHOL (Dimiao), SAMAR (Catubig River). Possibly near sea-level to c. 300m. Photos
30. Tectaria repanda (Willd.) Holttum, Fl. Malesiana, Ser. 2, Pterid. 2 (1991) 80; --Aspidium repandum Willd., Sp. Pl. 5 (1810) 216. --Tectaria crenata sensu Copel. (1960), pro parte. Bismarck Arch, ?Java, Lesser Sunda Isls, Moluccas, Moluccas, New Guinea, Pacific, Solomon Isls, Sulawesi, Philippines.
31. Tectaria sagenioides (Mett.) Christenh., Phytotaxa 10 (2010) 58; --Heterogonium sagenioides (Mett.) Holttum, Sarawak Mus. J. 5 (1949) 161; --Ctenitis sagenioides (Mett.) Copel., Gen. Fil. (1947) 124; --Aspidium sagenioides Mett., Abh. Senckenberg. Naturf. Ges. 2 (1858) 397. --Ctenitis obscura (Hook.) Copel., FFP (1960) 287; --Heterogonium obscurum (Hook.) Holttum, Kalikasan 4 (1975) 218; --Polypodium obscurum Hook., Sp. Fil. 4 (1862) 237. Borneo, China, Java, Laos, Lesser Sunda Isls, Myanmar, Peninsular Malaysia, Singapore, Sulawesi, Sumatra, Thailand, Vietnam, Philippines.
32. Tectaria samariana S.Y.Dong, Phytotaxa 178 (2014) 225; --Heterogonium lobulatum Holttum, Blumea 35 (1991) 557; --Type: Price & Hernaez 399 (K, holo; MICH, iso), Samar: western Samar. Endemic to the Philippines. SAMAR. Only known from the type.
33. Tectaria siifolia (Willd.) Copel., PJS 2 (1907) 414; --Polypodium siifolium Willd., Sp. Pl. 5 (1810) 196. Andaman Isls, Australia, Bismarck Arch, Borneo, Java, Lesser Sunda Isls, Moluccas, New Guinea, Peninsular Malaysia, Sulawesi, Sumatra, Thailand, Philippines. Photos
34. Tectaria stalactica M.G.Price, Kalikasan 3 (1974) 116. Endemic to the Philippines. LUZON: Cagayan, Ilocos Norte, Isabela, POLILLO. Conservation status: Endangered (DAO 2017-11).
35. Tectaria subaequalis (Rosenst.) Copel., PJS 9 c (1914) 5; --Aspidium subaequale Rosenst., Repert. Spec. Nov. Regni Veg. 12 (1913) 176. --Tectaria profereoides (C.Chr.) S.Y.Dong, Phytotaxa 178 (2014) 226; --Heterogonium profereoides (C.Chr.) Copel.; --Aspidium profereoides C.Chr., PJS 2 (1907) 158. --Heterogonium aspidioides sensu Holttum, Kalikasan 4 (1975) 224, pro parte. Borneo, New Guinea, Solomon Isls, Philippines. MINDANAO, MINDORO.
    
36. Tectaria sulitii Copel., PJS 81 (1952) 26. --Tectaria loheri Copel.,  PJS 81 (1952) 25. --Tectaria grandifolia sensu Copel. (1960), pro parte. According to Holttum, Fl. Males. ser. 2, 2 (1991) 91, there may be intermediates between this species and Tectaria crenata. Taiwan, Philippines. LUZON: Rizal (Montalban; Sumag), MINDANAO, NEGROS, PANAY. Photos
37. Tectaria suluensis Holttum, Blumea 35 (1991) 556; --Type: Kondo & Edaño 38624 (K, holo; SING, US, iso), Sulu: Siasi. Endemic to the Philippines. PALAWAN, SULU.
38. Tectaria tabonensis M.G.Price, Kalikasan 3 (1974) 118. Endemic to the Philippines. MINDANAO, PALAWAN. Conservation status: Other Threatened Species (DAO 2017-11).
39. Tectaria taccifolia (Fée) M.G.Price, Kalikasan 3 (1974) 175; --Hemigramma taccifolia (Fée) Copel., PJS 37 (1928) 406; --Leptochilus taccifolius Fée, Mém. Foug. 2 (1845) 89. Endemic to the Philippines. MINDORO. Photos
40. Tectaria trichotoma (Fée) Tagawa, Acta Phytotax. Geobot. 25 (1973) 180; --Aspidium trichotomum Fée, Mém. Foug. 5 (1852) 295. --Tectaria balabacensis (Christ) M.G.Price, Kalikasan 1 (1972) 37; --Ctenitis balabacensis (Christ) Copel., Gen. Fil. (1947) 124. Borneo, Vietnam, Philippines. BALABAC, LUZON: Quezon (Guinayangan), PALAWAN, PANAY, SIBUYAN. Petrophytic along rivers and streams in forests, 100-200m. Photos
41. Tectaria trifida (Fée) M.G.Price, Amer. Fern J. 64 (1974) 31; --Microsorum trifidum Fée, Mém. Foug. (1852) 269. --Tectaria irrigua (J.Sm. ex C.Presl) Copel., PJS 2 (1907) 413; --Aspidium irriguum C.Presl, Epim. Bot. (1851) 62. Endemic to the Philippines. LUZON, SAMAR.
42. Tectaria weberi Copel., PJS 2c (1912) 54; --Aspidium weberi (Copel.) C.Chr., Ind. Fil., Suppl. (1913) 9. Endemic to the Philippines. MINDANAO.
    
43. Tectaria wenzelii (Copel.) S.Y.Dong, Phytotaxa 178 (2014) 227; --Heterogonium wenzelii (Copel.) Holttum, Fl. Malesiana, Ser. 2, Pterid. 2 (1991) 111; --Ctenitis wenzelii Copel., PJS 81 (1952) 25. Endemic to the Philippines. LEYTE. Only known from the type. Conservation status: Critically Endangered (DAO 2017-11). Photos

Tectaridium Copel. = Tectaria

Thelypteridaceae

Amblovenatum J.P.Roux

1. Amblovenatum attenuatum (Kuntze) Barcelona & Pelser, Blumea 66 (2021) 97; --Amphineuron attenuatum (Kuntze) Holttum, Blumea 23 (1977) 215; --Dryopteris attenuata Kuntze, Revis. Gen. Pl. 2 (1891) 812; --Type: Cuming 327 (B, BM, BRI, E, G, K, L, LE, P, SING, UC, US, syn), Samar: 1841. --Cyclosorus alatellus auct. non (Christ) Copel., sensu Copel. (1960), excl. syn. Lastrea erubescens sensu Copel. (1960). Moluccas, Sulawesi, Philippines. LUZON, MINDANAO, SAMAR.
2. Amblovenatum immersum (Blume) Mazumdar, Int. J. Adv. Res. Innov. Ideas Educ. 3 (2017) 5060; --Amphineuron immersum (Blume) Holttum in B.K.Nayar & S.Kaur, Companion Handb. Ferns Brit. India (1974) 203; --Lastrea immersa (Blume) T.Moore, Index Fil. (1857) lxxxix; --Aspidium immersum Blume, Enum. Pl. Javae 2 (1828) 156. --Lastrea verrucosa (Kunze) C.Presl. Australia, Borneo, China, ?India, Java, Lesser Sunda Isls, Moluccas, Myanmar, New Guinea, Nicobar Isls, Peninsular Malaysia, Sulawesi, Sumatra, New Caledonia, Thailand, Vanuatu, Philippines. PALAWAN.
3. Amblovenatum opulentum (Kaulf.) J.P.Roux, Strelitzia 23 (2009) 201; --Thelypteris opulenta (Kaulf.) Fosb.in Fosb. & Sachet, Smithsonian Contr. Bot. 8 (1972) 3; --Amphineuron opulentum (Kaulf.) Holttum, Blumea 19 (1971) 45; --Aspidium opulentum Kaulf., Enum. Filic. (1824) 238. --Cyclosorus extensus (Blume) H.Ito, Bot. Mag. (Tokyo) 51 (1937) 713; --Aspidium extensum Blume, Enum. Pl. Javae 2 (1828) 156. Andaman Isls, Australia, Borneo, Chagos Arch, China, Christmas Isls, India, Java, Lesser Sunda Isls, Madagascar, Micronesia, Moluccas, Myanmar, New Guinea, Nicobar Isls, Pacific, Peninsular Malaysia, Singapore, Sri Lanka, Sulawesi, Sumatra, Thailand, Philippines. PALAWAN.
4. Amblovenatum terminans (Hook.) J.P.Roux, Strelitzia 23 (2009) 201; --Amphineuron terminans (Hook.) Holttum, Amer. Fern J. 63 (1973) 82. --Thelypteris wagneri Fosberg & Sachet, Smithsonian Contr. Bot. 8 (1972) 6. --Cyclosorus interruptus sensu Copel. (1960), non (Willd.) H.Ito Sri Lanka to Polynesia. Throughout the Philippines in the more seasonal regions. Terrestrial, common in thickets on hillsides and in valleys at low and medium elevation. Photos

Ampelopteris Kunze

1. Ampelopteris prolifera (Retz.) Copel., Gen. Fil. (1947) 144; --Hemionitis prolifera Retz., Observ. Bot. 6 (1791) 38. Tropical and subtropical regions of the Old World. In the Philippines recorded from LUZON: Abra, Rizal, Laguna, MINDANAO: Davao del Norte (Maco, Lake Leonard), Agusan del Sur (Agusan Marsh). Terrestrial, often forming mats or tangles in swampy sites, lake margins or river embankments. Low elevation. Photos  

Amphineuron Holttum = Amblovenatum

Chingia Holttum

1. Chingia christii (Copel.) Holttum, Kalikasan 3 (1974) 25; --Cyclosorus christii Copel., Fern Fl. Philip. 2 (1960) 362. Endemic to the Philippines. MINDANAO, MINDORO, NEGROS.
2. Chingia clavipilosa Holttum, Kalikasan 3 (1974) 23. Borneo, Java, Lesser Sunda Isls, Philippines.
   
   • var. clavipilosa. Borneo, Lesser Sunda Isls, Philippines.
3. Chingia ferox (Blume) Holttum, Blumea 19 (1971) 31; --Cyclosorus ferox (Blume) Ching, Bull. Fan Mem. Inst. Biol. Bot. 8 (1938) 167; --Aspidium ferox Blume, Enum. Pl. Javae 2 (1828) 153. Borneo, Java, Lesser Sunda Isls, Moluccas, New Guinea, Sulawesi, Taiwan, Philippines. MINDANAO: Bukidnon. Photos  
4. Chingia paucipaleata Holttum, Kalikasan 3 (1974) 26. Endemic to the Philippines. LUZON: Quezon. Conservation status: Vulnerable (DAO 2017-11 as Cyclosorus paucipaleatus (Holttum) Mazumdar & Mukhop.).
5. Chingia pricei Holttum, Kalikasan 3 (1974) 22. Endemic to the Philippines. LUZON: Benguet, MINDANAO: Misamis Occidental. Conservation status: Vulnerable (DAO 2017-11).
6. Chingia tenerior Holttum, Fl. Males. Ser. 2, 1 (1982) 395; --Type: Ramos BS 33066 (P, US), Luzon: Ilocos Norte, Baguio. Endemic to the Philippines. LUZON.
   
7. Chingia urens Holttum, Kalikasan 3 (1974) 28. Endemic to the Philippines. LUZON: Laguna, MINDANAO: Bukidnon. Conservation status: Critically Endangered (DAO 2017-11).

Christella H.Lév.

1. Christella acuminata (Houtt.) Holttum, Webbia 30 (1976) 192; --Cyclosorus acuminatus (Houtt.) Ching; --?Polypodium acuminatum Houtt., Nat. Hist. 2 (1783) 181. China, Japan, Ryukyu Isls, South Korea, Taiwan, Vietnam, Philippines. BABUYAN ISLS, LUZON. Photos
2. Christella arida (D.Don) Holttum, Companion Handb. Ferns Brit. India (1974) 206; --Cyclosorus aridus (D.Don) Ching, Bull. Fan Mem. Inst. Biol. Bot. 8 (1938) 194; --Aspidium aridum D.Don, Prodr. Fl. Nepal. (1825) 4. Australia, Bangladesh, Bhutan, Borneo, China, Fiji, India, Jammu & Kashmir, Lesser Sunda Isls, Moluccas, Myanmar, Nepal, New Caledonia, Samoa, Singapore, Sulawesi, Sumatra, Tibet, Taiwan, Thailand, Vanuatu, Vietnam, Vietnam. Photos
3. Christella dentata (Forssk.) Brownsey & Jermy, Brit. Fern Gaz. 10 (1973) 338; --?Polypodium dentatum Forssk., Fl. Aegypt.-Arab. (1775) 185. --Cyclosorus dentatus sensu Copel., 1960, pro parte. --Cyclosorus mindanaensis (Christ) Copel., Gen. Fil. (Copeland) (1947) 143; --Dryopteris mindanaensis Christ., PJS 2 (1907) 194. Pantropic. MINDANAO: Bukidnon. Throughout the Philippines. Terrestrial in exposed sites, light forests, along stream embankments, etc. Common at low elevation.
4. Christella parasitica (L.) H.Lév. ex Y.H.Chang, Taiwania 64 (2019) 380; --H.Lév., Fl. Kouy-Tcheou (1915) 473; --Polypodium parasiticum L., Sp. Pl. 2 (1753) 1090. --Cyclosorus dentatus sensu Copel. 1960, pro parte. --Cyclosorus benguetensis Copel., 81 (1952) 28. --Cyclosorus procurrens (Mett.) Copel., Fern. Fl. Philip. 2 (1960) 340; --Aspidium procurrens Mett., Ann. Mus. Bot. Lugduno-Batavi 1 (1864) 231. Africa, Australia, Bonin Isl, Borneo, Bangladesh, China, Guam, India, Japan, Laos, Lesser Sunda Isls, Moluccas, Myanmar, New Guinea, Nicobar Isls, Norfolk Isl, Pacific Isls, ?Pakistan, Peninsular Malaysia, Ryukyu Isls, South Korea, Sri Lanka, Sulawesi, Taiwan, Thailand, Vietnam, Volcano Isl, Philippines. Throughout the Philippines. PALAWAN. Terrestrial, open sites and degraded habitats generally, along stream embankments.
5. Christella quadrangularis (Fée) Holttum, Companion Beddome's Handb. Ferns Brit. India (1974) 207; --Nephrodium quadrangulare Fée, Gen. Fil. (1852) 308. --Christella hispidula (Decne.) Holttum, Kew Bull. 31 (1976) 312; --Aspidium hispidulum Decne., Nouv. Ann. Mus. 3 (1834) 346. --Cyclosorus falcatulus (Christ) Copel., Fern Fl. Philip. 2 (1960) 339; --Dryopteris parasitica var. falcatula Christ. Africa, Australia, Borneo, Central America, India, Java, Lesser Sunda Isls, Moluccas, Myanmar, New Guinea, Peninsular Malaysia, South America, Sri Lanka, Sulawesi, Sumatra, Vietnam, Philippines. PALAWAN.
   
6. Christella subdentata Holttum, Kew Bull. 31 (1976) 335. Endemic to the Philippines. LUBANG, LUZON, MINDORO.
7. Christella subpubescens (Blume) Holttum, Webbia 30 (1976) 193; --Cyclosorus subpubescens (Blume) Ching, Bull. Fan Mem. Inst. Biol. Bot. 8 (1938) 211; --Aspidium subpubescens Blume, Enum. Pl. Javae 2 (1828) 149. Andaman Isls, Australia, Bangladesh, Borneo, Cambodia, China, India, Japan, Lesser Sunda Isls, Moluccas, Myanmar, ?New Guinea, Pacific Isls, Peninsular Malaysia, Ryukyu Isls, Singapore, Sri Lanka, Sulawesi, Sumatra, Thailand, Vietnam, Philippines. PALAWAN.

Coryphopteris Holttum

1. Coryphopteris borealis Holttum, Blumea 23 (1976) 35. Endemic to the Philippines. LUZON: Benguet, Ifugao. Conservation status: Critically Endangered (DAO 2017-11).
2. Coryphopteris dura (Copel.) Holttum, Blumea 23 (1976) 44; --Lastrea dura (Copel.) Copel., Gen. Fil. (1947) 138; --Dryopteris dura Copel., LPB 3 (1910) 805. Endemic to the Philippines. MINDANAO (Mt Apo).
3. Coryphopteris pubirachis (Baker) Holttum, Blumea 23 (1976) 37. Bismarck Arch, Borneo, New Guinea, Pacific, Solomon Isls, Sulawesi, Philippines.
   
   • var. philippinensis Holttum, Blumea 23 (1976) 37. --Lastrea viscosa sensu Copel. (1960), auct. non (Baker) Bedd., pro specim. philipp. Endemic to the Philippines. Widespread, incl. PALAWAN. Photos
4. Coryphopteris squamipes (Copel.) Holttum, Blumea 23 (1976) 35; --Lastrea squamipes (Copel.) Copel., Fern Fl. Philip. 2 (1960) 325; --Dryopteris squamipes Copel., PJS 56 (1935) 99. Bismarck Arch, Moluccas, Philippines. LUZON: Benguet, Zambales, MINDANAO: Bukidnon, Davao del Sur, South Cotabato. Conservation status: Vulnerable (DAO 2017-11).

Cyclogramma Tagawa

1. Cyclogramma auriculata (J.Sm.) Ching., Acta Phytotax. Sin. 8 (1963) 317; --Cyclosorus auriculatus (J.Sm.) C.M.Kuo, Taiwania 47 ((2002)) 171; --Phegopteris auriculata J.Sm., Hist. Fil. (1875) 233. --Thelypteris simulans Ching, Bull. Fan Mem. Inst. Biol. Bot. 6 (1936) 280. Bhutan, China, India, Java, Nepal, Myanmar, Taiwan, Philippines. LUZON. Conservation status: Vulnerable (DAO 2017-11).

Cyclosorus Link

1. Cyclosorus interruptus (Willd.) H.Ito, Bot. Mag. (Tokyo) 51 (1937) 714; --Pteris interrupta Willd., Phytographia (1794) 13. --Cyclosorus gongylodes auctt., doubtfully based on Aspidium gongylodes Schkuhr. Africa, Americas, Andaman Isls, Australasia, Bangladesh, Bismarck Arch, Borneo, Cambodia, China, India, Japan, Java, Laos, Lesser Sunda Isls, Madagascar, Maldives, Micronesia, Moluccas, Myanmar, New Caledonia, Nepal, New Guinea, Pacific, Peninsular Malaysia, Ryukyu Isls, Singapore, Solomon Isls, South Korea, Sri Lanka, Sulawesi, Sumatra, Taiwan, Thailand, Vietnam, Philippines. PALAWAN. Photos

 Haplodictyum C.Presl = Pronephrium, Sphaerostephanos

Macrothelypteris Ching

1. Macrothelypteris polypodioides (Hook.) Holttum, Blumea 17 (1969) 29; --Alsophila polypodioides Hook.in Nightingale, Oceanic Sketches (1835) 131. --Lastrea leucolepis C.Presl, Epimel. Bot. (1851) 39. Australia, India, Lesser Sunda Isls, Micronesia, Moluccas, New Guinea, Pacific, Solomon Isls, Sulawesi, Taiwan, Philippines. PALAWAN. Photos
2. Macrothelypteris torresiana (Gaudich.) Ching, Acta Phytotax. Sin. 8 (1963) 310; --Lastrea torresiana (Gaudich.) T.Moore, Index Fil. (1858) 106; --Polystichum torresianum Gaudich., Voy. Uranie [Freycinet] (1828) 333. Australasia, Bangladesh, Bhutan, Borneo, China, Comoros, India, Japan, Java, Laos, La Reunion, Lesser Sunda Isls, Madagascar, Mauritius, Micronesia, Moluccas, Myanmar, Nepal, New Guinea, Pacific, Ryukyu Isls, Seychelles, Sumatra, Taiwan, Tibet, Vietnam, Solomon Isls, Sri Lanka, Sulawesi, Thailand, Philippines. PALAWAN. Photos

Mesophlebion Holttum

1. Mesophlebion crassifolium (Blume) Holttum, Blumea 22: 232 (1975); --Lastrea crassifolia (Blume) Moore; --Aspidium crassifolium Blume, Enum. Pl. Javae 2 (1828) 158. Borneo, Moluccas, Myanmar, New Guinea, Peninsular Malaysia, Singapore, Sulawesi, Sumatra, Thailand, Philippines. PALAWAN.
2. Mesophlebion rufescens Holttum, Blumea 22 (1975) 230. New Guinea, Philippines. BILIRAN, LUZON.

Metathelypteris Ching

1. Metathelypteris dayi (Bedd.) Holttum, Companion Handb. Ferns Brit. India (1974) 205; --Nephrodium dayi Bedd., J. Bot. 25 (1887) 323. --Thelypteris singalanensis sensu Holttum, auct. non (Baker) Ching. Borneo, Java, Moluccas, New Guinea, Peninsular Malaysia, Singapore, Sumatra, Thailand, Vietnam, Philippines. LUZON, MINDANAO.
2. Metathelypteris gracilescens (Blume) Ching, Acta Phytotax. Sin. 8 (1963) 305; --Lastrea gracilescens (Blume) T.Moore, Index Fil. (1858) 93; --Aspidium gracilescens Blume, Enum. Pl. Javae 2 (1828) 155. --Lastrea calva (Copel.) Copel., Gen. Fil. (1947) 138; --Dryopteris calva Copel., LPB 3 (1910) 808. Borneo, China, Java, Lesser Sunda Isls, Moluccas, New Guinea, Peninsular Malaysia, Sulawesi, Sumatra, Taiwan, Yakushima Isl, Philippines. LUZON, MINDANAO, MINDORO. Photos
3. Metathelypteris uraiensis (Rosenst.) Ching, Acta Phytotax. Sin. 8 (1963) 306; --Dryopteris uraiensis Rosenst., Hedwigia 56 (1915) 341. China, Ryukyu Isls, Taiwan, Yakushima Isls, Philippines. LUZON.

Nannothelypteris Holttum

1. Nannothelypteris aoristisora (Harrington) Holttum, Blumea 19 (1971) 38; --Cyclosorus aoristisorus (Harrington) Copel., Gen. Fil. (1947) 142; --Polypodium aoristisorum Harrington, J. Linn. Soc., Bot. 16 (1877) 30. Endemic to the Philippines. LUZON, PANAY. Conservation status: Vulnerable (DAO 2017-11). Photos
2. Nannothelypteris camarinensis Holttum, Kalikasan 5 (1976) 119. Endemic to the Philippines. LUZON. Conservation status: Vulnerable (DAO 2017-11).
3. Nannothelypteris inaequilobata Holttum, Kalikasan 2 (1973) 67. --Lastrea nervosa auct. non (Fée) Copel., sensu Copel. (1960), pro parte. Endemic to the Philippines. LUZON. Conservation status: Vulnerable (DAO 2017-11). Photos
4. Nannothelypteris nervosa (Fée) Holttum, Kalikasan 2 (1973) 66; --Lastrea nervosa (Fée) Copel., Fern Fl. Philip. 2 (1960) 323; --Phegopteris nervosa Fée -- Mém. Foug., 5. Gen. Filic. (1850-52) 244. Endemic to the Philippines. MINDANAO, SAMAR. Conservation status: Vulnerable (DAO 2017-11).
5. Nannothelypteris philippina (C.Presl) Holttum, Kalikasan 2 (1973) 66; --Physematium philippinum C.Presl, Epimel. Bot. (1851) 34. --Lastrea confusa (Copel.) Copel., Fern Fl. Philip. 2 (1960) 324; --Dryopteris confusa Copel., PJS c 6 (1911) 146. Endemic to the Philippines. LUZON. Conservation status: Vulnerable (DAO 2017-11).

Parathelypteris (H.Ito) Ching

1. Parathelypteris beddomei (Baker) Ching, Acta Phytotax. Sin. 8 (1963) 302; --Amauropelta beddomei (Baker) Y.H.Chang, Taiwania 64 (2019) 380; --Lastrea beddomei (Baker) Bedd., Ferns Brit. Ind. Corr. 2(supp. 16) (1870); --Nephrodium beddomei Baker, Syn. Fil. (Hooker & Baker) (1867) 267. Borneo, China, India, Japan, Java, Lesser Sunda Isls, Moluccas, Myanmar, New Guinea, Peninsular Malaysia, South Korea, Sri Lanka, Sulawesi, Sumatra, Taiwan, Yakushima Isl, Philippines. Perhaps best classified as Amauropelta beddomei based on unpublished data (TPG 2019). Photos
   • var. beddomei.
   • var. eugracilis (Copel.) Holttum; --Lastrea eugracilis Copel., Gen. Fil. (1947) 138. Endemic to the Philippines. MINDANAO.
2. Parathelypteris glanduligera (Kunze) Ching, Acta Phytotax. Sin. 8 (1963) 301; --Amauropelta glanduligera (Kunze) Y.H.Chang, Taiwania 64 (2019) 380; --Aspidium glanduligerum Kunze, Anal. (1837) 44. Bhutan, China, India, Japan, Korea, Laos, Myanmar, Nepal, Ryukyu Isls, Taiwan, Thailand, Vietnam, Philippines. LUZON. Perhaps best classified as Amauropelta glanduligera based on unpublished data (TPG 2019).
3. Parathelypteris grammitoides (Christ) Ching, Acta Phytotax. Sin. 8 (1963) 302; --Lastrea grammitoides (Christ) Copel., Fern Fl. Philip. 2 (1960) 320; --Aspidium grammitoides Christ, Bull. Herb. Boissier 6 (1898) 193. Japan, Java, Ryukyu Isls, Philippines. LUZON. Photos

Phegopteris Fée

1. Phegopteris aurita (Hook.) J.Sm., Cat. cult. Ferns (1857) 17; --Pseudophegopteris aurita (Hook.) Ching, Acta Phytotax. Sin. 8 (1963) 314; --Gymnogramma aurita Hook., Icon. Pl. 6 (1854) t. 974. Borneo, Bhutan, China, India, Japan, Myanmar, Nepal, New Guinea, Peninsular Malaysia, Tibet, Vietnam, Philippines. LUZON. Photos
   
2. Phegopteris decursive-pinnata Fée, Mém. Foug., 5. Gen. Filic. 242 (1850) t.20. China, Jammu & Kashmir, Japan, Java, Nepal, Korea, Ryukyu Isls, Sulawesi, Taiwan, Vietnam, Philippines. BATAN. Known only from one collection on Mt. Iraya, Batan I., Batanes (Barcelona 990).
3. Phegopteris paludosa (Blume) J.Sm., Hist. Fil. (1875) 233; --Pseudophegopteris paludosa (Blume) Ching, Acta Phytotax. Sin. 8 (1963) 315; --Aspidium paludosum Blume, Enum. Pl. Javae 2 (1828) 168. --Lastrea pyrrhorhachis sensu Copel., Fern Fl. Philipp. (1960) 330, pro parte. Borneo, China, Fiji, India, ?Java, Lesser Sunda Isls, Moluccas, New Guinea, Sri Lanka, Sulawesi, ?Sumatra, Thailand, Vietnam, Philippines. Photos

Possibly present:

1. Phegopteris microstegia (Hook.) Christenh., The Global Flora, Special edition, part 1 (2018) 25; --Nephrodium microstegium Hook., Sp. 4 (1862) 119.
   
   • ssp. microstegia. Bhutan, China, India, Japan, Myanmar, Nepal, Tibet, Vietnam, ?Philippines.

Plesioneuron (Holttum) Holttum

1. Plesioneuron savaiense (Baker) Holttum, Blumea 22 (1975) 240; --Nephrodium savaiense Baker, Ann. Bot. (Oxford) 5 (1891) 318. --Lastrea quadriaurita (Christ) Copel., Gen. Fil. (1947) 139; --Dryopteris quadriaurita Christ, PJS 2 c (1907) 209. Bismarck Arch, Moluccas, New Guinea, Pacific, Solomon Isls, Philippines. MINDANAO, NEGROS. Photos

Pneumatopteris Nakai

1. Pneumatopteris costata (Brack.) Holttum, Blumea 21 (1973) 305; --Goniopteris costata Brack., U.S. Expl. Exped., Filic. 16 (1854) 28. --Lastrea cavitensis Copel., PJS 81 (1952) 26. Australia, New Caledonia, New Guinea, Pacific, Solomon Isls, Lesser Sunda Isls, Philippines. LUZON.
2. Pneumatopteris glabra (Copel.) Holttum, Blumea 21 (1973) 301; --Cyclosorus glaber Copel., PJS 81 (1952) 34. --Cyclosorus viridis Copel., 81 (1952) 35. Endemic to the Philippines. BOHOL, LUZON, MINDANAO, SAMAR. Photos
3. Pneumatopteris laevis (Mett.) Holttum, Blumea 21 (1973) 308; --Aspidium laeve Mett., Abh. Senckenberg. Naturf. Ges. 2 (1858) 388. --Cyclosorus nitidulus auct. non (C.Presl) Copel., sensu Copel. (1960), pro parte. --Cyclosorus microlonchus auct. non (Christ) Ching, sensu Copel. (1960), pro parte. Endemic to the Philippines. LUZON, LEYTE, MINDANAO: Bukidnon, NEGROS, SAMAR. Photos
4. Pneumatopteris ligulata (C.Presl) Holttum, Blumea 21 (1973) 320; --Lastrea ligulata C.Presl. --Lastrea foxii (Christ) Copel., Fern Fl. Philip. 2 (1960) 328. Moluccas, Philippines. LUZON to MINDANAO, PALAWAN. Photos 
5. Pneumatopteris lithophila Holttum, Fl. Males., Ser. 2, Pterid. 1 (1982) 422. Endemic to the Philippines. SAMAR.
6. Pneumatopteris michaelis Holttum, Blumea 21 (1973) 313. Endemic to the Philippines. LUZON (Mt Makiling).
7. Pneumatopteris microloncha (Christ) Holttum, Blumea 21 (1973) 308; --Cyclosorus microlonchus (Christ) Ching, Bull. Fan Mem. Inst. Biol. Bot. 10 (1941) 246; --Dryopteris microloncha Christ, PJS 2 c (1907) 202. Endemic to the Philippines. LUZON, NEGROS.
8. Pneumatopteris nitidula (C.Presl) Holttum, Blumea 21 (1973) 318; --Cyclosorus nitidulus (C.Presl) Copel., Fern Fl. Philip. 2 (1960) 364; --Nephrodium nitidulum C.Presl, Epimel. Bot. (1851) 46. Endemic to the Philippines. LUZON to MINDANAO. Photos
9. Pneumatopteris truncata (Poir.) Holttum, Blumea 21 (1973) 314; --Cyclosorus truncatus (Poir.) Farw., Amer. Midl. Naturalist 12 (1931) 259; --Polypodium truncatum Poir., Encycl. 5 (1804) 534. Bangladesh, Borneo, ?Cambodia, China, Christmas Isl, India, Java, Laos, Lesser Sunda Isls, Myanmar, New Caledonia, Ryukyu Isls, Singapore, Sri Lanka, Sulawesi, Sumatra, Taiwan, Tibet, Thailand, Vietnam, Philippines. PALAWAN. Photos

Pronephrium C.Presl

Last edited by Julie F. Barcelona, 7 September 2020

1. Pronephrium amphitrichum Holttum, Fl. Males., Ser. 2, Pterid. 1 (1982) 521. Endemic to the Philippines. LUZON. Photos
2. Pronephrium asperum (C.Presl) Holttum, Blumea 20 (1972) 112; --Goniopteris aspera C.Presl, Tent. Pterid. (1836) 183. --Cyclosorus philippinarum (Fée) Copel., Fern Fl. Philip. 2 (1960) 372; --Abacopteris philippinarum Fée, Congr. sci. de France X sess. 1 (1843) 178. Australia, Borneo, Java, Lesser Sunda Isls, Moluccas, New Guinea, Solomon Isls, Sulawesi, Sumatra, Thailand, Vietnam, Philippines.
3. Pronephrium bakeri (Harrington) Holttum, Fl. Males., Ser. 2, Pterid. 1 (1982) 516; --Haplodictyum bakeri (Harrington) Ching, Sunyatsenia 5 (1940) 251; --Cyclosorus bakeri (Harrington) Copel., Gen. Fil. (1947) 142; --Nephrodium bakeri Harr. -- J. Linn. Soc., Bot. 16 (1877) 29. Endemic to the Philippines. LUZON, MINDANAO, NEGROS, PANAY. Photos
4. Pronephrium bulusanicum (Holttum) Holttum, Fl. Males., Ser. 2, Pterid. 1 (1982) 515; --Haplodictyum bulusanicum Holttum, Kalikasan 2 (1973) 61. Endemic to the Philippines. LUZON. Conservation status: Vulnerable (DAO 2017-11).
5. Pronephrium clemensiae (Copel.) Holttum, Blumea 20 (1972) 118; --Cyclosorus clemensiae (Copel.) Ching, Bull. Fan Mem. Inst. Biol. Bot. 10 (1941) 243; --Dryopteris clemensiae Copel., PJS 46 (1931) 213. Endemic to the Philippines. LUZON. Photos
   • var. clemensiae. LUZON.
   • var. degenerum (Christ) Holttum; --Cyclosorus degener (Christ) Copel.; --Dryopteris canescens var. degener Christ. LUZON.
6. Pronephrium cuspidatum (Blume) Holttum, Blumea 20 (1972) 123; --Cyclosorus cuspidatus (Blume) Copel., Gen. Fil. (1947) 142; --Meniscium cuspidatum Blume, Enum. Pl. Javae 2 (1828) 114. Borneo, Java, Lesser Sunda Isls, Moluccas, Nicobar Isls, Solomon Isls, Sumatra, Philippines.
7. Pronephrium granulosum (C.Presl) Holttum, Blumea 20 (1972) 119; --Cyclosorus granulosus (C.Presl) Copel., Fern Fl. Philip. 2 (1960) 373; --Polypodium granulosum C.Presl, Reliq. Haenk. 1 (1825) 24. --Cyclosorus subdimorphus Copel., PJS 81 (1952) 38. Lesser Sunda Isls, Philippines. BALABAC, LUZON, MINDANAO: Bukidnon, PALAWAN, PANAY. Photos
8. Pronephrium gymnopteridifrons (Hayata) Holttum, Blumea 20 (1972) 112; --Dryopteris gymnopteridifrons Hayata, Icon. Pl. Formosan. 8 (1919) 148. --Cyclosorus pustulosus Copel., PJS 81 (1952) 37. China, Taiwan, Philippines.
9. Pronephrium heterophyllum (C.Presl) Holttum, Fl. Males., Ser. 2, Pterid. 1 (1982) 515; --Haplodictyum heterophyllum C.Presl, Epimel. Bot. (1851) 51. Endemic to the Philippines. LUZON, MINDANAO, SAMAR. Photos
10. Pronephrium hosei (Baker) Holttum, Blumea 20 (1972) 120; --Cyclosorus hosei (Baker) Copel., Gen. Fil. (1947) 142; --Meniscium hosei Baker, J. Linn. Soc., Bot. 22 (1886) 230. Borneo, Philippines. BASILAN, MINDANAO: Zamboanga. Conservation status: Vulnerable (DAO 2017-11).
11. Pronephrium lineatum (Blume) C.Presl, Epimel. Bot. (1851) 259; --Aspidium lineatum Blume, Enum. Pl. Javae 2 (1828) 144. Borneo, Java, Moluccas, Philippines. TAWITAWI.
12. Pronephrium melanophlebium (Copel.) Holttum, Blumea 20 (1972) 108; --Cyclosorus melanophlebius (Copel.) Ching, Bull. Fan Mem. Inst. Biol. Bot. 10 (1941) 246; --Dryopteris melanophlebia Copel., PJS c 6 (1911) 147. Moluccas, Philippines. ?LUZON, MINDANAO, NEGROS.
13. Pronephrium menisciicarpon (Blume) Holttum, Blumea 20 (1972) 111; --Aspidium menisciicarpon Blume, Enum. Pl. Javae 2 (1828) 142. --Cyclosorus latifolius (C.Presl) Copel., Fern Fl. Philip. 2 (1960) 370; --Nephrodium latifolium C.Presl, Epimel. Bot. (1851) 45. Bismarck Arch, Borneo, Java, Moluccas, New Guinea, ?Nicobar Isls, Peninsular Malaysia, Singapore, Sulawesi, Sumatra, Thailand, Vietnam, Philippines. LEYTE, SAMAR.
14. Pronephrium merrillii (Christ) Holttum, Blumea 20 (1972) 117; --Cyclosorus merrillii (Christ) Ching, Bull. Fan Mem. Inst. Biol. Bot. 10 (1941) 246; --Dryopteris merrillii Christ, PJS 2 c (1907) 201. --Pronephrium balabacensis P.M.Zamora & Co, Nat. Appl. Sci. Bull. 32 (1980) 45. Endemic to the Philippines. BALABAC, PALAWAN.
15. Pronephrium nitidum Holttum, Blumea 20 (1972) 109. --Cyclosorus urophyllus auct. non (Wall. ex Hook.) Tardieu ex Tardieu & C.Chr., sensu Copel. (1960), excl. loc. Malacca. Borneo, Sulawesi, Sumatra, Philippines. MINDANAO: Bukidnon. Photos
16. Pronephrium ramosii (Christ) Holttum, Fl. Males., Ser. 2, Pterid. 1 (1982) 537; --Dryopteris ramosii Christ, PJS c 2 (1907) 203. Moluccas, New Guinea, Philippines. LUZON to MINDANAO. Photos
    • var. ramosii. --Cyclosorus cuspidatus auct. non (Blume) Copel., sensu Copel. (1960), excl. syn.
17. Pronephrium rhombeum (Christ) Holttum, Blumea 20 (1972) 120; --Cyclosorus rhombeus (Christ) Ching, Bull. Fan Mem. Inst. Biol. Bot. 8 (1938) 192. --Cyclosorus lanceola (Christ) Copel., Gen. Fil. (1947) 142; --Dryopteris diversiloba subvar. lanceola Christ. Borneo, Philippines. LUZON to MINDANAO. Photos
18. Pronephrium rubidum (J.Sm. ex Hook.) Holttum, Blumea 20 (1972) 111; --Cyclosorus rubidus (J.Sm. ex Hook.) Copel., Fern Fl. Philip. 2 (1960) 375; --Polypodium rubidum Hook., Sp. Fil. 5 (1863) 12. Endemic to the Philippines. LUZON, PALAWAN.
19. Pronephrium samarense (Copel.) Holttum, Blumea 20 (1972) 119; --Cyclosorus samarensis Copel., PJS 81 (1952) 35. Endemic to the Philippines. SAMAR.
20. Pronephrium solsonicum Holttum, Kalikasan 3 (1974) 197. Endemic to the Philippines. LUZON. Conservation status: Vulnerable (DAO 2017-11).
21. Pronephrium trachyphyllum Holttum, Fl. Males., Ser. 2, Pterid. 1 (1982) 525. Endemic to the Philippines. MINDANAO.
22. Pronephrium triphyllum (Sw.) Holttum, Blumea 20 (1972) 122; --Cyclosorus triphyllus (Sw.) Tardieu, Notul. Syst. (Paris) 7 (1938) 77; --Meniscium triphyllum Sw., J. Bot. (Schrader) (1800) 16. Australia, Bangladesh, Borneo, China, Hong Kong, India, Japan, Java, Laos, Lesser Sunda Isls, Moluccas, Myanmar, New Guinea, Nicobar Isls, Pacific, Peninsular Malaysia, Ryukyu Isls, Singapore, Sri Lanka, Sulawesi, Sumatra, Taiwan, Thailand, Vietnam, Philippines. PALAWAN.
23. Pronephrium xiphioides (Christ) Holttum, Blumea 20 (1972) 119; --Cyclosorus xiphioides (Christ) Ching, Bull. Fan Mem. Inst. Biol. Bot. 10 (1941) 250; --Dryopteris xiphioides Christ, PJS c 2 (1907) 201. --Cyclosorus edanyoi Copel., PJS 81 (1952) 37. --Cyclosorus rhombeus auct. non (Christ) Ching, sensu Copel. (1960), pro parte. Endemic to the Philippines. MINDANAO: Bukidnon. Photos

Pseudocyclosorus Ching

1. Pseudocyclosorus tylodes (Kunze) Ching, Acta Phytotax. Sin. 8 (1963) 323; --Lastrea tylodes (Kunze) Moore; --Aspidium tylodes Kunze, Linnaea 24 (1851) 281, as ‘xylodes’; --Li et al., Phytotaxa 424 (2019) 204; --Type: from India. Bhutan, China, India, Jammu & Kashmir, Lesser Sunda Isls, Myanmar, Nepal, Sri Lanka, Tibet, Taiwan, Thailand, Vietnam, Philippines. LUZON.

Pseudophegopteris Ching = Phegopteris

Sphaerostephanos J.Sm.

1. Sphaerostephanos acrostichoides (Desv.) Holttum, Fl. Males., Ser. 2, Pterid. 1 (1982) 456; --Nephrodium acrostichoides Desv., Mém. Soc. Linn. Paris 6 (1827) 255. --Sphaerostephanos smithianus (C.Presl) Holttum, Kalikasan 4 (1975) 65; --Nephrodium smithianum C.Presl -- Epimel. Bot. (1851) 46. --Cyclosorus boholensis Copel., PJS 81 (1952) 31. --Cyclosorus productus sensu Copel. (1960), pro parte. Bismarck Arch, Java, Lesser Sunda Isls, Moluccas, New Guinea, Sulawesi, Philippines. BOHOL, GUIMARAS, MINDANAO.
2. Sphaerostephanos alcasidii Holttum, Fl. Males., Ser. 2, Pterid. 1 (1982) 503. Endemic to Philippines. LUZON: Mountain Province (Mt Data).
3. Sphaerostephanos angustifolius (C.Presl) Holttum, Kalikasan 4 (1975) 66; --Cyclosorus angustifolius (C.Presl) Copel., Fern Fl. Philip. 2 (1960) 349; --Nephrodium angustifolium C.Presl, Epimel. Bot. (1851) 48. New Guinea, Philippines. Conservation status: Vulnerable (DAO 2017-11).
4. Sphaerostephanos appendiculatus (Blume) Holttum, Kalikasan 4 (1975) 62; --Gymnogramma appendiculata Blume, Enum. Pl. Javae 2 (1828) Add. Java, Lesser Sunda Isls, Sulawesi, Sumatra, Philippines. LUZON. Photos
5. Sphaerostephanos cartilagidens P.M.Zamora & Co, Nat. Appl. Sc. Bull. Univ. Philipp.  32 (1980) 43. Endemic to the Philippines. PALAWAN. Conservation status: Other Threatened Species (DAO 2017-11).
6. Sphaerostephanos convergens Holttum, Fl. Males., Ser. 2, Pterid. 1 (1982) 477. Endemic to the Philippines. SAMAR.
7. Sphaerostephanos dichrotrichoides (Alderw.) Holttum, Kalikasan 4 (1975) 65; --Lastrea dichrotrichoides (Alderw.) Copel., Fern Fl. Philip. 2 (1960) 329; --Dryopteris dichrotrichoides Alderw., Mal. Ferns Suppl. Corr. (1917) 48. Endemic to the Philippines. MINDANAO, ?PANAY. Conservation status: Other Threatened Species (DAO 2017-11).
8. Sphaerostephanos diminutus (Copel.) M.G.Price, Contr. Univ. Michigan Herb. 17 (1990) 277; --Cyclosorus diminutus (Copel.) Copel., Gen. Fil. (1947) 142; --Pronephrium diminutum (Copel.) Holttum, Blumea 20 (1972) 115; --Dryopteris diminuta Copel., PJS 40 (1929) 298. --Sphaerostephanos lucbanii Holttum, Fl. Males., Ser. 2, Pterid. 1 (1982) 463. Endemic to the Philippines. LUZON, MINDANAO.
9. Sphaerostephanos diversilobus (C.Presl) Holttum, Kalikasan 4 (1975) 57; --Cyclosorus diversilobus (C.Presl) Ching, Bull. Fan Mem. Inst. Biol. Bot. 10 (1941) 244; --Nephrodium diversilobum C.Presl, Epimel. Bot. (1851) 47. Sulawesi, Philippines. LUZON, PANAY. Photos
10. Sphaerostephanos ellipticus (Rosenst.) Holttum, Kalikasan 4 (1975) 66; --Cyclosorus ellipticus (Rosenst.) Copel., Gen. Fil. (1947) 142; --Dryopteris elliptica Rosenst., Meded. Rijks-Herb. 31 (1917) 6. --Sphaerostephanos ellipticus var. glabrior Holttum. Endemic to the Philippines. MINDANAO, NEGROS.
11. Sphaerostephanos erectus (Copel.) Holttum, Kalikasan 4 (1975) 64; --Cyclosorus erectus Copel., PJS 81 (1952) 30. Endemic to the Philippines. BILIRAN, LEYTE.
12. Sphaerostephanos foxworthyi Holttum, Fl. Males., Ser. 2, Pterid. 1 (1982) 475. Endemic to the Philippines. LUZON.
13. Sphaerostephanos grandescens Holttum, Fl. Males., Ser. 2, Pterid. 1 (1982) 465. Endemic to the Philippines. MINDORO (Mt Halcon).
14. Sphaerostephanos hernaezii Holttum, Fl. Males., Ser. 2, Pterid. 1 (1982) 459. Endemic to the Philippines. SAMAR. Conservation status: Vulnerable (DAO 2017-11).
15. Sphaerostephanos heterocarpus (Blume) Holttum in B.K.Nayar & S.Kaur, Companion Handb. Ferns Brit. India (1974) 209; --Aspidium heterocarpon Blume, Enum. Pl. Javae 2 (1828) 155. ?Andaman Isls, Australia, Borneo, Cambodia, China, Java, Lesser Sunda Isls, Micronesia, Moluccas, New Guinea, ?Nicobar Isls, Pacific, Peninsular Malaysia, Singapore, Solomon Isls, Sulawesi, Sumatra, Thailand, Vietnam, Philippines. Eight variants of this polymorphic species were enumerated (but no formal names given) by Holttum in the his FM revision. Two are definitely occurring in the Philippines, while another 2 resembles those ascribed to Borneo and New Guinea forms. Thus: -Mindanao form -Luzon form [as Cyclosorus bordenii (Christ) Ching in ‘Fern Flora’]. From Palawan is one collection resembling the Borneo form, while another one from San Ramon, Zamboanga is nearer to the New Guinea form. LUZON, MINDANAO, PALAWAN. Photos
16. Sphaerostephanos hirsutus (Kunze ex Mett.) Holttum, Kalikasan 4 (1975) 63; --Aspidium hirsutum Kunze ex Mett., Abh. Senckenberg. Naturf. Ges. 2 (1858) 391. Sulawesi, Philippines.
    
    • var. hirsutus. --Cyclosorus adenophorus (C.Chr.) Ching, Bull. Fan Mem. Inst. Biol. Bot. 10 (1941) 242; --Dryopteris adenophora C.Chr., Index Filic. (1905) 251.
17. Sphaerostephanos irayensis (Copel.) Holttum, Fl. Males., Ser. 2, Pterid. 1 (1982) 495; --Plesioneuron irayense (Copel.) Holttum, Blumea 22 (1975) 239; --Cyclosorus irayensis Copel., PJS 81 (1952) 28; --Type: Ramos BS 80344 (MICH, SING), Batanes: Mt Iraya. --Sphaerostephanos fenixii Holttum, Kalikasan 4 (1975) 55; --Type: Fenix BS 3780 (P, US), Batanes. Endemic to the Philippines. BATANES. Conservation status: Other Threatened Species (DAO 2017-11).
18. Sphaerostephanos lastreoides (C.Presl) Holttum, Kalikasan 4 (1975) 54; --Pronephrium lastreoides C.Presl, Epimel. Bot. (1851) 259. --Cyclosorus duplosetosus Copel., PJS 81 (1952) 31. Endemic to the Philippines. LUZON, MINDANAO, NEGROS, PALAWAN. Photos
19. Sphaerostephanos latebrosus (Kunze ex Mett.) Holttum in B.K.Nayar & S.Kaur, Companion Handb. Ferns Brit. India (1974) 209; --Aspidium latebrosum Kunze, Bot. Zeitung (Berlin) 6 (1848) 261. --Sphaerostephanos todayensis (Christ) Holttum, Kalikasan 4 (1975) 58; --Cyclosorus todayensis (Christ) Ching, Bull. Fan Mem. Inst. Biol. Bot. 10 (1941) 249; --Dryopteris todayensis Christ, PJS c 2 (1907) 193. --Cyclosorus sagittifolioides Copel., PJS 81 (1952) 29. --Cyclosorus halconensis Copel., PJS 81 (1952) 29. --Cyclosorus polypterus Copel., PJS 84 (1955) 161. Borneo, China, India, Nepal, Peninsular Malaysia, Sulawesi, Philippines. PALAWAN.
20. Sphaerostephanos lobatus (Copel.) Holttum, Kalikasan 4 (1975) 64; --Cyclosorus lobatus Copel., PJS 81 (1952) 33. --Cyclosorus bordenii auct. non (Christ) Ching, sensu Copel. (1960), pro parte. Endemic to the Philippines. LUZON, PALAWAN.
21. Sphaerostephanos loherianus (Christ) Holttum, Kalikasan 4 (1975) 61; --Cyclosorus loherianus (Christ) Copel., Gen. Fil. (1947) 142; --Aspidium loherianum Christ, Bull. Herb. Boissier 6 (1898) 191. Endemic to the Philippines. LUZON.
22. Sphaerostephanos magnus (Copel.) Holttum, Kalikasan 4 (1975) 61; --Cyclosorus magnus Copel., PJS 81 (1952) 30. Endemic to the Philippines. NEGROS.
23. Sphaerostephanos major (Copel.) Holttum, Fl. Males., Ser. 2, Pterid. 1 (1982) 464; --Haplodictyum majus Copel., LPB 9 (1920) 3110. --Cyclosorus dimorphus Copel., PJS 83 (1954) 99, nom. illeg., auct. non (Brause) Copel. Endemic to the Philippines. LUZON.
24. Sphaerostephanos norrisii (Rosenst.) Holttum, Fl. Males., Ser. 2, Pterid. 1 (1982) 485; --Dryopteris norrisii Rosenst., Meded. Rijks-Herb. 31 (1917) 8. --Sphaerostephanos elmerorum (Copel.) Holttum, Kalikasan 4 (1975) 55; --Cyclosorus elmerorum (Copel.) Copel., Gen. Fil. (1947) 142; --Dryopteris elmerorum Copel., PJS 40 (1929) 295. --Cyclosorus ellipticus auct non (Rosenst.) Copel., sensu Copel. (1960), pro parte. Borneo, Sumatra, Peninsular Malaysia, Philippines. MINDANAO.
25. Sphaerostephanos perglanduliferus (Alderw.) Holttum, Kalikasan 4 (1975) 59; --Dryopteris perglandulifera Alderw., Bull. Buit. III. 2 (1920) 150. Borneo, Micronesia (Palau), Sulawesi, Sumatra, Philippines. NEGROS.
26. Sphaerostephanos polisianus Holttum, Kalikasan 4 (1975) 65. Endemic to the Philippines. LUZON.
27. Sphaerostephanos polycarpus (Blume) Copel., Univ. Calif. Publ. Bot. 16 (1929) 60; --Aspidium polycarpon Blume, Enum. Pl. Javae 2 (1828) 156. --Sphaerostephanos asplenioides J.Sm. Bismarck Arch,Borneo, Java, Lesser Sunda Isls, New Guinea, Nicobar Isls, Peninsular Malaysia, Samoa, Singapore, Solomon Isls, Sulawesi, Sumatra, Thailand, Vietnam, Philippines.
28. Sphaerostephanos porphyricola (Copel.) Holttum, Kalikasan 4 (1975) 59; --Cyclosorus porphyricola (Copel.) Ching, Bull. Fan Mem. Inst. Biol. Bot. 10 (1941) 248; --Dryopteris porphyricola Copel., PJS c 7 (1912) 60. Borneo, Moluccas, Philippines.
29. Sphaerostephanos productus (Kaulf.) Holttum, Kalikasan 4 (1975) 59; --Cyclosorus productus (Kaulf.) Ching, Bull. Fan Mem. Inst. Biol. Bot. 10 (1941) 248; --Aspidium productum Kaulf., Enum. Filic. (1824) 237. --Cyclosorus serratus Copel., PJS 81 (1952) 36. --Cyclosorus weberi Copel., PJS 81 (1952) 36. --Cyclosorus rigidus Copel., PJS 81 (1952) 27, nom. illeg., auct. non (Ridl.) Copel. --Cyclosorus leucadenius Copel., PJS 81 (1952) 27. --Sphaerostephanos punctatus Holttum, Kalikasan 4 (1975) 60. --Cyclosorus megaphyllus auct. non (Mett.) Ching, sensu Copel. (1960). --Cyclosorus nitidulus auct. non (Presl) Copel., sensu Copel. (1960) pro parte. Taiwan, Philippines. LUZON to MINDANAO.
30. Sphaerostephanos santomasii Holttum, Kalikasan 4 (1975) 62. Endemic to the Philippines. LUZON: Mountain Province.
31. Sphaerostephanos sessilipinna (Copel.) Holttum, Kalikasan 4 (1975) 53; --Cyclosorus sessilipinnus (Copel.) Ching, Bull. Fan Mem. Inst. Biol. Bot. 10 (1941) 249; --Dryopteris sessilipinna Copel., PJS c 6 (1911) 145. --Cyclosorus austrophilippinus (Copel.) Copel., Gen. Fil. (1947) 142; --Dryopteris austrophilippina Copel., PJS 40 (1929) 300. Endemic to the Philippines. MINDANAO, NEGROS.
32. Sphaerostephanos simplicifolius (J.Sm. ex Hook.) Holttum, Kalikasan 4 (1975) 61; --Cyclosorus simplicifolius (J.Sm. ex Hook.) Copel., J. Arnold Arbor. 30 (1949) 438; --Aspidium simplicifolium J.Sm. ex Hook., Icon. Pl. 6 (1854) t. 919. Endemic to the Philippines. BILIRAN, LEYTE, SAMAR.
33. Sphaerostephanos spenceri (Christ) Holttum, Kalikasan 4 (1975) 58; --Cyclosorus spenceri (Copel. ex Christ) Copel., Gen. Fil. (1947) 143; --Dryopteris spenceri Copel. ex Christ, PJS c 2 (1907) 206. Endemic to the Philippines. MINDANAO. Conservation status: Other Threatened Species (DAO 2017-11).
34. Sphaerostephanos stenodontus (Copel.) Holttum, Kalikasan 4 (1975) 56; --Cyclosorus stenodontus Copel., PJS 81 (1952) 28. Endemic to the Philippines. PANAY. Conservation status: Other Threatened Species (DAO 2017-11).
35. Sphaerostephanos subcordatus Holttum, Fl. Males., Ser. 2, Pterid. 1 (1982) 469. Endemic to the Philippines. SAMAR.
36. Sphaerostephanos tephrophyllus (Copel.) Holttum, Kalikasan 4 (1975) 58; --Cyclosorus tephrophyllus (Copel.) Copel., Gen. Fil. (1947) 143; --Dryopteris tephrophylla Copel., PJS 40 (1929) 296. Endemic to the Philippines. MINDANAO. Conservation status: Other Threatened Species (DAO 2017-11).
37. Sphaerostephanos unitus (L.) Holttum, J. S. African Bot. 40 (1974) 165; --Cyclosorus unitus (L.) Ching, Bull. Fan Mem. Inst. Biol. Bot. 8 (1938) 192; --Polypodium unitum L., Syst. Nat., ed. 10. 2 (1759) 1326. MINDANAO: Bukidnon. Photos
    • var. unitus. Sri Lanka, S India, across Malesia. Throughout the Philippines. Terrestrial. Degraded habitats generally, along waterways, etc. at low to medium elevation. Very common.
    • var. mucronatus (Christ) Holttum; --Dryopteris cucullata var. mucronata Christ. Borneo, Lesser Sunda Isls, Moluccas, New Guinea, Solomon Isls, Sulawesi, Philippines.
38. Sphaerostephanos urdanetensis (Copel.) Holttum, Fl. Males., Ser. 2, Pterid. 1 (1982) 467; --Cyclosorus urdanetensis (Copel.) Copel., Gen. Fil. (1947) 143; --Dryopteris urdanetensis Copel., LPB Bot. 5 (1913) 1682. --Cyclosorus matutumensis (Copel.) Copel., Gen. Fil. (1947) 143; --Dryopteris matutumensis Copel., PJS 40 (1929) 299. Endemic to the Philippines. MINDANAO.
39. Sphaerostephanos williamsii (Copel.) Holttum, Kalikasan 4 (1975) 53; --Lastrea williamsii (Copel.) Copel., Gen. Fil. (1947) 140; --Dryopteris williamsii Copel., Brittonia 1 (1931) 67. Endemic to the Philippines. MINDANAO. Conservation status: Other Threatened Species (DAO 2017-11).

Uncertain status:

1. Sphaerostephanos mindorensis Holltum, Kalikasan 4 (1975) 54; --Type: M.G.Price 690 (K,  holotype; isotype in PNH). Mindoro: Mt Halcon. Endemic to the Philippines. MINDORO: Mt Halcon, in forest, 750m. Known from two collections from the type locality. Not accounted for in Holltum’s Flora Malesiana revision.

Stegnogramma Blume

1. Stegnogramma dissitifolia Holttum, Fl. Males., Ser. 2, Pterid. 1 (1982) 543. Endemic to the Philippines. LUZON.
2. Stegnogramma gymnocarpa (Copel.) K.Iwats., Acta Phytotax. Geobot. 19 (1963) 122; --Lastrea gymnocarpa (Copel.) Copel., Gen. Fil. (1947) 139; --Dryopteris gymnocarpa Copel., LPB 3 (1910) 807.  Philippines, ?Sulawesi. MINDANAO.
   
   • ssp. gymnocarpa

Literature

Alston, AHG. 1959. Isoetaceae. Flora Malesiana series 2, 1: 63-64

Amoroso, VB, FP Coritico & PW Fritsch. 2020. Actinostachys minuta, a new species of grass fern from Mindanao, Philippines. PhytoKeys 151: 59-66

Amoroso, VB, YLL Cariño, JC Nobleza & FP Coritico. 2020. Ophioderma subsessile (Ophioglossaceae), a new snake tongue fern species from Mindanao, Philippines. Philippine Journal of Science 150 (S1): 215-221

Amoroso, VB, YLL Cariño, JC Nobleza & FP Coritico. 2021. A new species of Actinostachys (Schizaeaceae) from Mindanao, Philippines. Phytotaxa 508: 101-106.

Barcelona, JF. Picasa photogalery of Philippine pterophytes and lycophytes.

Barcelona, JF. 2002. Philippine pteridophyte collections as a resource for conservation planning. Proceedings of the “International Symposium on the Fern Flora Worldwide: Threats and Responses” sponsored by the British Pteridological Society and IUCN- Species Survival Commission.” Fern Gazette 16(6, 7 & 8): 307-312.

Barcelona, JF. 2003. The taxonomy and ecology of the pteridophytes of Mt. Iraya & vicinity, Batan Island, Batanes Province, Northern Philippines, in Pteridology in the New Millenium (Chandra and Srivastava, eds). Kluwer Academy Publishers. The Netherlands, 299-325.

Barcelona, JF. 2003. Preliminary report on the ferns and fern allies (pteridophytes) of Mt. Bali-it, Balbalasang-Balbalan National Park, Kalinga, Northern Luzon, Philippines. Sylvatrop: The Technical Journal of Philippine Ecosystems and Natural Resources 13: 81-92

Barcelona, JF. 2004. Collection and conservation status of pteridophytes in Panay Island, Philippines. Philipp. Scient. 41:57-73.

Barcelona, JF. 2005. Noteworthy fern discoveries in the Philippines at the turn of the 21st Century. Fern Gazette 17: 139-146

Barcelona, JF, B. Hernaez, & MG Price. 1996. Philippine Schizaea. Asia Life Sciences 5: 27-34

Barcelona, JF & MG Price. 1999. A new Philippine Goniophlebium. Fern Gazette 15: 261-264.

Barcelona, JF, NE Dolotina, GS Madroñero, WG Granert & DD Sopot. 2006. The ferns and fern allies of the karst forests of Bohol Island, Philippines. American Fern Journal 96: 1-20

Barcelona, JF, DL Nickrent, JV LaFrankie, JRC Callado & PB Pelser. 2013. Co’s Digital Flora of the Philippines: plant identification and conservation through cybertaxonomy. Philippine Journal of Science 142: 57-67

Barcelona, JF & PB Pelser. 2014. Phanerosorus (Matoniaceae), a new fern genus record for the Philippines. Phytotaxa 170: 133-135.

Bosman, MTM. 1991. A monograph of the fern genus Microsorum (Polypodiaceae) including an attempt towards a reconstruction of the phylogenetic history of the microsoroids. Leiden Botanical Series 14: 1-161

Brownsey PJ. 1987. A review of the fern genus Hypolepis (Dennstaedtiaceae) in the Malesian and Pacific regions. Blumea 32: 227-276

CABI Invasive Species Compendium

Chambers TC & PA Farrant. 2001. Revision of Blechnum (Blechnaceae) in Malesia. Blumea 46: 283-350

Chen, C-W, LT Ngan, A Hidayat, L Evangelista, HP Nooteboom & W-L Chiou. 2014. First insights into the evolutionary history of the Davallia repens complex. Blumea 59: 49-58

Chen, C-W, CJ Rothfels, AMA Mustapeng, M Gubilil, DK Karger, M Kessler & Y-M Huang. 2018. End of an enigma: Aenigmopteris belongs in Tectaria (Tectariaceae: Polypodiopsida). Journal of Plant Research 131: 67-76

Chen, C-W, S Lindsay, CM Mynssen, L-Y Kuo, VB Amoroso & Y-M Huang. 2018. A new Haplopteris species from the Philippines and clarification of the status of H. amboinensis. Garden's Bulletin Singapore 70: 177-190

Chen, C-W, S Lindsay, KT Yong, AMA Mustapeng, VB Amoroso, VD Dang & Y-M Huang. 2019. Clarification of two poorly known vittarioid ferns (Pteridaceae): Haplopteris angustissima and H. capillaris. Systematic Botany 44: 483-493

Christenhusz, MJM, X-C Zhang & H Schneider. 2011. A linear sequence of extant families and genera of lycophytes and ferns. Phytotaxa 19: 7-54.

Coritico FP, VB Amoroso & YC Liu. 2019. Athyrium nakanoi Makino (Athyriaceae), a new record from the Philippines and an identification key to the Malesian Athyrium sect. Polystichoides. Philippine Journal of Systematic Biology 13: 46-50

Coritico, FP, VB Amoroso, FM Acma, YLL Cariño & PW Fritsch. 2020. Ferns and lycophytes of Mt. Tago Range, Bukidnon, southern Philippines: species richness, distribution, and conservation status. Philippine Journal of Science 149: 773-790

Coritico FP & VB Amoroso. 2020. Threatened lycophytes and ferns in four protected areas of Mindanao, Philippines. Nature Conservation Research 5: 78-88

Crane, E.H. 1997. A revised circumscription of the genera of the fern family Vittariaceae. Systematic Botany 22: 509-517

DENR Administrative Order. 2017. Updated national list of threatened Philippine plants and their categories. DAO 2017-11

Ding, H-H, Y-S Chao & S-Y Dong. 2013. Taxonomic novelties in the fern genus Tectaria (Tectariaceae). Phytotaxa 122: 61-64.

Dong, S-H. 2014. Nomenclatural novelties in the fern genus Tectaria (Tectariaceae). Phytotaxa 178: 225-228.

Ebihara A, CR Fraser-Jenkins, BS Parris, XC Zhang, YH Yang, WL Chiou, HM Chang, S Lindsay, D Middleton, M Kato, TN Praptosuwiryo, VB Amoroso, JF Barcelona, RHG Ranil, CH Park, N Murakami & A Hoya. 2012. Rare and threatened pteridophytes of Asia 1. An enumeration of narrowly distributed taxa. Bulletin of the National Museum of Nature and Science, Series B. 38: 93-119

Ebihara, A, J-Y Dubuisson, K Iwatsuki, S Hennequin, & M Ito. 2006. A taxonomic revision of the Hymenophyllaceae. Blumea 51: 221-280

Field, AR, W Testo, PD Bostock, JAM Holtum & M Waycott. 2016.  Molecular phylogenetics and the morphology of the Lycopodiaceae subfamily Huperzioideae supports three genera: Huperzia, Phlegmariurus and Phylloglossum. Molecular Phylogenetics and Evolution 94: 635-657

Gruezo, WS. 2012. Tmesipteris zamorae: a new species of Tmesipteris Bernh. (Psilotaceae) from the Philippines. Asia Life Sciences 21: 653-670.

Hassler, M & B Schmitt. World ferns: checklist of ferns and lycophytes of the world

Hassler, M. 2019. World ferns: checklist of ferns and lycophytes of the world. Digital resource at www.catalogueoflife.org/col. Species 2000: Naturalis, Leiden, the Netherlands. ISSN 2405-8858.

Hassler, M & B Schmitt. 2019. Checklist of ferns and lycophytes of the world

Hauke, RI. 1963. A taxonomic monograph of the genus Equisetum subgenus Hippochaete. Beih. Nova Hedwigia 8: 1-123.

Hennipman, E. 1970. Some novelties in Bolbitis from Asia and the Pacific (Filices). Blumea 18: 147-149

Hennipman, E. 1974. The lectotypification of Acrostichum acuminatum Willd. and the reinstatement of the name Photinopteris speciosa (Bl.) Presl. Blumea 22: 51-52

Hennipman, E. 1977. A monograph of the fern genus Bolbitis (Lomariopsidaceae). Leiden Botanical Series 2: 1-329

Hennipman, E. 1987. A new species of Lecanopteris (Polypodiaceae) from the Philippines. Blumea 32: 311-312

Hennipman, E & MC Roos. 1983. Phylogenetic Systematics of the Polypodiaceae (Filicales). Mededelingen van het Botanisch Museum en Herbarium van de Rijksuniversiteit te Utrecht 525: 321-342

Hennipman, E & WLA Hetterscheid. 1984. The emendation of the fern genus Christiopteris, including the transference of two taxa to the microsorioid Polypodiaceae. Mededelingen van het Botanisch Museum en Herbarium van de Rijksuniversiteit te Utrecht 539: 1-10

Holttum, RE. 1959. Keys to the genera of Pteropsida. Flora Malesiana series 2, 1: 15-19

Holttum, RE. 1959. Gleicheniaceae. Flora Malesiana series 2, 1: 1-36

Holttum, RE. 1959. Schizaeaceae. Flora Malesiana series 2, 1: 37-61

Holttum, RE. 1959. Cyatheaceae. Flora Malesiana series 2, 1: 65-176

Holttum, RE. 1966. The genera Lomariopsis, Teratophyllum, and Lomagramma in the islands of the Pacific and Australia. Blumea 14: 215-223

Holttum, RE. 1966. The genus Arthropteris J.Sm. in Malesia. Blumea 14: 225-229

Holttum. RE. 1966. Florae Malesianae Precursores XLIV. The fern-genus Elaphoglossum in Malesia, with descriptions of new species. Blumea 14: 317-326

Holttum, RE. 1968. A re-definition of the fern-genus Taenitis Willd. Blumea 16: 87-95

Holttum, RE. 1969. Studies in the family Thelypteridaceae. The genera Phegopteris, Pseudophegopteris, and Macrothelypteris. Blumea 17: 5-32

Holttum, RE. 1971. Studies in the Family Thelypteridaceae. III. A new system of genera in the Old World. Blumea 19: 17-52

Holttum, RE. 1972. Studies in the family Thelypteridaceae IV. The genus Pronephrium Presl. Blumea 20: 105-126

Holttum, RE. 1973. Studies in the family Thelypteridaceae V. The genus Pneumatopteris Nakai. Blumea 21: 293-325

Holttum, RE. 1975. A comparative account of the fern-genera Syngramma J.Sm. and Taenitis Willd., with discussion of their relationships to each other and to other genera. Kew Bulletin 30: 327-343.

Holttum, RE. 1975. Studies in the family Thelypteridaceae VIII. The genera Mesophlebion and Plesioneuron. Blumea 22: 223-250

Holltum, RE. 1975. Studies in the family Thelypteridaceae. IX. The genus Sphaerostephanos in the Philippines. Kalikasan 4: 47-68.

Holttum, RE. 1976. Studies in the family Thelypteridaceae X. The genus Coryphopteris. Blumea 23: 18-47

Holttum, RE. 1977. Studies in the family Thelypteridaceae. XII. The genus Amphineuron Holttum. Blumea 23: 205-218

Holttum, RE. 1981. Thelypteridaceae. Flora Malesiana Ser. 2, 1: 331-599

Holttum, RE. 1984. Studies in the fern-genera allied to Tectaria. III. Aenigmopteris and Ataxipteris, two new genera allied to Tectaria Cav., with comments on Psomiocarpa Presl. Blumea 30: 1-11

Holttum, RE. 1985. Studies in the fern genera allied to Tectaria Cav. IV. The genus Ctenitis in Asia, Malesia and the Western Pacific. Blumea 31: 1-38

Holttum, RE. 1991. Tectaria group. Flora Malesiana series 2, 2: 1-132

Holttum, RE. 1991. New taxa in the Tectaria Group (Polypodiaceae) from Malesia. Blumea 35: 547-557

Holttum, RE & E Hennipman. 1978. Lomariopsis group. Flora Malesiana series 2, 1: 255-330

Holttum, RE, U Sen & D Mittra. 1970. Studies in the family Thelypteridaceae II. A comparative study of the type-species of Thelypteris Schmidel, Cyclosorus Link, and Ampelopteris Kunze. Blumea 18: 195-215

Hovenkamp, PH. 1986. A monograph of the fern genus Pyrrosia (Polypodiaceae). Leiden Botanical Series 9: 1-280

Hovenkamp, PH. 1998. An account of the Malay-Pacific species of Selliguea (Polypodiaceae). Blumea 43: 1-108

Hovenkamp, PH. 1998. Lepisorus in Malesia. Blumea 43: 109-115

Hovenkamp, PH. 2019. Flora of Singapore precursors, 13. New names and lectotypifications in Athyriaceae and Polypodiaceae. Gard. Bull. Singapore 71: 61-67

Hovenkamp, PH, MTM Bosman, E Hennipman, HP Nooteboom, G. Rodl-Linder & MC Roos. 1998. Polypodiaceae. Flora Malesiana series 2, 3: 1-234

Hovenkamp, PH & H Boon-Chuan. 2012. A revision of the fern genus Oleandra (Oleandraceae) in Asia. PhytoKeys 11: 1-37

Hovenkamp, PH & NAP Franken. 1993. An account of the fern genus Belvisia Mirbel (Polypodiaceae). Blumea 37: 511-527

Hovenkamp, PH & F Miyamoto. 2005. A conspectus of the native and naturalized species of Nephrolepis (Nephrolepidaceae) in the world. Blumea 50: 279-322

Hovenkamp, PH & F Miyamoto. 2012. Nephrolepidaceae. Flora Malesiana series 2, 4: 97-122

Hsu, T-C, Y-F Huang & Y-S Chao. 2019. Systematics of Hymenophyllum subgenus Mecodium (Hymenophyllaceae) in Taiwan. Systematic Botany 44: 753-767.

Karger, DN, S Lehtonen, VB Amoroso, M Kessler. 2012. A new species of Lindsaea (Lindsaeaceae, Polypodiopsida) from Mt. Hamiguitan, Mindanao, Philippines. Phytotaxa 56: 15-20

Kato, M. 1993. A taxonomic study of the genus Matonia (Matoniaceae). Blumea 38: 167-172

Kato, M. 1998. Matoniaceae. Flora Malesiana series  2, 3: 289-294

Kato, M, Y Yatabe, N Sahashi & N Murakami. 2001. Taxonomic studies of Cheiropleuria (Dipteridaceae). Blumea 46: 513-525

Kinosian, SP, WD Pearse & PG Wolf. 2020. Cryptic diversity in the model fern genus Ceratopteris (Pteridaceae). Molecular Phylogenetics and Evolution 152: 106938.

Knapp, R. Picasa albums of Taiwanese pterophytes and gynosperms.

Kramer, KU. 1967. The Lindsaeoid ferns of the Old World II. A revision of Tapeinidium. Blumea 15: 545-556

Kramer, KU. 1967. The Lindsaeoid ferns of the Old World III. Notes on Lindsaea and Sphenomeris in the Flora Malesiana area. Blumea 15: 557-574

Kramer, KU. 1971. Lindsaea Group. Flora Malesiana series 2, 1: 177-254

Laferrière, JE. 1998. Equisetaceae. Flora Malesiana series 2, 3: 287-288

Laferrière, JE. 1998. Cheiropleuriaceae. Flora Malesiana series 2, 3: 285-286

Lehnert, M & FP Coritico. 2018. The genus Dicksonia (Dicksoniaceae - Cyatheales) in western Malesia. Blumea 63: 268-278

Lin, YX & R Viane. 2013. Aspleniaceae. In: Pp. 267–316 in Wu, ZY, PH Raven & DY Hong (eds.), Flora of China, Vol. 2-3 (Pteridophytes). Beijing: Science Press; St. Louis: Missouri Botanical Garden Press

Lindsay, S, PH Hovenkamp & DJ Middleton. 2017. New combinations and typifications in Aglaomorpha (Polypodiaceae). Gardens' Bulletin Singapore 69: 149-155

Liu, H-M, R-H Jiang, J Guo, P Hovenkamp, LR Perrie, L Shepherd, S Hennequin & H Schneider. 2013. Towards a phylogenetic classification of the climbing fern genus Arthropteris. Taxon 62: 688-700

Liu, Y-C, CR Fraser-Jenkins, VB Amoroso & W-L Chiou. 2008. Athyrium erythropodum (Woodsiaceae, Pteridophyta), a new Philippine record. Blumea 53: 447-452

Luong, TT, PH Hovenkamp & MSM Sosef. 2015. Revision of the fern genus Orthiopteris (Saccolomataceae) in Malesia and adjacent regions. PhytoKeys 53: 39–71

Masuyama, S & Y Watano. 2010. Cryptic species in the fern Ceratopteris thalictroides (L.) Brongn. (Parkeriaceae). IV. Taxonomic revision. Acta Phytotaxonomica et Geobotanica 61: 75-86

Moody, K. 1989. Weeds reported in rice in South and Southeast Asia. IRRI, Philippines

Murdock, AG. 2008. A taxonomic revision of the eusporangiate fern family Marattiaceae, with description of a new genus Ptisana. Taxon 57: 737-755

Nooteboom, HP. 1992. Notes on Davalliaceae I. The genera Araiostegia, Davallodes, Leucostegia, and Gymnogrammitis. Blumea 37: 165-187

Nooteboom, HP. 1994. Notes on Davalliaceae II. A revision of the genus Davallia. Blumea 39: 151-214

Nooteboom, HP. 1997. The microsoroid ferns (Polypodiaceae). Blumea 42: 261-395

Nooteboom, HP. 1998. Davalliaceae. Flora Malesiana series 2, 3: 235-276

Paclibar, GCB & ER Tadiosa. 2020 Plant species diversity and assessment in Quezon Protected Landscape, Southern Luzon, Philippines. Philippine Journal of Systematic Biology 4: 1-19

Panigrahi, G.1993. Parahemionitis, a new genus of Pteridaceae. American Fern Journal 83: 90-92.

Parris, BS. 2007. Five new genera and three new species of Grammitidaceae (Filicales) and the re-establishment of Oreogrammitis. Garden's Bulletin Singapore 59: 233-274

Parris, BS. 2009. New genera of Malesian Grammitidaceae (Monilophyta). Blumea 54: 217-219

Price, MG. 1990. Philippine fern notes. Contr. Univ. Mich. Herb. 17: 267-278.

Ravensberg, WJ & E Hennipman. 1986. The Pyrrosia species formerly referred to Drymoglossum and Saxiglossum (Filicales, Polypodiaceae). Leiden Botanical Series 9: 281-310

Reid, JD, GM Plunkett, GA Peters. 2006. Phylogenetic relationships in the heterosporous fern genus Azolla (Azollaceae) based on DNA sequence data from three noncoding regions. International Journal of Plant Sciences 167(3): 529-538.

Rödl-Linder. G. 1990. A monograph of the fern genus Goniophlebium (Polypodiaceae). Blumea 34: 277-423

Rödl-Linder, G. 1994. A monograph of the fern genus Thylacopteris (Polypodiaceae). Blumea 39: 351-364

Rolleri, C. 1993. Revision of the genus Christensenia. American Fern Journal 83: 3-19

Roos, MC. 1985. New combinations in Aglaomorpha Schott (Polypodiaceae). Blumea 31: 153

Ruhfel, B, S Lindsay & CC Davis. 2008. Phylogenetic placement of Rheopteris and Monogramma (Pteridaceae, s.l.): evidence from Rbcl sequence data. Systematic Botany 33: 37-43

Salgado, AE. 2017. Does Asplenium falcatum exist in the Philippines? Edinburgh Journal of Botany 74: 1-4

Saunders, RMK. 1998. Azollaceae. Flora Malesiana series 2, 3: 277-284

Schuettpelz, E & KM Pryer. 2007. Fern phylogeny inferred from 400 leptosporangiate species and three plastid genes. Taxon 56: 1037-1050

Schuettpelz, E, H Schneider, L Huiet, MD Windham & KM Pryer. 2007. A molecular phylogeny of the fern family Pteridaceae: assessing overall relationships and the affinities of previously unsampled genera. Molecular Phylogenetics and Evolution 44:1172-1185

Senterre B, G Rouhan, C Morel & JY Dubuisson. 2017. New species and records in the fern genus Didymoglossum for the flora of Seychelles, with notes on the Southeast Asian D. motleyi complex (Hymenophyllaceae). Phytotaxa 292: 201-217

Shang, H, Z-L Liang & L-B Zhang. 2021. Taxonomy of the fern genus Didymochlaena (Didymochlaenaceae) from Asia and Pacific islands based on morphological and molecular evidence with the description of four new species and one new status. Phytotaxa 479: 71-82.

Smith AR, Pryer KM, Schuettpelz E, Korall P, Schneider H & Wolf PG. 2006. A classification for extant ferns. Taxon 55: 705–731

Tan, BC. 2013. The Selaginella flora, a good indicator of the Philippine pleistocene island groups. Philippine Journal of Science 142: 211-213

Testo, WL, AR Field, AB Sessa & M Sundue. 2019. Phylogenetic and morphological analyses support the resurrection of Dendroconche and the recognition of two new genera in Polypodiaceae subfamily Microsoroideae. Systematic Botany 44: 1-16

TPG. 2019. Updating Taiwanese pteridophyte checklist: a new phylogenetic classification. Taiwania 64: 367-395

Tryon. 1941. A revision of the genus Pteridium. Rhodora 43: 1-70.

Wang, LI, Z-Q Wu, QP Xiang, J Heinrichs, H Schneider & XC Zhang. 2010. A molecular phylogeny and a revised classification of tribe Lepisoreae (Polypodiaceae) based on an analysis of four plastid DNA regions. Botanical Journal of the Linnean Society 162: 28-38

Wardani, W & B Adjie. 2018. The genus Athyrium Roth in Malesia: a checklist with working names and distributions. Sibbaldia 16: 39-48

Wieffering, JH. 1964. A preliminary revision of the Indo-Pacific species of Ophioglossum (Ophioglossaceae). Blumea 12: 321-337

Yesilyurt, JC & H Schneider. 2010. The new fern genus Calciphilopteris (Pteridaceae). Phytotaxa 7: 52-59.

Zhang, L-B. 2012. Reducing the fern genus Dryopsis to Dryopteris and the systematics and nomenclature of Dryopteris subgenus Erythrovariae section Dryopsis (Dryopteridaceae). Phytotaxa 71: 17-27.

Zhang, L-B & L Zhang. 2012. The inclusion of Acrophorus, Diacalpe, Nothoperanema, and Peranema in Dryopteris: The molecular phylogeny, systematics, and nomenclature of Dryopteris subg. Nothoperanema (Dryopteridaceae). Taxon, 61: 1199-1216

Zhang, XC. 2003. New combinations in Haplopteris (Pteridophyta: Vittariaceae). Annales Botanici Fennici 40: 459-461

Zhang XC & HP Nooteboom. 1997. Some new combinations in Plagiogyria (Plagiogyriaceae, Pteridophyta). Blumea 42: 483-484

Zhang XC & HP Nooteboom. 1998. A taxonomic revision of Plagiogyriaceae (Pteridophyta). Blumea 43: 401-469

Zhang XC & HP Nooteboom. 1998. Plagiogyriaceae. Flora Malesiana series 2, 3: 295-316

Zhou, S, W Dong, X Chen, X Zhang, J Wen & H Schneider. 2014. How many species of bracken (Pteridium) are there? Assessing the Chinese brackens using molecular evidence. Taxon 63: 509-521

Zhou, X-M & L-B Zhang. 2019. A monograph of the fern genus Pteridrys (Pteridryaceae). Systematic Botany 44 (2019) 243-273.

Acknowledgements

Barcelona, J.F. (2011 onwards). Pteridophytes (Lycophytes and Monilophytes). In: Pelser, P.B., J.F. Barcelona & D.L. Nickrent (eds.). Co's Digital Flora of the Philippines. www.philippineplants.org

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Last updated 11 July 2021